ABSTRACTS CIRMOUNT SESSION AGU 2009 Climate Change and Biogeophysical Impacts Across Elevation and Latitude: Are Mountains Different From Poles? Presiding: J S Littell, University of Washington, Seattle; J A Hicke, University of Idaho, Moscow; J D Lundquist, Civil and Environmental Engineering, University of Washington, Seattle; C T Driscoll, Syracuse; J L Campbell, USDA Forest Service, Durham; G T Pederson, Northern Rocky Mountain Science Center, U.S. Geological Survey, Bozeman Water and geochemical responses to seasonal changes across the rain-snow transition in the Southern Sierra Nevada Bales, R C (rbales@ucmerced.edu), University of California, Merced, CA, United States Hunsaker, C T (chunsaker@fs.fed.us), Pacific Southwest Research Station, USDA Forest Service, Fresno, CA, United States Meadows, M (mmeadows@ucmerced.edu), University of California, Merced, CA, United States Kerkez, B (bkerkez@gmail.com), University of California, Berkeley, CA, United States Glaser, S D (glaser@ce.berkeley.edu), University of California, Berkeley, CA, United States Liu, F (fliu@ucmerced.edu), University of California, Merced, CA, United States The hydrologic and geochemical response of rain-dominated versus snow-dominated catchments was investigated using multi-year measurements of precipitation, snow accumulation and melt, streamflow, soil moisture and meteorological variables. Research was carried out at the Kings River Experimental Watersheds (KREW), an integrated ecosystem project for long-term research on nested headwater streams in the Southern Sierra Nevada. KREW is also the site of the Southern Sierra Critical Zone Observatory (CZO). Snow at lower elevations exhibits multiple accumulation and melt cycles throughout the cold season, with soil moisture response similar across events and locations. Snow at higher elevations exhibits a single main melt period in spring, with streamflow lagging that at lower elevations. Soil moisture declines rapidly in the first week after snowmelt is completed, followed by a more-gradual decline thereafter. Local differences in the timing of snowmelt and soil drying between north versus south aspects and shaded versus open sites are both about one month, comparable to elevation differences in the average response. Interannual variability in timing is also similar. Evapotranspiration follows snowmelt and temperature patterns. Ionic concentrations are consistently higher in the rain-dominated sites. Two aspects of the elevational gradients offer lessons for how catchments will respond to climate warming, precipitation and snowmelt patterns. Soil moisture response to climate change is also indicated by aspect differences. Interannual differences provide indications of how geochemical fluxes in streamwater will respond to warming. Expectations and reality for high latitude versus high elevation global change Bunn, A G (andy.bunn@wwu.edu), Environmenal Sciences, Western Washington University, Bellingham, WA, United States Lloyd, A H (lloyd@middlebury.edu), Biology, Middlebury College, Middlebury, VT, United States Arctic and alpine ecosystems are often treated as analogs of each other, in large part because they share a similar vegetation transition from forested to low-stature tundra communities. Despite the superficial similarities, the response of the two types of ecosystems to future climate change will likely differ because of differences in ecosystem history, function, and extent. The role of feedbacks differs substantially between the two as the Arctic terrestrial system is dominated by feedbacks which have the potential to significantly alter the rate and magnitude of future climate change. If invoked, these feedbacks will substantially alter and augment northern high latitude change far above the background forcing from increased greenhouse gas concentrations. The same is not obviously true for mountains, both because of the difference in areal extent and because of differences in soil characteristics that affect the potential for carbon cycle feedbacks. The climatic controls over biophysical processes may differ in subtle but important ways between the two systems despite the overriding importance of temperature as a control in both ecosystems. For example, changes in the position of the treeline ecotone in the Sierra Nevada during the late Holocene occurred in response to variation in both temperature and moisture, whereas treeline advance and retreat in Arctic regions appears to be primarily a function of temperature. Despite those differences, it appears likely that changes in Arctic and alpine ecosystems will have large influences on the global system. The consequences of changes in alpine ecosystems will be amplified by their large importance in controlling global water supplies. More than 50% of the world’s freshwater supplies, for example, are derived from mountainous regions. Any change to those regions might have disastrous effects on human welfare. Global impacts of changes in Arctic regions are amplified by the aforementioned feedbacks on the climate system, which have the potential to increase the rate of warming in high latitudes by several fold, with cascading effects on the global climate system. We will review some of the similarities and differences in arctic and alpine systems by showing data on predicted changes to the physical, floral, and faunal aspects of both systems paying particular attention to the role of feedbacks and forcings. The Persistent Life of Snow Hiemstra, C A (hiemstra@cira.colostate.edu), Cooperative Institute for Research in the Atmosphere, Colorado State University, Fort Collins, CO, United States Liston, G E (liston@cira.colostate.edu), Cooperative Institute for Research in the Atmosphere, Colorado State University, Fort Collins, CO, United States Elder, K (kelder@fs.fed.us), Rocky Mountain Research Station, USDA Forest Service, Fort Collins, CO, United States Sturm, M (matthew.sturm@usace.army.mil), Cold Regions Research and Engineering Laboratory, US Army Corps of Engineers, Fairbanks, AK, United States Snow is an essential element linking mountains and poles. In high-elevation and high-latitude environments, snow is the dominant precipitation form, and observations suggest snowpacks in both these areas are being altered with climate change directly (higher temperatures) and indirectly (vegetation change). Snow’s substantial control on energy balance, water resources, and ecosystem processes make it a key variable in understanding climate change ramifications in both mountain and polar systems. In spite of its broad importance, snow remains difficult to accurately quantify on many landscapes and over a wide range of spatial and temporal scales. Because of its interactions with the atmosphere and surrounding landscape, snow is inherently dynamic and a challenge to measure and model. In both mountainous and Arctic domains, it is often transported by wind and interacts with topography and vegetation to form a heterogeneous distribution in both space and time. This heterogeneous distribution imparts numerous effects on ecosystem structure and function and land-atmosphere surface fluxes; it also obfuscates analyses of long-term trends. Both middle latitude mountains and the Arctic are experiencing changes in vegetation, precipitation, and air temperature. These accompany attendant changes in the timing and spatial distributions of snow properties, characteristics, and quantities. We will describe tools, techniques, challenges, and outcomes of measuring and modeling snow accumulation and ablation in snowy environments ranging from low to high elevations and middle to high northern latitudes, with a particular focus on the common snow-related impacts of climate and vegetation changes in these two environments. We will look at middleand high-latitude snow-vegetation interactions within shrubland environments and present improved ways to represent snow-atmosphere interactions within these landscapes. We will describe the potential ramifications of widespread bark beetle outbreaks in mountainous environments; Arctic snow trends; and how emerging Arctic vegetation and sea-ice changes are modifying terrestrial snow covers and their subsequent influences on atmospheric, hydrologic, and ecosystem processes. Interactions of climate, vegetation, and fire during the Holocene: lessons from high-latitude and high-elevation ecosystems Higuera, P E (phiguera@uidaho.edu), Forest Resources, University of Idaho, Moscow, ID, United States Chipman, M L (mchipman@life.illinois.edu), Plant Biology, University of Illinois, Urbana, IL, United States Allen, J (Jennifer_allen@nps.gov), Regional Fire Ecologist, National Park Service, Fairbanks, AK, United States Brubaker, L (lbru@u.washington.edu), Forest Resources, University of Washington, Seattle, WA, United States Whitlock, C L (whitlock@montana.edu), Earth Sciences, Montana State University, Bozeman, MT, United States Hu, F (fshu@life.illinois.edu), Plant Biology, University of Illinois, Urbana, IL, United States Recent climate change in high-latitude and high-elevation ecosystems has led to significant increases in the frequency of large, severe wildfires, highlighting the need to understand the biophysical controls of fire regimes across multiple temporal scales. Paleoecological records reveal the interactive impacts of climate and vegetation on fire regimes, and they provide a context for evaluating the consequences of ongoing and future climate change. Here we highlight new findings from high-resolution fire-history records from lake sediments in arctic ecosystems of Alaska and subalpine forests of the Rocky Mountains. Post-glacial climate change (last 15 ky) in each region influenced fire regimes directly, by altering summer temperatures and moisture patterns, and indirectly, by promoting large-scale shifts in vegetation. In the Alaskan Arctic, vegetation change strongly mediated the impacts of millennial-scale climate change on fire regimes, by changing the quantity and quality of fuels on the landscape. For example, fire frequencies decreased in the south-central Brooks Range with the development of deciduous woodlands ca. 10 ka BP, despite evidence of warmer, drier summers than at present. In the U.S. Rocky Mountains, Holocene vegetation change was less dramatic, and fire-history records from subalpine forests underwent more subtle changes. Nonetheless, the direct and indirect