Chapter 16 Evolution Chapter 16 Evolution ESSENTIAL QUESTIONS Who was Charles Darwin and what is his theory of Natural Selection What is Evolution and how do mutations allow it to happen? Are humans still evolving? Why or Why not? Why is the concept of Evolution just a theory and not considered a Scientific Law? What is the difference between these two types of Scientific Certainty? Chapter 16 Evolution • Charles Darwin is the father of evolutionary theory! • MUTATIONS are natural changes in organisms, occasionally(very rarely,) they’re beneficial. • Humans are still evolvinghere’s a photo of me from high school. Just kidding…INDIVIDUALS can’t evolve! ….but our species is. • Evolution is a theory but some believe it may be the most influential in modern scientific thought because of its very broad implications Emergence of Evolutionary Thought •Evolution is the process by which SPECIES change over time. A Theory is a BROAD explanation that has been tested and supported. •Like most theories, evolutionary theory keeps developing and expanding. Most of Darwin’s ideas remain scientifically supported. The Emergence of Evolutionary Thought Early Explanations of life’s diversity If species were individually and perfectly created at one time and one place Then…. Why were organisms different in different regions of earth? Why are current organisms different from ancient organisms? If each species is different, why are there the same fundamental structures and body plans to do different things in animals? Charles Darwin 1809-1882 Charles Darwin was the NATURALIST aboard the HMS Beagle. His job was to collect animals and plants for identification and potential commercial value. The Beagle’s mission was to map the ports of South America. They left in 1831 and returned FIVE years later in 1836. Young bull Charles Darwin 1809-1882 Darwin witnessed a tremendous diversity of animal life both living and FOSSILIZED He traveled to the Galapagos Islands and collected a bunch of different birds, but it turned out they were all FINCHES! He became convinced that species change over time Old bull Darwin’s Influences Darwin was influenced by a number of Scientists and their thoughts including: Jean Baptiste Lamarck Thomas Malthus Charles Lyell & James Hutton Georges Cuvier Darwin’s Influences: Jean Baptiste Lamarck Lamarckian Inheritance Lamarck noticed that each organism is usually well ADAPTED to its environment. He proposed that organisms change over time as they adapt to changing environments. Darwin once accepted this idea because it proposed a role for inheritance in evolution. Larmack thought early Giraffes stretched their necks to feed and they’re newly acquired Long neck was then passed off to offspring…..WRONG!!! Acquired traits are not hereditable (or are they?) Lamarckian Inheritance Some of Lamacks ideas still have relevance today though and he may be right in a way…check out Epigenetics for an extra credit project! Darwin’s Influences: Thomas Malthus Malthus: “The tragedy of the COMMONS” Populations tend to grow faster than the food supply needed to feed it Darwin’s Influences: Charles Lyell & James Hutton Lyell & Hutton- GRADUALISM Changes occur over vast periods of time in geology; land formations move and topography changes Darwin’s Influences: Georges Cuvier Cuvier - CATASTROPHISM Geologic changes occur rapidly and have catastrophic outcomes for living things, …..like extinction Darwin’s Theory: DESCENT with Modification Darwin’s Dangerous Idea: He didn’t publish his findings until he was 50! “On the Origin of Species” 1859 Evolutionary pathway resembled a branched tree NOT a ladder from lower forms to higher forms All CONTEMPORARY species have descended from earlier species Darwin’s Observations Individuals in a population VARY, some of these variations are inheritable Populations produce more OFFSPRING than the environment’s food, space, and other assets. Therefore, species COMPETE for resources EVOLUTION BY NATURAL SELECTION Evolution is a change in the inherited characteristics of a population from one generation to the next. Darwin’s theory predicts that; Over time, the number of individuals that carry ADVANTAGEOUS traits will increase in a population. Every living thing has the potential to produce many offspring, but not all of those offspring are likely to SURVIVE and reproduce. -Individuals that have traits that suit their environment are more likely to survive. -Individuals that have certain traits tend to produce more offspring than others do. -These differences are part of natural selection. The Mechanism of