Synthesis activities of the Western Mountain Initiative • Don McKenzie • Nate Stephenson • Dave Peterson • Jill Baron • Craig Allen • Dan Fagre and the younger cohort • Lindsey Christensen – Postdoctoral reearch associate, Colorado State University • Jeff Hicke – Faculty, University of Idaho • Jeremy Littell – Postdoctoral reearch associate, University of Washington • Phil van Mantgem – Ecologist, USGS Sierra Nevada WMI Project Organization Products and Management Implications Cross-site Interpretations Discharge, Water Quality, Aquatic response Snow, Treeline, Productivity, Fire, Forest Dieback Books, Specialized and Synthetic Papers, Datasets, Models, Website, Graduate degrees, Presentations to managers and scientific meetings Remote Sensing Statistical Analyses Modeling Synthesis Workshops Regional Synthesis Fire-climate Relationships, Forest Dynamics, Climate Variability, Snow and Glaciers Northern Rockies Pacific NW we are here Site Specific Studies Monitoring Paleostudies Modeling Sierra Nevada Disturbance Vegetation dynamics Hydrologic processes Central Rockies Southern Rockies WMI synthesis (West-wide) Fire-climate interactions Wildfire & regional air quality Fire-BGC simulations (2) Insect outbreak dynamics Disturbance interactions & Paleofire-climate workshops Disturbance Vegetation dynamics Hydrologic processes Climate & growth in Douglas-fir Treeline dynamics workshop Demography, productivity, mortality RhesSys simulations (2) Broad-scale NPP RhesSys simulations (2) Regional glacier mapping (2) = cross-site interpretations Hypotheses (sneak preview) • Increasing moisture limits on productivity will alter (tree) species composition by – – – – locally favoring more xeric species. exacerbating episodes of vegetation dieback. altering mortality and turnover rates. Underlying ecological mechanism = large-scale shift to a negative water balance. • Disturbance will be the principal agent of ecosystem change – late 20th-century trends such as increasing insect mortality or fire area burned may be replaced by more abrupt changes. – underlying mechanisms here are complex, operate at multiple scales, and may be constrained by physical limits. Themes • Empirical studies of climate, growth, & productivity • Disturbance will be the principal agent of ecosystem change • Watershed modeling of vegetation & hydrology Climate dimensions of the sample transect The magnitude of the correlation between seasonal hydrological variables and tree-growth depends on the position of the plot along a gradient of surplus water in the environment. Themes • Empirical studies of climate, growth, & productivity • Disturbance will be the principal agent of ecosystem change • Watershed modeling of vegetation & hydrology Projected increases in area burned -- Montana (McKenzie et al. 2004) Northern Ecosystems Eco-provinces with similar climate correlations • More annual precip → less area burned. • Higher summer and growing-season temperature → more area burned. • Current climate is important. • Previous years’ climate has less influence. Littell et al. (2006) Arid/Semi-arid Ecosystem Pattern • Four ecoprovinces with semi-arid vegetation types • Very strong, positive lagged relationships with PPT / PDSI, esp. winter • Weaker - Summer / GS PPT or PDSI Littell et al. (2006) Climate-limited or fuel-limited? • Different fuel types respond differently to climate. • Two mechanisms: drying of fuels and production of fuels. • Drying happens seasonally, whereas production affects fire on scales from years to decades. Rapid climatic change will send ecosystems across disturbance thresholds For example Drastic increase in fire severity Doubling of insect reproductive cycles Mountain pine beetle Pandora moth But threshold behavior is complex For example If insect population cycles are not synchronized with seasons, outbreaks are not likely. Mountain pine beetle Pinus contorta (lodgepole pine) Hicke et al., J. Geophys. Res.-Biogeosciences, 2006 Hicke et al., J. Geophys. Res.-Biogeosciences, 2006 Increasing T leads to… 2. Initial increase in adaptive seasonality at highest elevations 1. Decrease in adaptive seasonality at lowest elevations ∆T = 5° Hicke et al., J. Geophys. Res.-Biogeosciences, 2006 MPB outbreaks in high-elevation whitebark pine Railroad Ridge, ID Today Photo by J. Logan July 2005, Railroad Ridge, ID Quickbird Satellite Imagery Themes • Empirical studies of climate, growth, & productivity • Disturbance will be the principal agent of ecosystem change • Watershed modeling of vegetation & hydrology Climate time series GIS Inputs DEM Initial states Meteorological Processes Canopy Processes Soils Veg Hydrologic Processes Drain LAI Baseline runs Sensitivities Cross-site comparisons Future climate experiments Cross-site comparisons West-wide patterns, gradients, and interpretations Daily streamflow Norm alized Streamflow (mm per day) • • • • • • 90 80 70 60 50 40 30 20 10 0 10 11 12 1 Modeled 2 3 4 Month 5 6 7 8 Observed 9 Stehekin River Lake McDonald RhesSys watersheds Merced River undecided Snake River Transpiration: sensitivity to snowpack at middle elevations 400 600 1200-1800 m 800 1000 1200 1850-2150 m 1000 Merced River 800 Energy limited Transpiration (mm) Water limited 600 400 200 0 2200-2550 m 2600-3900 m 1000 800 600 Energy limited Water limited 400 200 0 400 600 800 1000 1200 Peak snow depth (mm) Christensen et al. in prep Hypotheses (WMI synthesis) • Increasing moisture limits on productivity will alter (tree) species composition by – – – – locally favoring more xeric species. exacerbating episodes of vegetation dieback. altering mortality and turnover rates. Underlying ecological mechanism = large-scale shift to a negative water balance. • Disturbance will be the principal agent of ecosystem change – late 20th-century trends such as increasing insect mortality or fire area burned may be replaced by more abrupt changes. – underlying mechanisms here are complex, operate at multiple scales, and may be constrained by physical limits. Changing relationship over time between precipitation and growth Precipitation coefficient Long-term trend (due to gradual warming) 0 Energy-limited? High variability, sensitive to e.g., excessive snowpack Decoupling Water-limited? Time Very sensitive to e.g., drought Recoupling Jemez Mts., May 2004 Hypotheses (WMI synthesis) • Increasing moisture limits on productivity will alter (tree) species composition by – – – – locally favoring more xeric species. exacerbating episodes of vegetation dieback. altering mortality and turnover rates. Underlying ecological mechanism = large-scale shift to a negative water balance. • Disturbance will be the principal agent of ecosystem change – late 20th-century trends such as increasing insect mortality or fire area burned may be replaced by more abrupt changes. – underlying mechanisms here are complex, operate at multiple scales, and may be constrained by physical limits. Fire Disturbance synergy Climate Vegetation Disturbance drives ecosystem changes 25-100 yr Climatic change 100-500 yr Habitat changes Broad-scale homogeneity Truncated succession Loss of forest cover Loss of refugia Fire-adapted species New fire regimes More frequent fire More extreme events Greater area burned Species responses Fire-sensitive species Annuals & weedy species Specialists with restricted ranges Biggest challenge: integrating across scales Scale mismatch between drivers and responses Multiple stochastic elements with no equilibrium Neilson, Lenihan, Bachelet, et al. Stands ? Landscapes ? Continents Thanks!