influence of climate is evident on a range of time scales in Yellowstone and Rocky Mountain national parks. Area burned over the past 750 years in Yellowstone, for example, was significantly above average during decades with extreme drought. Overall, both systems display evidence of strong climate-fire relationships at multiple time scales, but large-scale shifts in vegetation have had a greater impact on fire regimes in Alaska than in the Rocky Mountains. Relationships of Periglacial Processes to Habitat Quality and Thermal Environment of Pikas (Lagomorpha, Ochotona) in Alpine and High-Latitude Environments Millar, C I (cmillar@fs.fed.us), Sierra Nevada Research Center, USDA Forest Service, Albany, CA, United States Smith, A T (a.smith@asu.edu), School of Life Sciences, Arizona State University, Tempe, AZ, United States Hik, D S (david.hik@ualberta.ca), Department of Biological Sciences, University of Alberta, Edmonton, AB, Canada Patterned-ground and related periglacial features such as rock-glaciers and fractured-rock talus are emblematic of cold and dry arctic environments. The freeze-thaw processes that cause these features were first systematically investigated in the pioneering work of Linc Washburn. Unusual internal and autonomous micro-climatic and hydrologic processes of these features, however, are only beginning to be understood. Such features occur also in temperate latitude mountains, often in surprising abundance in regions such as the Great Basin (NV, USA) and San Juan Mtns (CO, USA), where they occur as active as well as relict (neoglacial or Pleistocene) features. Rock-dwelling species of pikas (/Ochotona/) in temperate North American and Asian mountains and in North American high-latitudes have long been known for their preference for talus habitats. We are investigating geomorphic, climatic, and hydrologic attributes of these periglacial features for their role in habitat quality and thermal environment of pikas. PRISM-modeled and observed climatic conditions from a range of talus types for /Ochotona princeps/ in California and the western Great Basin (USA) indicate that, 1) thermal conditions of intra-talus-matrix in summer are significantly colder than talus-surface temperatures and colder than adjacent slopes and forefield wetlands where pika forage; 2) near-talus-surface locations (where haypiles are situated) are warmer in winter than intra-talus-matrix temperatures; 3) high-quality wetland vegetation in talus forefields is promoted by year-round persistence of outlet springs, seeps, and streams characteristic of active taluses. The importance of snowpack to winter thermal conditions is highlighted from these observations, suggesting a greater sensitivity of habitat in dry temperate regions such as eastern California and Nevada USA to warming winter minimum temperatures than in regions or elevations where snowpacks are more persistent. In regions where warming air temperatures cause declining snowpacks over talus in the future, pika winter habitats may be exposed not only to colder surface- air temperatures, but to sub-freezing air circulation from within talus matrices. An important implication is that periglacial features in which pikas inhabit have different thermal environments and unique internal air circulation compared to slopes and ambient surface air above taluses. This suggests that talus environments might have micro-climatic conditions in the future quite different from those modeled for the general mountain or arctic contexts in which they occur. We bring together new observations from North American temperate mountain- and high-latitude pika species, and from rock- and soil-dwelling species in the highlands of Asia, to speculate on population ecology in the face of changing climates in these disparate regions. Managing for Climate Change in Western Forest Ecosystems; The Role of Refugia in Adaptation Strategies Millar, C I (cmillar@fs.fed.us), Sierra Nevada Research Center, USDA Forest Service, Albany, CA, United States, Morelli, T (tmorelli@fs.fed.us), Sierra Nevada Research Center, USDA Forest Service, Albany, CA, United States Managing forested ecosystems in western North America for adaptation to climate change involves options that depend on resource objectives, landscape conditions, sensitivity to change, and social desires. Strategies range from preserving species and ecosystems in the face of change (resisting change); managing for resilience to change; realigning ecosystems that have been severely altered so that they can adapt successfully; and enabling species to respond to climate changes. We are exploring one extreme in this range of strategies, that is, to manage locations, species, communities, or ecosystems as refugia. This concept is familiar from the Quaternary literature as isolated locations where climates remained warm during cold glacial intervals and wherein species contracted and persisted in small populations. References to refugia have been made in the climate-adaptation literature but little elaborated, and applications have not been described. We are addressing this gap conceptually and in case-studies from national forest and national park environments in California. Using a classification of refugium categories, we extend the concept beyond the original use to include diverse locations and conditions where plant or animal species, or ecosystems of concern, would persist during future changing climatic backgrounds. These locations may be determined as refugial for reasons of local microclimate, substrate, elevation, topographic context, paleohistory, species ecology, or management capacity. Recognizing that species and ecosystems respond to climate change differently, refugium strategies are appropriate in some situations and not others. We describe favorable conditions for using refugium strategies and elaborate specific approaches in Sierra Nevada case studies. Patterns of cold-air drainage and microclimate in mid-latitude versus high-latitude mountains: contrasts and implications for climate change Pepin, N C (nicholas.pepin@port.ac.uk), Geography, University of Portsmouth, Portsmouth, United Kingdom Predictions of current spatial patterns of climate are difficult in areas of complex relief in all parts of the world, because of the interweaving influences of topography, elevation and aspect. These influences vary temporally as a result of the seasonal and diurnal cycles in radiation balance. In periods of negative energy balance, surface decoupling can occur as cold air drainage develops low-level temperature inversions, and the surface temperature regime beneath the inversion becomes divorced from free atmospheric forcing. Both the spatial scale and temporal persistence of this decoupling vary according to latitude, and although the physical processes that influence inversion formation are similar in polar areas and mid-latitude mountains, the contrasting seasonal and diurnal forcings make the end results very different. Examples are contrasted from detailed field temperature measurements (~50 sites per field area) taken over several years in areas of complex relief in the eastern Pyrenees (~42.5 deg N), the Oregon Cascades (also ~42.5 deg N) and Finnish Lapland (70 deg N and above the Arctic circle). In the former two locations decoupling is mostly diurnally driven, and small-scale topography is important in mediating the effects. Summer decoupling is brief and spatially limited, whereas winter decoupling can be more spatially extensive. There are strong relationships between synoptic conditions, as measured by objective flow indices at the 700 mb level (derived from NCEP/NCAR reanalysis fields) and the patterns of decoupling, which allow us to assess the effects of past and potential future circulation change on spatial patterns of future climate warming. In Finnish Lapland the decoupling regime most clearly approaches the mid-latitude pattern around the equinoxes when there are clear day and night periods. In winter and summer however (the polar night and polar day) with the muting of the diurnal cycle, processes are more poorly understood. Winter cold pools can develop and strengthen over days until eventually they extend over and above the topography. Strangely, there are also indistinct relationships with circulation indices at this time. While build-up can take days, destruction is often immediate and is dynamically forced. In summer, localized decoupling occurs on clear nights even though the sun is above the horizon, but micro-scale patterns are different than in mid-latitudes. The above comparison shows that polar areas are very different in their microtemperature regimes than mid-latitude mountains and in their relationships of these regimes with circulation. Thus we expect detailed spatial patterns of climate change may be very different in the two regions. Potential effects of Rising CO2 and Climate Change Interactions with Atmospheric Deposition in Northeastern U.S. over the 21st Century Using a Dynamic Biogeochemical Model (PnET-BGC) Pourmokhtarian, A (apourmok@syr.edu), Civil & Environmental Engineering, Syracuse University, Syracuse, NY, United States Driscoll, C T (ctdrisco@syr.edu), Civil & Environmental Engineering, Syracuse University, Syracuse, NY, United States Campbell, J L (jlcampbell@fs.fed.us), Northern Research Station, US Forest Service, Durham, NH, United States Hayhoe, K (katharine.hayhoe@ttu.edu), Geosciences, Texas Tech University, Lubbock, TX, United States Climate is an important regulator of hydrology and biogeochemical processes within forest ecosystems, therefore the functionality of terrestrial ecosystems is altered by changes in climate. The ecological responses to climate change have been assessed by observational, gradient, laboratory and field studies; however, models are the only practical approach that can predict concurrent exposure to multiple environmental factors and consider interactive effects between climate change, atmospheric deposition and CO2 fertilization effects. Therefore, dynamic biogeochemical watershed models are potentially powerful tools to help to understand the long-term effects of climate change on ecosystems. In this study, we use a biogeochemical model (PnET-BGC) coupled with long-term measurements to evaluate the effects of potential future changes in temperature, precipitation, solar radiation, atmospheric deposition and atmospheric CO2 on pools