Natural Selection Selection and Adaptation Darwin’s theory explains why living things vary in form and match their environment. Each HABITAT presents unique challenges and opportunities to survive and reproduce. So, each species evolves because of the “selection” of those individuals that survive the challenges or make best use of the opportunities. The Mechanism of Natural Selection Put another way, each species becomes adapted to its environment as a result of living in it over time. An ADAPTATION is an inherited trait that is present in a population because the trait helps individuals survive and reproduce in a given environment. Darwin’s theory explains evolution as a gradual process of adaptation. Note that Darwin’s theory refers to populations and species—not individuals— as the units that evolve. –Also, keep in mind that a species is a group of populations that can INTERBREED. DARWINS THEORY OF EVOLUTION BY NATURAL SELECTION 4 Steps in Darwin’s Theory of Evolution by Natural Selection. Step 1 Overproduction: Every population is capable of producing more offspring than can possibly survive. Step 2 Variation: Variation exists within every population. Much of this variation is in the form of inherited traits. Step 3 Selection: In a given environment, having a particular trait can make individuals more or less likely to survive and have successful offspring. So, some individuals leave more offspring than others do. Step 4 Adaptation: Over time, those traits that improve survival and reproduction will become more common. The Mechanism of Natural Selection VOCABULARY PART I CHARLES DARWIN : The Father of Evolution through Natural Selection he observed change thorugh DESCENT WITH MODIFICATION SPECIES : Organisms so similar they can interbreed and produce VIABLE (fertile) offspring NATURAL SELECTION: greater reproductive success displayed by individuals with ADAPTIVE traits VARIATION: differences within a population ADAPTATIONS: traits that are selected for because they help an organism survive and reproduce EVOLUTION: Changes ins species over time- Occurs when genes in a population change- shift to enhance survival and reproduction JEAN BAPTISTE LAMARCK: Observed organisms change/evolve by passing down AQUIRED TRAITS LYELL AND HUTTON: Geologists who proposed GRADUALISM as the source of change CUVIER: Geologist who proposed CATASTROPHISM as the source of change EVIDENCE FOR EVOLUTION: Comes from Fossils, Comparative Anatomy, Embryology, Biochemistry, Biogeography, and others ! Evidence for Evolution Evidence for Evolution comes from each of the following disciplines The Fossil Record Biogeography Development Biology or Embryology Comparative Anatomy Biochemistry Evidence for Evolution A Fossil is a record of an animal that is preserved in rock Found in SEDIMENTARY Rock Fossils show overall structural scheme of how organism evolved Including: Body scheme Feeding behavior Mobility Social Structure New fossils in upper layers, older fossils in lower layers Evidence for Evolution Evidence from the fossil record allows us to piece together an Evolutionary SEQUENCE The Evolutionary Sequence of Modern Cetacea (Whales) Social Structure New fossils in upper layers, older fossils in lower layers Comparative Anatomy HOMOLOGOUS Structure: Similar structures in related organisms suggests organisms share a common ancestor Deer vs Kangaroo Human Hand = Whale flipper ANALOGOUS Structures Similar structures in nonrelated organisms show evolution recreates similar solutions to different environmental challenges Vestigial Organs CONTEMPORARY Organs with no apparent use, but resemble functional structures in ancestors Tailbones Wisdom Teeth Appendix Whale Pelvis Comparative Embryology Ontogeny Recapitulates Phylogeny- Organisms with common descent have similar organs Embryos undergo many physical and genetic changes as they develop into mature forms. Scientists compare the EMBRYONIC development of species to look for similar patterns and structures. Such similarities most likely derive from an ancestor that the species have in common. For example, at some time during development, all vertebrate embryos have a tail. Vertebrates are animals that have backbones. Vertebrates breathe though lungs but as embryos, all vertebrates have gills that becomes the Eustachian tube in man Biochemistry To explain the patterns of change seen in anatomy, scientists make testable predictions. For example, if species have changed over time, the GENES that determine their characteristics should also have changed. Genes can change by mutation and that such