Chapter 46 Osmoregulation + waste disposal • Water!!!!! Is important to all critters!! • Homeostasis: maintaining a constant internal environment - Metabolic wastes also need to be controlled (excretion) •CO 2 Nitrogenous wastes: Proteins AA A A N (Nitrogen-containing amino group) •Water •Nitrogenous wastes How do critters deal with this problem? • Fish eliminate Ammonia - TOXIC Ammonia Osmosis: The movement of water across a semi-permeable membrane dilution is the solution • Mammals and Amphibians - Urea in water Hypertonic: membrane High conc. solutes • Birds - Uric Acid (Crystalline Paste) Hypotonic: Low conc. solutes - protonephridia w/ flame cells • Proto - nephridial, flame cells (closed tubules) • Metanephridia • Malpighian tubules • Green glands • Kidneys 1 Fig. 46.4 - metanephridia (opened tubules) Fig. 46.5 Solution for fresh H2O fishes : Vertebrates • Kidney (osmoregulation and excretion) - Cleans blood of wastes and maintains water balance Fish??? Fresh water and salt water fishes - Drink no water - Salts leave by diffusion/Active uptake of salts by gills - excrete large quantities of dilute urine Fresh H2O fish body fluids are Hypertonic - i.e., Body fluids have high solutes (salt conc.) - H2O will enter osmotically 2 Salt H2O fishes Osmoregulation in Freshwater Fish - Body fluids are hypotonic (body fluids has low solute conc. (salt conc.) - water tends to leave the fish by osmosis Solution: - drink lots of water - excrete the salt (through gills) - small volumes of isotonic urine Fresh H 2O fish body fluids are Hypertonic Mammalian Kidney Marine Types - Body fluids are hypotonic • The mammalian kidney helps maintain homeostasis – Excretes the majority of the nitrogenous wastes and helps maintain fluid balance • Other excretory organs are the lungs, skin, and digestive system - Kidney reabsorbs urea – tissue is hypertonic Excretory Functions and Organs The Urinary System • The kidneys, urinary bladder, and their ducts make up the urinary system • The ureters carry urine to the urinary bladder, which is a distensible, muscular sac • During urination, urine flows through the urethra • In males, the urethra also carries semen 3 Human Urinary System Mammalian Kidney Human Urinary System Mammalian kidneys have nephrons • The outer portion of the kidney is the cortex; the inner is the medulla • The medulla is composed of 8-10 pyramids, each with nephrons, the ends of which are the collecting ducts • The urine flows from a collectting ducts to renal pelvis - 4 parts 1) Bowman’s capsule (which houses the glomerulus) 2) Proximal tube 3) Loop of Henle 4) Distal tube - Many nephrons empty into a single Collecting duct then to ureter Filtration, Secretion, and Reabsorption Proximal tubule Bowman’s capsule Distal tubule Glomerulus Fluid Solutes: Glucose A.A. Na K+ Urea No to: Blood cells proteins H2O Collecting duct H2 O Na Cl H2 O NaCl NaCl NaCl NaCl U r e a H2 O Loop of Henle Fig. 46.13 4 • Water balance controlled by brain and hormones. - e.g. Water stress Blood pressure decreases Hormones released by Pituitary (antidiuretic hormone) (ADH) Collecting duct becomes more permeable to H 2O (i.e., water is reabsorbed) Additionally when under water stress • Decrease in BP causes Adrenal Glands to release Aldosterone • Aldosterone stimulates sodium reabsorption across the nephron walls • This causes increased osmosis out of nephron into the kidney tissue of water • Too much water Pituitary and Adrenals stop release of hormones 5