and fluxes of major elements at the Hubbard Brook Experimental Forest (HBEF) in New Hampshire. Future emissions scenarios were developed from monthly output from two atmosphere-ocean general circulation models (AOGCMs; HadCM3, PCM) in conjunction with potential lower and upper bounds of projected atmospheric CO2 (550 and 970 ppm by 2099, respectively). We also evaluated the interaction of climate change with changes in acidic deposition. Estimates of atmospheric deposition are based on a “business-as-usual” deposition scenario. We compared the results of the “business as usual” scenario with two additional scenarios which consider additional moderate and aggressive emission controls on sulfate and nitrate. AOGCM results over the 21st century indicate an average increase in temperature ranging from 1.9 to 6.9°C with simultaneous increases in precipitation ranging from 12.5 to 13.9% above the long term mean (1970-1999). Long-term measurements and watershed modeling results show a significant shift in hydrology with earlier spring discharge (snowmelt), greater evapotranspiration and longer growing season (due to CO2 fertilization), and later snowpack development. Model results also show an increase in NO3- leaching over the second half of the century due to increases in net mineralization and nitrification. The extent of this response is dependent on the fertilization effect that increasing atmospheric CO2 has on forest vegetation. The watershed responses of other major elements such as SO42and Ca2+, and chemical characteristics such as pH and ANC varied based on future climate and deposition scenarios. Model predictions showed that equivalent decreases in sulfate deposition were twice as effective as decreases in nitrate on recovery of soil and streamwater chemistry. Model projections also suggest marked decreases in soil exchangeable calcium, magnesium and potassium with simultaneous decline in soil base saturation and Ca/Al ratio over the next century. Hydrochemical responses to climate change in high-elevation catchments of the Colorado Front Range. Williams, M W (markw@snobear.colorado.edu), University of Colorado, Boulder, CO, United States Potential climate impacts on the hydrochemistry of two seasonally snow covered catchments is evaluated using 24 years of data from the Niwot Ridge Long Term Ecological Research Site, Colorado. At the larger (220 ha), higher elevation (3570 m) GL4 catchment annual discharge did not change significantly based on nonparametric trend testing. However, October streaflow volumes and groundwater storage did increase, despite drought conditions near the end of the record in 2000-2004. In contrast, at the smaller (8 ha), lower elevation (3400 m) MART catchment, annual discharge decreased significantly over the study period with the most substantial changes in July-September. The study period was separated into "wet", "normal", and "dry" years based on the 75th and 25th quartiles of annual precipitation. Results indicate that MART is particularly sensitive to changes in precipitation with dry years exhibiting decreased snowmelt peak flows, earlier snowmelt timing, decreased annual discharge, and reduced late-season flows. GL4 was less susceptible to changes in precipitation and surprisingly late-season flow volumes (Sept.-Oct.) were not significantly different between wet, normal, and dry conditions. Surprisingly, during dry years both the concentrations and annual fluxes of Ca2+ and SO42- increased in the outflow of GL4, but not at the Martinelli catchment. These changes in hydrochemistry were particularly pronounced during the low-flow period. Streamwater chemistry in GL4 during drought years resembled that of permafrost, suggesting augmented flow during the fall due to permafrost melt. This study shows that seasonally snow covered catchments are particularly sensitive to changes in climate, but the hydrochemical response may depend on landscape characteristics. POSTER SESSION Recent increase in maximum temperature at the tropical treeline of North America Biondi, F (fbiondi@unr.edu), University of Nevada, Reno, NV, United States There are only a handful of weather stations above 3000 m in the entire American Cordillera, from Alaska to Tierra del Fuego. I present a surface instrumental record of high elevation (treeline) ecoclimatic variables for the tropics of North America. Besides its high elevation (3760 m) and tropical (19.5°N) features, this site is also located in the North American Monsoon System, making the data relevant to a broad suite of environmental issues. Automated half-hour data collected on Nevado de Colima, Mexico, from 2001 to 2009 show an increase in maximum temperature during the dry winter season, while incoming solar radiation remained stationary. Since minimum temperature did not increase as much, the daily range of air temperature has expanded over time. At this elevation, with average daily barometric pressure of 655 ± 1.4 hPa, maximum temperatures reflect the annual and daily energy cycle because of the dominant role of ground heating caused by incoming shortwave radiation. In fact, spring is the warmest season in this area, as it is followed by pronounced cooling during the summer monsoon because of increased cloudiness. The observed warming is associated with reduced wind speed, especially around solar noon, and is therefore most likely driven by reduced atmospheric flow, suggesting that the energy and water balance of high elevation tropical ecosystems are changing in unexpected ways. Further measurements and regional modeling experiments are therefore needed, given the staggering consequences this could have for any resource managers and policy makers concerned with trans-boundary (Mexico-US) terrestrial, coastal, and oceanic issues. Small mammals and high elevation vegetation in Yosemite National Park could be responding to smaller temperature increases than previously reported Conklin, D R (david.conklin@oregonstate.edu), Oregon State University, Corvallis, OR, United States Daly, C (daly@nacse.org), Oregon State University, Corvallis, OR, United States Recent research related the dynamics of Sierra Nevada subalpine conifers to 20th century warming, and used temperature records from Yosemite National Park headquarters and two other locations outside the Park to estimate a 3.7°C rise in average minimum temperature. More recently, observed elevational shifts of small mammals in Yosemite National Park were linked to the same upward trend in minimum monthly temperatures. However, our analysis of spatially explicit, monthly time series of temperatures, derived using the PRISM model, suggests that the large increase in minimum monthly temperatures may be limited to Yosemite Valley, where the Park headquarters itself is located. PRISM bases its interpolations on observations from 195 temperature recording stations within and near Yosemite National Park. Minimum monthly temperatures over most of the Park do not show a centennial-scale trend, but for the final quarter of the century they do trend upwards by 1°C. Our new estimate of the spatial and temporal pattern of 20th century changes in minimum temperatures in the Park could affect conclusions about the relative importance for subalpine conifers of the centennial trend compared to interdecadal variability of temperature. It also raises a question of whether the elevational shifts of the mammals took place only in the latter part of the century, and in response to smaller temperature increases. It challenges us to accept that these plants and animals are responding to smaller changes in minimum temperatures than previously estimated or else to find reasons other than an increase in minimum temperatures for the changes that have been documented, by these and other studies, over the last century in Yosemite. Using an Inverse Geochemical Reaction Path Model to Analyze the Effects of Climate Change on High-elevation Catchments in the Southern Rocky Mountains Driscoll, J M (driscoll@hwr.arizona.edu), Hydrology and Water Resources, University of Arizona, Tucson, AZ, United States Meixner, T (tmeixner@hwr.arizona.edu), Hydrology and Water Resources, University of Arizona, Tucson, AZ, United States Brooks, P D (brooks@hwr.arizona.edu), Hydrology and Water Resources, University of Arizona, Tucson, AZ, United States McIntosh, J C (mcintosh@hwr.arizona.edu), Hydrology and Water Resources, University of Arizona, Tucson, AZ, United States Williams, M W (markw@snobear.colorado.edu), Institute of Arctic and Alpine Research, University of Colorado, Boulder, CO, United States Molotch, N P (noah.molotch@colorado.edu), Institute of Arctic and Alpine Research, University of Colorado, Boulder, CO, United States Sickman, J O (jsickman@ucr.edu), Environmental Studies, University of California, Riverside, CA, United States High-elevation catchments are particularly responsive to changes in climate, which alters the snowmelt dynamics that drive geochemical reactions in these sensitive ecosystems. Snowmelt timing and duration can be altered by the amount of snowpack available for melt as well as the rate at which melt occurs. This study aims to assess the impact of changes to these snowmelt dynamics through the use of a reaction path model (RPM). The inverse geochemical RPM approach within PHREEQC was used to find estimates of mineral weathering rates and stoichiometry and infer the hydrologic structure of the catchments studied. Chemical and hydrologic data from two NSF Critical Zone Observatories, Green Lake 4 (GL4) in the Boulder Creek watershed in Colorado and the Valles Caldera (Valles) at Redondo Peak in New Mexico were chosen to evaluate the impact of physical differences in snowmelt dynamics on weathering and hydrologic connectivity. The influence of total snow amount on weathering and connectivity was evaluated using wet (1996) and dry (2002) winters in GL4, the impact of the rate of snowmelt was evaluated by assessing a variety of aspect directions (around Redondo Peak) in the Valles, and a comparison of the relative latitude of these catchments was also used to determine the amplitude of effects from south to north in the Rocky Mountains. We hypothesize that increased snowmelt rates due to variations in aspect will lead to differences in catchment residence times, and that longer residence times will lead to more contact time between water and weathering minerals and therefore produce