change can make new varieties appear. A comparison of DNA or amino-acid sequences shows that some species are more genetically similar than others. Chimps –Humans 99% Homology These comparisons, like those in anatomy, are evidence of hereditary relationships among the species. Biogeography Study of species distribution on Galapagos Islands look similar to South American species Each island would have to had it own special creation event for each species. Darwin contended one species MIGRATED from South America and underwent Adaptive Radiation How Do Variations Arise? •Natural selection IS NOT the cause of variation •Mutation - Random changes in DNA sequences in the parents genome •Gene Shuffling - mixing of parental chromosomes can create new genetic combinations •Variation arises randomly. Variations are then SELECTED on by nature. •Variations which are selected for are called ADAPTATIONS Adaptations and Speciation Types of Adaptations STRUCTURAL Adaptations: Involve structure or anatomy Bird’s beak, Anteaters sticky tongue PHYSIOLOGICAL Adaptations: Functions in organisms Poison Venom in a snake, ink of an octopus BEHAVIORAL Adaptations: Behavior aids in survival and reproduction Wildebeasts/ Caribou/ Bird Migration Wolves Hunt in Packs Fish swim in schools Species & Speciation Species: A group of organisms that are so similar they can interbreed and produce FERTILE offspring Some Organisms are similar enough to form Hybrids-they can breed but the offspring is infertile Horses + Donkey = Mule Lion + Tiger = Liger …Not the same species but a HYBRID Human male + human female = human baby…same species! 3 Types of Natural Selection Directional Selection Shifts a population toward one extreme form of a trait Stabilizing Selection: Shifts a population toward intermediate (medium) traits Disruptive Selection: Extreme phenotypes are more common 3 Types of Natural Selection DIRECTIONAL Selection Shifts a population toward one extreme form of a trait 4 million years ago; heavy cheetahs Over time, light, fast animals reproduced more successfully Natural selection favored the genes that pushed the cheetah’s weight in one direction…lighter Or the Human Brain…it has continued to grow larger with more convolutions (gyrii) over time 3 Types of Natural Selection STABILIZING Selection: Shifts a population toward intermediate (medium) traits Example: Most human babies weight ~ 7pounds 3 Types of Natural Selection DISRUPTIVE Selection: Extreme phenotypes are more common Example: African butterflies appear as either bright orange or bright blue These colors mimic other butterflies that are foul-tasting; less likely to be eaten Intermediate butterflies look like neither, and are more likely to be eaten Patterns of Evolution SPECIATIONOrganisms which are separated by distance + gene pool will no longer interbreed. Each group becomes new species Species become dissimilar or more alike over time through gradual Change 5 Types of Evolution Divergent Evolution: Adaptive Radiation: Convergent Evolution: Coevolution Extinction Patterns of Evolution Divergent Evolution: Single population is split into 2 or more populations. Patterns of Evolution Convergent Evolution: 2 dissimilar species evolve in ways to make them LOOK more similar Sugar Gliders- winged marsupials in Australia Flying Squirrels - Winged squirrels in Northern Hemisphere Flying Squirrel Sugar Glider Patterns of Evolution COEVOLUTION Organisms are part of one other’s environment, so they can affect one another’s evolution. Species that live in close contact often have clear adaptations to one another’s existence. ADAPTIVE RADIATION Over time, species may split into two or more lines of descendants, or lineages. The process tends to speed up when a new species enters an environment that contains few other species. (FINCHES) EXTINCTION If all members of a lineage die off or simply fail to reproduce, the lineage is said to be extinct. The fossil record shows 99% of all species have become EXTINCT. The Pace of Evolution and Earth History Gradualism: Over time, new species arise as environment change If true, we should see intermediate species in fossil record…not always true Catastrophism: Rapid catastrophic events shape earth and living thing THE CURRENT ACCEPTED THEORY Punctuated Equilibrium: Species exist for long periods of time without changes, then large changes occur over short periods of time Populations & Evolution Larger populations tend to have more genetic variation Smaller populations tend to have less genetic variation Genetic Drift/Founder Effect- Loss of genetic variation