more weathering products. The amount of mineral weathering within a catchment is therefore indicative of relative residence time; however, when comparing catchments, assumptions regarding bedrock lithology, temperature and other chemical inputs are necessary. The geochemical connectivity of waters in the catchments is dependent on the amount of snowmelt and transit time, with the number of connections hypothesized to be fewer with less snowpack. Further analysis of the changes in the specific mineral weathering reactions that occur in these conditions will lead to conclusions on the impact of changing snowmelt dynamics due to climate change on the buffering capacity of alpine catchments. Multi-Scale Influences of Climate, Spatial Pattern, and Positive Feedback on 20th Century Tree Establishment at Upper Treeline in the Rocky Mountains, USA Elliott, G P (elliottg@missouri.edu), Geography, University of Missouri, Columbia, MO, United States The influences of 20th century climate, spatial pattern of tree establishment, and positive feedback were assessed to gain a more holistic understanding of how broad scale abiotic and local scale biotic components interact to govern upper treeline ecotonal dynamics along a latitudinal gradient (ca. 35°N-45°N) in the Rocky Mountains. Study sites (n = 22) were in the Bighorn, Medicine Bow, Front Range, and Sangre de Cristo mountain ranges. Dendroecological techniques were used for a broad scale analysis of climate at treeline. Five-year age-structure classes were compared with identical fiveyear bins of 20th century climate data using Spearman’s rank correlation and regime shift analysis. Local scale biotic interactions capable of ameliorating broad scale climate inputs through positive feedback were examined by using Ripley’s K to determine the spatial patterns of tree establishment above timberline. Significant correlations (p < 0.01) between tree establishment and climate were confined to the Front Range, where a positive correlation exists with summer (June-Aug) and cool season (Nov-Apr) temperature range (Tmax-Tmin). Additionally, trees in the Front Range are almost exclusively situated in a random spatial pattern above timberline (4/5 sites). Random spatial patterns imply that positive feedback is of minimal importance and that trees are more closely aligned with broad scale changes in abiotic conditions. This tight coupling between climate and treeline vegetation in the Front Range helps explain synchronous ecological (tree establishment) and climate regime shifts (temperature) during the early 1950s. Similar to the Front Range, a majority of trees at upper treeline in the Bighorn Mountains are in a random spatial pattern, but their existence appears to be dependent on shelter availability in the lee of boulders. This contingency helps explain the lag time between a regime shift to more favorable temperatures and subsequent peaks in tree establishment. The Medicine Bow and Sangre de Cristo Mountains primarily contain clustered spatial patterns of trees above timberline, which indicates a strong reliance on the amelioration of abiotic conditions through positive feedback with nearby vegetation. Although clustered spatial patterns likely originate in response to harsh abiotic conditions such as drought or constant strong winds, the local scale biotic interactions within a clustered formation of trees appears to override the immediate influence of broad scale climate. This is evidenced both by a lack of significant correlations between tree establishment and climate in these mountain ranges, as well as the considerable lag times between initial climate regime shifts and corresponding shifts in tree age structure. Taken together, this research suggests that the influence of broad scale climate on upper treeline ecotonal dynamics is contingent on the local scale spatial patterns of tree establishment and related influences of positive feedback. These findings have global implications for our understanding of how vegetation patterns will respond to various global climate change scenarios. Plant Water Use Efficiency Response to the Atmospheric CO2 Concentration is Greater in High Altitude Environments Feng, X (xiahong.feng@dartmouth.edu), Earth Sciences, Dartmouth College, Hanover, NH, United States Wang, G (gawang@cau.edu.cn), Resources and Environment, China Agricultural University, Beijing, China Intrinsic water-use efficiency of plants (A/g ratio, where A is CO2 assimilation rate and g is stomatal conductance of H2O) quantifies the amount of carbon assimilated per unit leaf area per unit time per unit cost of water. There has been a large body of work showing that intrinsic plant water use efficiency (WUE) increases with increasing atmospheric CO2 concentration. This conclusion has strong implications for quantifying the effects of terrestrial carbon sequestration and plant transpiration under the condition of continuously increasing anthropogenic CO2. Less attention has been given to assessing whether the plant response to the atmospheric CO2 increase differs as a function of environmental variables, such as temperature, precipitation and altitude. One would expect interactions between the CO2 concentration and other environmental variables, and the joint effects on the plant WUE might be different from the effect of CO2 concentration alone. However, these interactions can be quite complicated and difficult to predict; even the sign of response remains uncertain. For example, one would expect that plants growing under a dry climate may benefit from the CO2 increase more than those under wet climate, and thus A would increase more in a dry climate. Stomata density of leaves typically decreases with increasing CO2 concentration, causing g to decrease, and A/g to increase. However, it is not known if stomata density decreases more or less under dry or wet climate conditions. Similar uncertainties or lack of knowledge apply to temperature effects. In this work, we adopt an empirical approach using carbon isotopic ratios in tree rings. Over 50 tree-ring δ13C series are compiled from the literature. The response of δ13C to atmospheric conditions (CO2 concentration and δ13C) is obtained, and the rates of change of the WUE are obtained at several different times between AD 1800 and 2000. These rates are then compared statistically with location’s mean annual temperature, annual precipitation and altitude, in addition to the rate of change in the atmospheric CO2 concentration. The multiple linear regression results show that the majority of the WUE increase is explained by the change in CO2 concentration, and the magnitude of the CO2 effect is close to the theoretically predicted value. The rate of change of WUE has no significant response to either temperature or precipitation. However, there is a significant positive contribution of altitude to the rate of increase of WUE; i.e., the WUE increases faster in high altitude than in low altitude locations. This is probably caused by the fact that high altitude plants are CO2 starved due to low CO2 partial pressure. It is possible that an increase in CO2 concentration induces a greater boost in A in high altitude plants relative to low altitude ones. Climate Change Responses of Hydrologic Flowpaths in Mountainous and Polar Regions Godsey, S (godseys@eps.berkeley.edu), UC-Berkeley, Berkeley, CA, United States Gooseff, M N (mgooseff@engr.psu.edu), Pennsylvania State University, State College, PA, United States Kirchner, J W (james.kirchner@wsl.ch), Swiss Federal Institute for Forest, Snow, and Landscape Research (WSL), Birmensdorf, Switzerland Tague, C (ctague@bren.ucsb.edu), UC-Santa Barbara, Santa Barbara, CA, United States Hydrologic processes in mountainous and polar regions may respond differently to changes in the catchment energy budget that are anticipated to occur as climate changes. In the Sierra Nevada Mountains of California, warmer winter temperatures are expected to shift the phase of precipitation from snow to rain across a range of elevations. We examine whether this phase change will alter subsequent low flow regimes during the dry Mediterranean summers of this region. We show that changes in the phase of precipitation as well as changes in evapotranspiration losses from vegetation are key drivers in the hydrologic response of these mountains to climate change. In northern Alaska, the depth of the active layer above permafrost evolves over space and time, affecting subsurface flowpaths. Large changes in water or energy flows may lead to catastrophic loss of ground ice, known as thermokarst development. Thermokarst features can deliver large pulses of sediment and nutrients to lakes and streams, and further alter the hydrology because they expose previously insulated permafrost to the ground surface, and thus higher heat fluxes. Here we show the spatial and temporal development of the active layer inside of and outside of thermokarst features over the course of the warming season. We explore the importance of changes in subsurface topography as a driver of hydrologic response to climate change in arctic tundra catchments. Biotic Responses of Headwater Streams to Geophysical Alteration and Disturbance Related to Climate Change Gresswell, R E (bgresswell@usgs.gov), US Geological Survey, Northern Rocky Mountain Science Center, Bozeman, MT, United States Sedell, E R (tsedell@gmail.com), Department of Ecology, Montana State University, Bozeman, MT, United States Cannon, S (cannon@usgs.gov), US Geological Survey Landslide Hazards Program, Denver, CO, United States Hostetler, S W (steve@coas.oregonstate.edu), US Geological Survey, National Research Program, Water Resources Center, Corvallis, OR, United States Williams, J E (JWilliams@tu.org), Trout Unlimited, Ashland, OR, United States Haak, A L (ahaak@tu.org), Trout Unlimited, Boise, ID, United States Kershner, J L (jkershner@usgs.go), US Geological Survey, Northern Rocky Mountain Science Center, Bozeman, MT, United States Climate change will potentially alter physical habitat availability for trout species (both native and nonnative) in the western USA, and ultimately affect population distribution and abundance in watersheds across the region. To understand the biological consequences of habitat alteration associated with climate change, we have developed models linking contemporary patterns of occurrence and abundance to geomorphic variables (e.g., aspect, elevation, and slope) and stream conditions derived from the habitat (e.g., temperature, discharge, and flood regimes). Because headwater streams may be especially susceptible to catastrophic disturbances in the form of debris flow torrents that have the potential to radically alter the physical structure of channels and sometimes extirpate local fish populations, we are focusing fine-scale spatial analyses in the high elevation systems. Risks of such disturbances increase exponentially in landscapes that have experienced recent wildfires when high-intensity precipitation or runoff events occur. Although predicting the timing, extent, and severity of future wildfires or subsequent precipitation and runoff events is difficult, it is possible to identify channels within stream networks that may be prone to debris flows. These channels can be identified using models based on characteristic storm and burn scenarios and geographic information describing topographic, soil, and vegetation characteristics. At-risk channels are being mapped throughout the stream networks within the study areas in the headwaters of the Colorado River to provide information about the potential for catastrophic population disturbance in response to variety of wildfire and post-wildfire storm scenarios. Explaining the Spatial Variability in Stream Acid Buffering Chemistry and Aquatic Biota in the Neversink River Watershed, Catskill Mountains, New York State Harpold, A A (aah38@cornell.edu), Biological & Environmental Engineering, Cornell University, Federal Way, NY, United States Walter, M T (mtw5@cornell.edu), Biological & Environmental Engineering, Cornell University, Federal Way, NY, United States The Neversink River Watershed (NRW) originates at the highest point in the Catskill Mountains and is sensitive to changing patterns in acidic deposition, precipitation, and air temperature. Despite reductions in fossil fuel emission since the Clean Air Act, past acidic deposition has accelerated the leaching of cations from the soil and reduced the stores of base cations necessary for buffering stream acidity. The goal of this study was to investigate connections between different watershed ‘features’ and the apparently complex spatial patterns of stream buffering chemistry (specifically, acid neutralizing capacity ANC and Ca concentrations) and aquatic biota (macroinvertebrate and fish populations). The ten nested NRW watersheds (2.0 km^2 to 176.0 km^2) have relatively homogeneous bedrock geology, forested cover, and soil series; therefore, we hypothesized that differing distributions of hydrological flowpaths between the watersheds control the variability in stream buffering chemistry and aquatic biota. However because the flowpath distributions are not directly measurable, this study used step-wise linear regression to develop relationships between watershed ‘features’ and buffering chemistry. The regression results showed that the mean ratio of precipitation to stream runoff (or runoff ratio) from twenty non-winter storm events explained more than 81% of the variability in mean summer ANC and Ca concentrations. The results also suggested that steeper (higher mean slope) more channelized watersheds (larger drainage density) are more susceptible to stream acidity and negative impacts on biota. A simple linear relationship (using no discharge or water chemistry measurements) was able to explain buffering chemistry and aquatic biota populations in 17 additional NRW watersheds (0.3 km^2 to 160.0 km^2), including 60-80% of the variability in macroinvertebrate populations (EPT richness and BAP) and 50-60% of the variability in fish density and species richness. These results have several important implications for understanding the effects of climate change on buffering chemistry and aquatic biota in this well-studied watershed. First, the results demonstrate that geomorphological and hydrogeological ‘features’ control the spatial variability of stream buffering chemistry, suggesting that acidification ‘hot-spots’ could be predicted a priori. Second, the connection between event-scale processes (runoff ratio) and average stream chemistry imply that changing precipitation patterns in the Catskills may have uneven effects on long-term buffering chemistry between ‘flashy’ and ‘damped’ watersheds. Specifically, an increasing trend in precipitation in the last 25 years in the Catskill Mountains makes it difficult to compare base cation recovery across NRW streams, even if the concentrations are normalized by discharge. The results of this study could improve the modeling of base cation recovery in surface waters in other mountainous Northeastern U.S. watersheds with future reductions in acidic deposition and differing climate scenarios. Predicting Individual Tree and Shrub Species Distributions with Empirically Derived Microclimate Surfaces in a Complex Mountain Ecosystem in Northern Idaho, USA Holden, Z (zaholden@fs.fed.us), U.S. Forest Service, Missoula, MT, United States Cushman, S (scushman@fs.fed.us), USFS Rocky Mountain Research Station, Missoula, MT, United States Evans, J (jeffrey_evans@tnc.org), The Nature Conservancy, Ft. Collins, CO, United States Littell, J S (jlittell@u.washington.edu), CSES Climate Impacts group, University of Washington, Seattle, WA, United States The resolution of current climate interpolation models limits our ability to adequately account for temperature variability in complex mountainous terrain. We empirically derive 30 meter resolution models of June-October day and nighttime temperature and April nighttime Vapor Pressure Deficit (VPD) using hourly data from 53 Hobo dataloggers stratified by topographic setting in mixed conifer forests near Bonners Ferry, ID. 66%, of the variability in average June-October daytime temperature is explained by 3 variables (elevation, relative slope position and topographic roughness) derived from 30 meter digital elevation models. 69% of the variability in nighttime temperatures among stations is explained by elevation, relative slope position and topographic dissection (450 meter window). 54% of variability in April nighttime VPD is explained by elevation, soil wetness and the NDVIc derived from Landsat. We extract temperature and VPD predictions at 411 intensified Forest Inventory and Analysis plots (FIA). We use these variables with soil wetness and solar radiation indices derived from a 30 meter DEM to predict the presence and absence of 10 common forest tree species and 25 shrub species. Classification accuracies range from 87% for Pinus ponderosa , to > 97% for most other tree species. Shrub model accuracies are also high with greater than 90% accuracy for the majority of species. Species distribution models based on the physical variables that drive species occurrence, rather than their topographic surrogates, will eventually allow us to predict potential future distributions of these species with warming climate at fine spatial scales. Phosphorous in the Sierra Nevada: Forms, mechanisms, and timing of release in high-elevation soils Homyak, P M (Peter.homyak@email.ucr.edu), Environmental Sciences, University of California, Riverside, Riverside, CA, United States Sickman, J O (jsickman@ucr.edu), Environmental Sciences, University of California, Riverside, Riverside, CA, United States Melack, J M (melack@lifesci.ucsb.edu), Ecology, Evolution & Marine Biology, University of California, Santa Barbara, Santa Barbara, CA, United States In high-elevation lakes of the Sierra Nevada (California) a change in nutrient loading has resulted in mild eutrophication with concomitant shifts from P to N limitation, but the source of P is currently unknown. Temperature, runoff patterns, and the timing of snowmelt influence N and P biogeochemistry in high-elevation ecosystems, which can modify cycling of P in soils and result in altered P availability. To determine whether changes in P cycling, in response to climatic changes, can lead to the mild eutrophication documented in Sierran lakes, we analyzed P pools in entisols and inceptisols in the Emerald Lake Watershed, a representative high-elevation catchment, in Sequoia National Park. Our objective is to address how P is mobilized and transformed in soils and how these processes are modified by variations in climate and hydrology. Results from sequential P fractionation extractions indicate that on average 692 µg P g^-1 of soil are available in organic soils and 547 µg P g^-1 of soil are available in mineral soils. In organic soils, 71 % of the total P is freely exchangeable or associated with Fe and Al, 19 % is Ca-associated P, and 10 % exists in recalcitrant pools. In mineral soils, 58 % of the total P is freely exchangeable or associated with Fe and Al, 32 % is associated with Ca, and 10 % exists in recalcitrant pools. Our results suggest that the majority of the total P in high-elevation soils is found in pools that can be affected by climatic and hydrologic changes. Future research will incorporate lake sediment P chemistry as well as freezethawing and drying-rewetting experiments on soils to assess microbial P turnover and the potential effect of climate change on P availability in Sierran soils. IPY-Back to the Future: Determining decadal time scale change in ecosystem structure and function in high latitude and high altitude tundra ecosystems Johnson, D R (drjohnson2@utep.edu), Systems Ecology Lab, University of Texas at El Paso, El Paso, TX, United States Villarreal, S (svillarreal3@miners.utep.edu), Systems Ecology Lab, University of Texas at El Paso, El Paso, TX, United States Lara, M (mjlara@miners.utep.edu), Systems Ecology Lab, University of Texas at El Paso, El Paso, TX, United States Webber, P J (webber@msu.edu), Department of Plant Biology, Michigan State University, East Lansing, MI, United States Callaghan, T (terry_callaghan@binternet.com), Abisko Scientific Research Station, Abisko, Sweden Hik, D (dhik@ualberta.ca), Department of Biology, University of Alberta, Edmonton, AB, Canada Tweedie, C E (ctweedie@utep.edu), Systems Ecology Lab, University of Texas at El Paso, El Paso, TX, United States In the absence of long-term monitoring, revisiting, re-sampling and assessing environmental change that has occurred at Arctic and alpine research sites established several decades ago represent a largely untapped change detection capacity. The primary objective of the “International Polar Year - Back