due small population sizes. Extinction more likely 10,000 years ago, almost all cheetahs go extinct due to climate changes. Resulting cheetahs have little genetic diversity Artificial Selection The Mechanism of Natural Selection VOCABULARY PART II EVIDENCE FOR EVOLUTION FOSSIL EVIDENCE: Records of past organisms show common ancestry COMPARATIVE ANATOMY: the study of similarities in organisms of common ancestry Vestigial Structures- appendix, whale pelvis Analogous Structures – bee wing v. bird wing Homologous structure - your hands-whale flipper EMBRYOLOGY: The study of how embryos develop to show common ancestry- you had gills and a tail! BIOCHEMISTRY: The similarity in gene sequence BIOGEOGRAPHY: The study of how organisms are distibuted today and in the fossil record 3 TYPES OF ADAPTATIONS STRUCTURAL ADAPTATIONS: Physical feature like a wing to fly or fins to swim PHYSIOLOGICAL ADAPTATION: Body functions like the production of venom or the ability to tolerate heat BEHAVIORAL ADAPTATION: When organisms work together like a wolf pack, or school of fish 3 TYPES OFNATURAL SELECTION DIRECTIONAL SELECTION: When a direction in phenotypes is favored- bigger brains STABILIZING SELECTION: When 1 average phenotype is selected for – 7 lb babys DISRUPTIVE SELECTION: When 2 extreme phenotypes are selected for simultaneously- different colored butterflies SPECIATION: The formation of new species through evolution 5 TYPES OF EVOLUTION DIVERGENT: When new species are formed that have different features- Tiger & Lion CONVERGENT: When species change to be similar to other species- Kangaroo & Deer ADAPTIVE RADIATION: When species form to fill new or open ecological niches- Darwin’s Finches COEVOLUTION: When species evolve together and rely on one another for survival- birds & flowers EXTINCTION: When species evolutionary sequence ends-99% of all species over time have become extinct Chapter 17 Population Genetics and Speciation KEY IDEAS How is microevolution studied? How is phenotypic variation measured? How are genetic variation and change measured? How does genetic variation originate? POPULATION GENETICS How is microevolution studied? Microevolution can be studied by observing changes in the numbers and types of alleles in populations. The study of microevolution in this sense is POPULATION genetics. GENETICS AND EVOLUTION Charles Darwin knew that heredity influences characteristics, but he did know about GENES. We can now study and predict genetic variation and change that underlie evolution. MICROEVOLUTION Microevolution or POPULATION GENETICS is evolution at the level of genetic change in populations. The study of genetics and evolution are advancing together. The link from microevolution to macroevolution—SPECIATION can be studied in detail. PHENOTYPIC VARIATION How is phenotypic variation measured? Biologists study polygenic phenotypes by measuring each INDIVIDUALS in the population and then analyzing the distribution of the measurements. The variety of phenotypes that exists for a given characteristic depends on how many genes affect it. PHENOTYPIC VARIATION Polygenic TRAITS or characters are influenced by several genes. Examples include human eye color and height. A distribution is an overview of the relative frequency and range of a set of values. Often, some values in a range are more common than others. A normal distribution, or BELL curve, is one that tends to cluster around an average value in the center of the range. MEASURING VARIATION AND CHANGE How are genetic variation and change measured? Genetic variation and change are measured in terms of the ALLELE FREQUENCY in the GENE POOL of a population. A frequency is the proportion or ratio of a group that is of one type. To study genetic change, the frequency of each allele in a population can be tracked over time. The particular combination of alleles in a population at any one point in time makes up a gene pool. WHAT IS THE ALLELE FREQUENCY FOR THE b eye color gene in your class?? SOURCES OF GENETIC VARIATION How does genetic variation originate? The major source of new alleles in natural populations is MUTATION in Gametes or “germ cells.” Evolution cannot proceed if there is no variation. Mutation generates new alleles at a slow rate. Only mutations in germ cells (egg and sperm) are passed on to offspring. https://www.youtube.com/watch?v=L-cxg8mF_Lw Section 2: Genetic Change KEY IDEAS What does the Hardy-Weinberg principle predict? How does sexual reproduction influence evolution? Why does population size matter? What are the limits of the force of natural