to the Future” project, a three-year IPY project (#214), is to determine how key structural and functional characteristics of high latitude/altitude tundra ecosystems have changed over the past 25 or more years and assess if such trajectories of change are likely to continue in the future. This poster presents an update of the resampling efforts undertaken in several Arctic (Barrow, Alaska, Atqasuk, Alaska and Baffin Island, Canada) and alpine locations (Niwot Ridge, Colorado and Kluane Lake, Canada). This poster presentation will highlight several shared and contrasting patterns of change documented the IPY-BTF sites highlighted above, as well as several other IPY-BTF sites (Greenland, northern Russia, and Scandinavia), and presented at a fall 2009 IPY-BTF synthesis meeting. Climate controls on anomalously high productivity in the mixed conifer forests of the Sierra Nevada Kelly, A E (a.kelly@uci.edu), Earth System Science, University of California, Irvine, Irvine, CA, United States Goulden, M L (mgoulden@uci.edu), Earth System Science, University of California, Irvine, Irvine, CA, United States The Mediterranean climate of California’s Sierra Nevada Mountains supports a dense conifer forest that contains some of the largest trees in the world. Well-established ecological relationships, such as the Miami Model, predict relatively low NPPs for these forests (~250 g/m^2 /yr to 1300 g/m^2 /yr) due to winter cold limitation and summer drought. However, the observed rates of NPPs are quite high (up to 2000 g/m^2 /yr), raising the question of what environmental conditions and plant adaptations promote such a high NPP. We hypothesize that the trees in these forests are neither as cold-limited nor water-limited as surface weather station data suggest. Eddy covariance observations at the top of a 55 m tall micrometeorological tower located at 2050 m elevation indicate daytime CO2 uptake continues year round, and is not limited by winter cold or summer drought. Comparisons of temperature measurements on the tower with operational balloon soundings indicate that tree canopies are often in the free troposphere, which buffers the temperatures they experience and moderates winter cold limitation and summer evapotranspiration. Meteorological Controls On The Energy Balance And CO2 Over Alpine Tundra At Niwot Ridge, Colorado Knowles, J F (John.Knowles@colorado.edu), Geography, University of Colorado, Boulder, CO, United States Blanken, P (blanken@colorado.edu), Geography, University of Colorado, Boulder, CO, United States Williams, M W (markw@snobear.colorado.edu), Geography, University of Colorado, Boulder, CO, United States Ecosystems in topographically complex (mountainous) terrain are likely responsible for a majority of land-atmosphere CO2 exchange (net ecosystem exchange; NEE) across the western United States due to high inputs of winter precipitation as snowfall. NEE in these regions has been historically difficult to quantify using the eddy covariance (EC) method, however, due to complexities in surface terrain that lead to irregularities in streamline air flow, particularly advective fluxes during periods of low turbulent mixing. This research evaluated the applicability of the EC method at an alpine tundra site in conjunction with the Niwot Ridge Long Term Ecological Research Project (LTER), and subsequently investigated meteorological factors controlling the observed NEE. Persistent high wind speeds, a relatively short turbulent flux footprint, and the nearly flat ridge-top location of the study site resulted in 85% mean annual energy balance closure and general fulfillment of basic EC methodological requirements. In all, the alpine tundra was a net source of C to the atmosphere during 2008. Peak rates of summertime CO2 sequestration and wintertime CO2 respiration were of comparable magnitude. Rates of CO2 sequestration during the growing season increased non-linearly with increasing soil temperature, in agreement with results from other alpine tundra studies. CO2 sequestration was 22% less during the relatively longer 2007 growing season, highlighting a possible inverse relationship between alpine growing season length and CO2 sequestration on Niwot Ridge. Climate and 20th century establishment in alpine treeline ecotones of the Western US Littell, J S (jlittell@u.washington.edu), University of Washington, Seattle, WA, United States Pederson, G T (gpederson@usgs.gov), United States Geological Survey, Bozeman, MT, United States Graumlich, L J (lisag@cals.arizona.edu), University of Arizona, Tucson, AZ, United States Germino, M J (germmatt@isu.edu), Idaho State University, Pocatello, ID, United States Rowland, E (erowland@cals.arizona.edu), University of Arizona, Tucson, AZ, United States Climate affects the location and demography of treeline by affecting the balance between seedling establishment and mortality. In this study, we evaluated the nature of and climatic associations with conifer establishment at nine treelines in the western US. Late twentieth century pulses of establishment occurred at all nine treelines, and the location of new establishment is almost entirely within 50-100m of trees that established prior to the 19th or early 20th century. The timing, species, and magnitude of the pulses varied among treelines. At the scale of the western US, the pulses in aggregate appear to be related to increasing temperature, but detailed analysis at the level of each treeline suggests a more complicated relationship between site-level climatic and biotic factors. We describe the role of growing season heat sums, spring-summer snowpack persistence, and summer water balance within the different treelines and relate establishment both to climate and to microsite conditions. Structural and Geomorphic Controls in Altitudinal Treeline: a Case Study in the Front Ranges of the Canadian Rocky Mountains Macias Fauria, M (mmaciasf@ucalgary.ca), Biogeoscience Institute, University of Calgary, Calgary, AB, Canada Johnson, E A (johnsone@ucalgary.ca), Biogeoscience Institute, University of Calgary, Calgary, AB, Canada Altitudinal treelines occur on mountain slopes. The geological history of mountain systems sets both the distribution of slope angles, aspects and lengths, and the physical characteristics of the bedrock and regolith on which trees have to establish and grow. We show that altitudinal treeline is largely controlled at an ecosystem level by structural and slope (i.e. gravitational) geomorphic processes operating at a range of temporal and spatial scales, which have direct influence on the hydrological properties of the substrate (affecting the trees’ water and energy budget), as well as on substrate stability, both of which affect recruitment and growth of trees. The study was conducted over a relatively large area of > 200 km2 in the Front Ranges of the Canadian Rocky Mountains, selected to contain the regional diversity of slopes and substrates, which is the result of hundreds of millions of years of sea deposition, subsequent mountain building, and deep erosion by glaciations. Very high-resolution remote sensing data (LiDAR), aerial orthophotos taken at several times since the late 1940s, and ground truthing were employed to classify the terrain into process-based geomorphic units. High resolution, landscape-scale treeline studies are able avoid potential biases in site selection (i.e. selection of sites that are not representative of the overall regional treeline), and consequently capture the coupling between trees and the environment at an ecosystem (regional) level. Moreover, explicitly accounting for slope and substrate-related processes occurring in the studied mountain region is paramount in order to understand the dynamics of trees at their altitudinal distribution limit. Presence of trees in each unit was found to be controlled by a set of parameters relevant to both hydrological and slope processes, such as contributing area, slope angle, regolith transmissivity, and aspect. Our results show no treeline advance over the last 60 years in the region, as most of the area is controlled by geological processes and not by physiological temperature thresholds. Temperature could potentially affect presence of trees at high elevations through its effects on the physical properties of the slopes on which trees grow. However, this effect is at a much longer timescale than those implied in current studies of treeline response to global warming. Finally, continuous recruitment of trees following lightning-caused wildfires during the first half of the 20th century has resulted in increased high altitude forest stand density. Climate induced changes in high-elevation lake chemistry and the importance of sulfide weathering Mast, A (mamast@usgs.gov), Colorado Water Science Center, US Geological Survey, Denver, CO, United States Holland-Sears, A (ahollandsears@fs.fed.us), White River National Forest, US Forest Service, Glenwood Springs, CO, United States Despite downward trends in precipitation sulfate concentrations across Colorado, high-elevation lakes in several wilderness areas in the region show sharp increases in lake-water sulfate concentrations during 1985-2008. Similar increases in sulfate concentrations have been reported for numerous alpine lakes in Europe, which have been attributed to enhanced weathering rates, increased biological activity, and/or melting of permanent ice features caused by increasing air temperatures. Analysis of climate records from Colorado SNOTEL stations shows increases in annual air temperature of 0.43 to 0.93 °C per decade over most mountainous areas suggesting climate also may be a factor for the Colorado lakes. Sulfur isotopic data for a subset of lakes reveals that sulfate is largely derived from the weathering of pyrite, which is associated with hydrothermally altered and mineralized bedrock. Unlike the weathering of silicate minerals, pyrite breakdown is largely dependent on oxygen availability and can be accelerated by fluctuating groundwater levels, which enhance exposure of mineralized rock to oxygen as water levels decline. We suggest that during warmer, drier years the water table declines enhancing pyrite oxidation and build up of soluble salts in the unsaturated zone. During the subsequent snowmelt, these salts are flushed from soils and sediments resulting in increased solute concentrations in lakes. If climate change in mountainous areas results in increased summer warming or a greater frequency of drought years, then the magnitude of sulfate export from mineralized watersheds may continue to increase. Because pyrite is often associated with other base-metal sulfides and its breakdown generates acidity, climate changes could result in increased acidity and trace metal concentrations in surface water to levels where impacts on aquatic life may become evident. Futhermore, climate change may act to decrease critical loads in these mineralized watersheds unlike the increases in critical loads predicted for high-elevation European watersheds. Climate Change and the Water Vapor Feedback at High Altitudes and Latitudes Miller, J R (miller@marine.rutgers.edu), Marine and Coastal Sciences, Rutgers University, New Brunswick, NJ, United States Rangwala, I (imtiazr@envsci.rutgers.edu), Environmental Sciences, Rutgers University, New Brunswick, NJ, United States Chen, Y (chen.yonghua@gmail.com), Columbia University, New York, NY, United States Russell, G (russell@giss.nasa.gov), NASA/GISS, New York, NY, United States Future climate change is likely to be enhanced in high northern latitudes, and there is evidence of similar enhancement at high altitudes. In both regions there are positive feedbacks on surface temperature associated with snow and ice cover, clouds, and atmospheric water vapor. We combine observations and global climate model simulations to examine some of the connections and feedbacks among these variables for the present climate and for a future climate in which atmospheric greenhouse gases are increasing. We focus on two sites—the Tibetan Plateau and the Arctic Ocean. The well-known snow/sea-ice albedo feedback mechanism accounts for some of the projected future temperature increases in both regions. Increased water vapor, cloud cover, and cloud optical depth also play roles by increasing the flux of downward longwave radiation. For the future climate, the strength of the feedbacks can change with time. Similarities and differences between the feedbacks at the high latitude and high altitude locations are discussed, with a particular emphasis on the water vapor feedback. Drivers of aboveground primary production and litter accumulation in grass dominated systems O'Halloran, L R (riesl@science.oregonstate.edu), Oregon State University, Corvallis, OR, United States Borer, E (borer@science.oregonstate.edu), Oregon State University, Corvallis, OR, United States Chu, C (chuchj04@lzu.cn), Lanzhou University, Lanzhou, China Cleland, E E (ecleland@ucsd.edu), UCSD, San Diego, CA, United States DeCrappeo, N (ndecrappeo@usgs.gov), USGS, Corvallis, OR, United States MacDougall, A (amacdo02@uoguelph.ca), University of Guelph, Guelph, ON, Canada McCulley, R L (rebecca.mcculley@uky.edu), University of Kentucky, Lexington, KY, United States Seabloom, E (seabloom@science.oregonstate.edu), Oregon State University, Corvallis, OR, United States Wolkovich, E M (wolkovich@nceas.ucsb.edu), NCEAS, Santa Barbara, CA, United States Global trends in vegetation structure and function across major biomes spanning broad latitudinal and climatic gradients have been well characterized in the literature. However, controls on these vegetation characteristics within a specific biome are not well understood. Grasslands, for example, persist in diverse biogeographic regions throughout the world and experience a range of abiotic conditions that are likely to control regional-scale patterns in biodiversity and primary production. Here, we explore the relationship of climatic characteristics such as mean annual precipitation, interannual variability in precipitation, and mean seasonal temperature to ecosystem traits including plant species diversity, cover, aboveground primary production, and litter accumulation and estimated loss using a global network of 45 grassland sites. Our preliminary results suggest that although all of the sites are grass-dominated ecosystems, the variables controlling major ecosystem functions, such as aboveground primary production and litter accumulation, vary significantly among sites. This global scale study demonstrates the importance of within-biome variation in response to abiotic conditions, highlights the need for incorporation of this type of variability into existing dynamic global vegetation models, and provides validation data for such models. Are All Headwater Catchments the Same? Elevational Controls on Organic and Inorganic Nutrients in Headwater Catchments in the Boulder Creek Watershed, Colorado Front Range Parman, J (jordan.parman@colorado.edu), Geography, University of Colorado, Boulder, CO, United States Williams, M W (markw@snobear.colorado.edu), Geography, University of Colorado, Boulder, CO, United States High-elevation ecosystems have become the focus of recent biogeochemical research due to their unique and complex processes, but also because these systems may serve as an early warning system for the potential effects of climate change. In the Colorado Front Range, it is expected that alpine areas will continue to experience greater annual precipitation, as well as an increase in atmospheric deposition of inorganic nitrogen (Williams and Tonnessen, 2000). Past studies have shown that these mountain systems tend to amplify such environmental changes in specific areas of the landscape. The Landscape Continuum Model (LCM, Seastedt et al., 2004) proposed a conceptual framework for how mountain ecosystems accumulate and redistribute exogenous material from the atmosphere and endogenous material derived from the mountain itself, emphasizing the importance of transport processes and redeposition of nutrients and water across highly varying and complex terrain. Here, we test the LCM by comparing and contrasting changes in organic and inorganic nutrients in stream waters of headwater catchments along an elevational gradient in the Colorado Front Range. We simultaneously collected water samples at four gauged headwater catchments: (1) Green Lakes Valley (3,500 m); (2) Como Creek (2,900 m); Gordon Gulch (2,400 m); and Betasso (1,830 m). All water samples were analyzed for DOC, DON, DOP, nitrate, and ammonium. Additionally, spectroscopic techniques were used to determine the quality of DOC. These measurements, along with supporting information on soil C:N ratios and climate data, allow us to determine how elevational position controls: (a) the redistribution of exogenous materials from the regional environment such as nitrate in wetfall; and (b) endogenous sources originating from montane areas such as DOC and DON, while controlling for catchment size, aspect, and underlying geology. Seastedt, T. R., W. D. Bowman, T. N. Caine, D. McKnight, A. Townsend & M. W. Williams (2004) The landscape continuum: A model for high-elevation ecosystems. Bioscience, 54, 111-121. Williams, M. W. & K. A. Tonnessen (2000) Critical loads for inorganic nitrogen deposition in the Colorado Front Range, USA. Ecological Applications, 10, 1648-1665. The role of cold-air drainage in explaining spatial patterns of temperature trends in the Western U.S. Pepin, N C (nicholas.pepin@port.ac.uk), Geography, University of Portsmouth, Portsmouth, United Kingdom Daly, C (daly@nacse.org), Geosciences, Oregon State University, Corvallis, OR, United States Lundquist, J D (jdlund@u.washington.edu), Civil Engineering, University of Washington, Seattle, WA, United States Understanding spatial patterns of climate change in the western U.S. is of fundamental importance to understanding impacts on the landscape. Because much of the western U.S. is topographically complex, and shows distinct microclimates, there is no reason therefore to expect past or future patterns of change to be spatially smooth. In particular cold air drainage in mountain valleys causes decoupling of the temperature regime in many locations from the free atmosphere. This is a major feature of the western U.S. climate which is often dominated by relatively cloud-free high pressure conditions. We investigate here whether this decoupling has seen significant temporal change in the past, and the implications of such a change for patterns of warming observed across the western U.S. We analyse temperature trends from 494 long-term weather stations in the western U.S. for the period 1948-2006, from the GHCNv2 dataset, supplemented by additional COOP stations. There is a range of topographic incision and elevation in the sites chosen. At each location we derive monthly synoptic indices representative of the degree of anticyclonicity vs cyclonicity using NCEP/NCAR 700 mb reanalysis pressure fields. The number of anticyclonic days minus the number of cyclonic days (AC) is strongly related to temperature anomalies at exposed convex sites and hilltops where the free atmosphere controls the temperature signal. At cold air drainage sites which tend to be in topographic concavities the relationship is much weaker. We use the gradient of the A-C/temperature relationship to represent a coupling index which is high at exposed free-air locations and low at cold-air drainage locations. There are clear relationships between the past rate of warming and this coupling index across the western U.S. but these are seasonally determined. On a mean annual basis there are no strong relationships between temperature trend magnitude, elevation, the degree of topographic incision or the coupling index. However, in winter the rate of warming is generally weaker at cold-air pool locations, but especially when snow cover plays a major role (an ice-box effect). In fall the relationship is reversed with cold air pools warming more rapidly relative to exposed free-air locations. Summer, when small-scale convection dominates the climate, shows no patterns. Our results show that future change may be different (either amplified or muted dependent on season and/or surface characteristics) in locations prone to cold-air drainage, in comparison with the free-atmosphere, and in order to achieve successful downscaling of future climate change such processes need to be understood. 