selection? What patterns can result from natural selection? EQUILIBRIUM AND CHANGE What does the Hardy-Weinberg principle predict? The Hardy-Weinberg principle predicts that the frequencies of alleles and genotypes in a population will NOT change unless at least one of five forces acts upon the population. The forces that can act against GENETIC EQUILIBRIUM are gene flow, nonrandom mating, genetic drift, mutation, and natural selection. HARDY WEINBERG EQUILIBRIUM A population in which no genetic change occurred would be in a state of genetic equilibrium. Genetic change in a population can be measured as a change in genotype frequency or ALLELE FREQUENCY. A change in one doesn’t necessarily mean a change in the other. THE FIVE CONDITIONS OF HARDY WEINBERG EQUILIBRIUM 1. GENE FLOW Gene flow occurs when genes are added to or removed from a population. Gene flow can be caused by MIGRATION, the movement of individuals from one population to another. THE FIVE CONDITIONS OF HARDY WEINBERG EQUILIBRIUM 2. NON RANDOM MATING In sexually reproducing populations, any limits or preferences of MATE choice will cause nonrandom mating. THE FIVE CONDITIONS OF HARDY WEINBERG EQUILIBRIUM 3. GENETIC DRIFT Chance events can cause RARE alleles to be lost from one generation to the next, especially when populations are small. Such random effects on allele frequencies are called GENETIC DRIFT THE FIVE CONDITIONS OF HARDY WEINBERG EQUILIBRIUM 4. MUTATION Mutation can add a new ALLELE to a population. 5. NATURAL SELECTION Natural selection acts to eliminate individuals with certain traits from a population. As individuals are eliminated, the alleles for those traits may become less frequent in the population. Thus, both allele and genotype frequencies may change. HOW LONG WOULD HE LAST ON THE SAVANNAH?? SEXUAL REPRODUCTION AND EVOLUTION How does sexual reproduction influence evolution? Sexual reproduction creates the possibility that mating patterns or behaviors can influence the gene pool of a population. For example, in animals, females sometimes select mates based on the male’s size, color, ability to gather food, or other characteristics. This kind of behavior is called sexual selection and is an example of nonrandom mating. SEXUAL REPRODUCTION AND EVOLUTION Another example of nonrandom mating is INBREEDING, in which individuals either selffertilize or mate with others like themselves. Inbreeding is more likely to occur if a population is small. In a small population, all members are likely to be closely related. POPULATION SIZE AND EVOLUTION Why does population size matter? Allele frequencies are more likely to remain stable in LARGE populations than in small populations. Population size strongly affects the probability of genetic change in a population. Genetic drift is a strong force in small populations and occurs when a particular allele DISAPPEARS. NATURAL SELECTION AND EVOLUTION What are the limits of the force of natural selection? Natural selection acts only to change the relative frequency of alleles that exist in a population. It acts on genotypes by removing UNSUCCESSFUL phenotypes from a population. NATURAL SELECTION AND EVOLUTION Natural selection is a result of the following facts: –All populations have genetic variation. –Individuals tend to produce more OFFSPRING than the environment can support. –Populations depend upon the reproduction of individuals. Genetic Results of Selection The result of natural selection is that each allele’s frequency may increase or decrease depending on the allele’s effects on survival and reproduction. Although natural selection is not the only force that can cause evolution, it is a powerful force. Why Selection is Limited The key lesson that scientists have learned about evolution by natural selection is that the ENVIRONMENT does the selecting. Only characteristics that are expressed can be targets of natural selection. If a mutation results in rare recessive alleles, for example, selection cannot operate against it. For this reason, genetic disorders (such as cystic fibrosis in humans) can persist in populations. PATTERNS OF NATURAL SELECTION What patterns can result from natural selection? Three major patterns are possible in the way that natural selection affects the distribution of polygenic characters over time. These patterns are Directional selection, Stabilizing selection, and Disruptive selection Directional Selection In directional selection, the “peak” of a normal distribution moves in one direction along its range. In this case, selection acts to eliminate on extreme from a range of