20th Century Trends In The Maximum And Minimum Temperatures In Colorado’s San Juan Mountains Rangwala, I (rangwala.imtiaz@gmail.com), UCAR, Boulder, CO, United States Miller, J R (miller@marine.rutgers.edu), Marine Sciences, Rutgers University, New Brunswick, NJ, United States We examine the maximum (Tmax) and minimum (Tmin) temperature changes in San Juan Mountain (SJM) region of southwestern Colorado between 1895-2005. We analyze monthly averaged observations from 6 National Weather Service (NWS) stations between 1895-1949, and 25 NWS stations between 1950-2005. These changes are evaluated on annual, seasonal and monthly bases. Annually, our results suggest a long-term gradual warming trend in Tmin and no such discernable trend in Tmax. However, between 1990 and 2005, the region experiences a rapid warming trend with both Tmax and Tmin increasing by 1 degree C. Between 1950 and 1985, there is a regional cooling trend during which there are significant decreases in Tmax and almost no trend in Tmin. Similar to the annual trends, only Tmin shows a gradual warming trend during the 20th century during all seasons. Furthermore, during fall and summer, there is a lower correlation between Tmax and Tmin as compared to winter and spring. Between 1990-2005, Tmax increases more than Tmin during summer and spring, whereas Tmin shows greater increases during winter. We also examine Tmax and Tmin trends from 23 Snow Telemetry (SNOTEL) sites in the region between 1984-2005. We find strong correlation between NWS and SNOTEL observations, both annually and seasonally. Between 1990-2005, the largest warming at the SNOTEL sites occurs during summer while it is largest during winter at the NWS sites. Spatially, there are similar increases in Tmax and Tmin except in the central mountain region, where increases in Tmin started earlier and are greater. Physical mechanisms for these observed trends in Tmax and Tmin will be discussed. Holocene Paleoenvironment of the North-central Great Basin: Preliminary Results from Favre Lake, Northern Ruby Mountains, Nevada Starratt, S (sstarrat@usgs.gov), U S Geological Survey, Menlo Park, CA, United States Wahl, D (dwahl@usgs.gov), U S Geological Survey, Menlo Park, CA, United States Wan, E (ewan@usgs.gov), U S Geological Survey, Menlo Park, CA, United States Anderson, L (la@gmail.gov), Environmental and Earth System Science, Stanford University, Stanford, CA, United States Wanket, J (jwanket@csus.edu), Geography, California State University, Sacramento, CA, United States Olson, H (holson@usgs.gov), U S Geological Survey, Menlo Park, CA, United States Lloyd-Davies, T (tlloyddavies@usgs.gov), U S Geological Survey, Menlo Park, CA, United States Kusler, J (jkusler@gmail.com), Geography, California State University, Sacramento, CA, United States Little is known about Holocene climate variability in north-central Nevada. This study aims to assess changes in watershed vegetation, fire history, lake levels and limnological conditions in order to understand secular to millennial-scale changes in regional climate. Favre Lake (2,899 m a.s.l.; 12 m deep; 7.7 hectares) is a flow-through lake in the northern Ruby Mountains. The primary sources of influent, both of which appear to be intermittent, are Castle Lake (2,989 m a.s.l.) and Liberty Lake (3,077 m a.s.l.). The bedrock of the three lake basins is early Paleozoic marble and Mesozoic granite and metamorphic rocks. Bathymetric maps and temperature, pH, salinity, and conductivity profiles have been generated for Favre Lake. Surface samples and a series of cores were also collected using a modified Livingstone piston corer. The presence of the Mazama ash in the basal sediment (~4 m below the sediment/water interface) indicates the record extends to ~7,700 cal yr B.P. Magnetic susceptibility (MS) and loss-on-ignition data indicate that the sediments in the lowest part of the core contain primary and reworked Mazama ash. About 2,000 years ago CaCO3 increased from 2 to 3% of the inorganic sediment. The upper 25 cm of the core are marked by an increase in MS which may indicate increased erosion due to grazing. Between about 7,700 and 6,000 cal yr B.P. the diatom flora is dominated by a diverse assemblage of benthic species. The remainder of the core is dominated by Fragilaria, suggesting that lake level rose and flooded the shelf that surrounds the depocenter of the lake. This is supported by changes in the abundance of the aquatic fern Isoetes. Pinus and Artemisia dominate the pollen record, followed by subordinate levels of Poaceae, Asteraceae, Amaranthaceae, and Sarcobatus. The late early Holocene (7,700-6,000 cal yr B.P.) is dominated by Pinus which is present in reduced amounts during the middle Holocene (6,000-3,000 cal yr B.P.) and then returns to dominance in the late Holocene (post-3,000 cal yr B.P.). Future research will include analysis of both macro- and micro-charcoal abundances. The charcoal record will augment the suite of data presented here by providing independent evidence of variability in precipitation regimes and drought history. An additional set of cores from a perennial wetland on the eastern edge of the range, Ruby Marsh, will provide a low elevation paleoclimatic counterpoint to this alpine site. Effects of Climate Change on Alpine Lakes in the Georgia Basin, British Columbia, Canada Strang, D M (donnastrang@trentu.ca), Environmental and Life Science, Trent University, Peterborough, ON, Canada Aherne, J (jaherne@trentu.ca), Environmental and Life Science, Trent University, Peterborough, ON, Canada Alpine lakes are sensitive to the effects of climate change due to their dilute nature, dependence on glacial processes as well as their susceptibility to changes in temperature and precipitation. The Georgia Basin, located in the south-western corner of British Columbia, Canada is an area influenced by four mountain ranges. In the fall of 2008, a synoptic water quality survey was conducted on alpine lakes in the basin (n = 72), with elevations ranging from 90 to 2005 m.a.s.l. (mean = 1145 m), and catchment glacier coverage ranging between 0 and 62 % (mean = 3.7%). The lakes were characterized by low conductivities (mean = 9.42 µS cm-1), low DOC levels (mean = 1.12 mg L-1) and low acid-neutralizing capacities (mean = 63.0 µeq L-1). During the 20th century, air temperature in the Georgia Basin increased by 1.5 degrees C and precipitation increased between 5-35 % (depending on season). General circulation models predict that both air temperature and precipitation will continue to increase; winter temperatures increasing between 2.4 and 6.0 degrees C, summer temperatures increasing between 0.6 and 4.2 degrees C, and precipitation increasing by 100-200 mm (depending on season) by 2050. The purpose of this study is to assess the sensitivity and response of alpine lake catchments in the Georgia Basin to increased temperature and precipitation. Specifically, the study will focus on the potential changes in alpine lake chemistry across biogeoclimatic zones based on current observations of water chemistry, soil mineralogy, soil carbon and nitrogen pools and measurements of air, surface water and soil temperatures. The study will also assess the potential changes in catchment weathering rates under increased temperature. The results of this study will aid in the understanding of how climate change will affect these relatively unstudied ecosystems as well as acknowledge data gaps. It will also serve as a platform for more in depth examination of the potential ramifications of climate change on the alpine regions in the Georgia Basin as well as other high elevation mountainous regions around the world. Downscaling to the Climate Near the Ground: Measurements and Modeling Along the Macro-, Meso-, Topo-, and Microclimate Hierarchy Van de Ven, C (cvandeven@albion.edu), Geological Sciences, Albion College, Albion, MI, United States Weiss, S B (stu@creeksidescience.com), Creekside Center for Earth Observation, Menlo Park, CA, United States Most climate models are expressed at regional scales, with resolutions on the scales of kilometers. When used for ecological modeling, these climate models help explain only broad-scale trends, such as latitudinal and upslope migration of plants. However, more refined ecological models require down-scaled climate data at ecologically relevant spatial scales, and the goal of this presentation is to demonstrate robust downscaling methods. For example, in the White Mountains, eastern California, tree species, including bristlecone pine (Pinus longaeva) are seen moving not just upslope, but also sideways across aspects, and downslope into areas characterized by cold air drainage. Macroclimate in the White Mountains is semi-arid, residing in the rain shadow of the Sierra Nevada. Macroclimate is modified by mesoscale effects of mountain ranges, where climate becomes wetter and colder with elevation, the temperature decreasing according to the regionally and temporally-specific lapse rate. Local topography further modifies climate, where slope angle, aspect, and topographic position further impact the temperature at a given site. Finally, plants experience extremely localized microclimate, where surrounding vegetation provide differing degrees of shade. We measured and modeled topoclimate across the White Mountains using iButton Thermochron temperature data loggers during late summer in 2006 and 2008, and have documented effects of microclimatic temperature differences between sites in the open and shaded by shrubs. Starting with PRISM 800m data, we derived mesoscale lapse rates. Then, we calculated temperature differentials between each Thermochron and a long-term weather station in the middle of the range at Crooked Creek Valley. We modeled month-specific minimum temperature differentials by regressing the Thermochronweather station minimum temperature differentials with various topographic parameters. Topographic position, the absolute value of topographic position, and slope combined to provide a very close fit (r2>0.9) to measured inversions of >8°C. Although topoclimatic maximum temperature models have been more elusive, regressions with degree hours greater than zero (DH>0) have been modeled with September insolation and slope (r2=0.7). In paired experiments, Thermochrons also recorded the temperature differences between the environment under sagebrush (Artemisia tridentata) and in the open, with an average minimum temperature difference of 2.1°C, and maximum temperature difference of 4.5°C. When we incorporate hourly weather station data, the strength of the inversion is weakened by wind, higher relative humidity, and cloudiness. This hierarchical modeling provides a template for downscaling climate and weather to ecologically relevant scales.