phenotypes, making them less common. Stabilizing Selection In stabilizing selection, the bellcurve shape becomes NARROWER. In this case, selection eliminates individuals that have alleles for any extreme type. Stabilizing selection is very common in nature. •Disruptive Selection In disruptive selection, the bell curve is “disrupted” and pushed apart into TWO peaks. In this case, selection acts to eliminate individuals with average phenotype values. Section 3: Speciation KEY IDEAS How can species be defined? How do we know when new species have been formed? Why is studying extinction important to understanding evolution? DEFINING SPECIES A species is generally defined as a POPULATION that can interbreed and usually produce fertile offspring. Other definitions for species may be used for fossils or for organisms that reproduce asexually. Species may be instead be defined based on their physical features, their ecological roles, and their genetic relatedness. FORMING NEW SPECIES How do we know when new species have been formed? Speciation has occurred when the net effects of evolutionary forces result in a population that has unique features and is REPRODUCTIVELY isolated. SPECIATION Each population of a single species lives in a different place. In each place, natural selection acts upon each population and tends to result in offspring that are better adapted to each specific environment. If the ENVIRONMENTS differ, the adaptations may differ. This is called divergence and can lead to the formation of new species. Speciation is the process of forming new species by evolution from preexisting species Reproductive Isolation Reproductive isolation is a state in which two populations can no longer interbreed From this point on, the groups may be subject to different forces, so they will tend to diverge over time. Through divergence over time, populations of the same species may differ enough to be considered SUBSPECIES. Subspecies are simply populations that have taken a step toward speciation REPRODUCTIVE ISOLATION Any of the following mechanisms may contribute to the reproductive isolation of populations: Geography Ecological Niche Mating Behavior and Timing Polyploidy Hybridization EXTINCTION: THE END OF SPECIES Why is studying extinction important to understanding evolution? The species that exist at any time are the net result of both speciation and extinction. Extinction occurs when a species fails to produce any more descendants. Extinction, like speciation, can only be detected after it is complete. EXTINCTION THE END OF SPECIES More than 99% of all of the species that have ever lived becoming extinct. Many cases of extinction are the result of ENVIRONMENTAL change. If a species cannot adapt fast enough to changes, the species may be driven to extinction. CH17 VOCAB BIO CH17 VOCAB Chapter 17 Population Genetics and Speciation • MICROEVOLUTION: the study of changes in allele frequency of a population • POPULATION: A group of organism of the same species that are breeding • PHENOTYPIC VARIATION: the allele frequencies within a population • NORMAL DISTRIBUTION: the bell curve • ALLELE FREQUENCY: the ratio of specific alleles in a population • GENETIC VARIATION: the different forms of alleles and traits in a population HARDY WEINBERG EQUILIBRIUM: an IDEAL condition in which no evolution or allele frequency change occurs - when all of the following DO NOT HAPPEN • • • • • GENE FLOW: the gain or loss of new alleles through immigration or emmigration MUTATION: change in DNA sequence NONRANDOM MATING:reproductive selection, females choose mates selectively GENETIC DRIFT: small populations can have major changes in allele frequency NATURAL SELECTION: when the environment favors a particular allele over another CH17 VOCAB BIO CH17 VOCAB Chapter 17 Population Genetics and Speciation •FOUNDER EFFECT: the initial alleles/individuals in a population have great effect on allele frequency •POPULATION SIZE: the larger the population the smaller the impact of individual changes in allele frequency •SPECIATION: the development of new species through directional, disruptive or stabilizing selection •REPRODUCTIVE ISOLATION: a barrier to reproduction amoung individuals that leads to new species – Geography: physical barriers like rivers, canyons, oceans –squirrels in grand canyon – Ecological Niche: organisms are performing different roles in an ecosystem – Darwin Finches – Mating Behavior and Timing: migrations, mating dances in birds – Polyploidy: changes in chromosome number – Hybridization : pairings between similar species- liger