Colville National Forest Macro-fungi Surveys Interagency Special Status/Sensitive Species Program

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Colville National Forest Macro-fungi Surveys
Interagency Special Status/Sensitive Species Program
Final Report, December 7, 2009
Kathy Ahlenslager,
Colville National Forest, 765 S. Main, Colville, WA,
509-684-7178, kahlenslager@fs.fed.us.
Introduction
Through this project a list of 228 macro-fungi from the Colville National Forest was created from spring
and fall surveys in 2009 (Table 1). One, Cudonia monticola, is listed as Sensitive on the Regional
Forester’s Special Status Species List (2008). This is the first list of macro-fungi taxa for the Forest. Two
cooperators, Dr. James Groth and Drew Parker, collected and identified specimens. Parker considers
another species, Volvariella smithii, quite rare.
Methods
Each of the two cooperators surveyed two locations for a total of 169 acres, as shown in Table 2. The sites
occurred across the Colville National Forest (Maps 1-7). Surveys were conducted according to the Region
6 “Macro-fungi Sporocarp Survey Protocol” (USDA FS and USDI BLM 2008). Each of the four sites was
surveyed three times, once in the spring and twice in the fall. Specimens were collected, prepared, and
identified by the cooperators.
Results
Tables 4-7 are taxa lists for each of the four sites with the date observed. Sighting forms and GPS locations
for sensitive taxa were provided by the cooperators. The Colville National Forest GIS layer and NRIS
database will be updated for sensitive taxa by Forest Service employees. A reference collection of two rare
fungi were sent to Darci Pankratz-Rivers, ISSSSP Inventory Coordinator Assistant, for curation at the
University of Washington Herbarium at the Burke Museum.
References
U.S. Department of Agriculture, Forest Service. 2008. Regional Forester’s Special Status Species List,
Sensitive Nonvascular Plants, USDA Forest Service Pacific Northwest Region January 2008. Available
online at http://www.fs.fed.us/r6/sfpnw/issssp/agency-policy/.
U.S. Department of Agriculture, Forest Service and U.S. Department of the Interior, Bureau of Land
Management. 2008. Sporocarp Survey Protocol for Macrofungi Version 1.0, December 2008. Available
online at http://www.fs.fed.us/r6/sfpnw/issssp/documents/inventories/inv-sp-fu-ver1-2008-12.pdf.
1
Table 1. Taxa by Survey Area and Season.
Bunchgrass
Lake Leo
Taxa
Meadow
Trail
Spring Fall
Spring Fall
Agrocybe praecox
Albatrellus flettii
Albatrellus ovinus
F
Amanita muscaria var. alba
F
Amanita muscaria var.
S
F
flavivolvata
Amanita pantherina
S
F
Amanita porphyria
Armillaria mellea
F
Armillaria ostoyae
Armillaria sinapina
Auricularia auricula
S
Baeospora myosura
Bisporella citrina
Boletus coniferarum
F
Boletus edulis
S
F
Boletus rubripes
Boletus smithii
S
F
Bondarzewia montana
F
Byssonectria terrestris
S
Calvatia fumosa
S
F
Camarophyllus recurvatus
F
Catathelasma ventricosum
F
Chalciporus piperatus
F
Chroogomphus ochraceus
F
Chroogomphus vinicolor
F
Chrysomphalina
chrysophylla
Clavariadelphus truncatus
Clavulinopsis laeticolor
Climacocystis borealis
Clitocybe clavipes
Clitocybe fragrans
Clitocybe gibba
F
F
Clitocybe glacialis
S
Clitocybe nebularis
Clitocybe odora var.
pacifica
Clitocybe squamulosa
F
Collybia tuberosa
F
F
Coltricia perennis
F
Conocybe lactea
Coprinus comatus
Cortinarius ahsii
S
2
Sherman
Peak Trail
Spring Fall
S
Sullivan
Creek
Spring Fall
S
F
F
F
F
F
F
F
F
F
F
S
F
F
F
F
S
F
F
S
F
F
Taxa
Cortinarius caperatus
(=Rozites caperatus)
Cortinarius cf. calochrous
Cortinarius cf. flavoroseus
Cortinarius cf. glaucopus
Cortinarius cf.
subolivascens
Cortinarius clandestinus
Cortinarius croceus
Cortinarius malachius
Cortinarius mutabilis
Cortinarius phoeniceus
Cortinarius semisanguineus
Cortinarius sp.
Cortinarius traganus
Cortinarius vibratilis
Cortinarius violaceus
Cudonia circinans
Cudonia monticola
Cystoderma amianthinum
Cystoderma fallax
Cystoderma granulosum
Dacromyces chrysospermus
(=Dacromyces palmatus)
Discina perlata
Elaphomyces muricatus
Flammulina velutipes
Floccularia albolanaripes
Fomitopsis pinicola
Fuscoboletinus
ochraceoroseus
Galerina autumnalis
Ganoderma applanatum
Gautieria graveolens
Gautieria monticola
Geoglosum umbratile
Gloeophyllum sepiarium
Gomphidius glutenosus
Gomphidius subroseus
Gymnopus acervatus
Gomphus floccosus
Guipinia helvelloides
(=Tremiscus helvelloides)
(=Phlogiotis helvelloides)
Guipiniopsis alpina
(=Heterotextus alpinus)
Bunchgrass
Meadow
Spring Fall
F
Lake Leo
Trail
Spring Fall
Sherman
Peak Trail
Spring Fall
Sullivan
Creek
Spring Fall
F
F
S
S
F
S
S
F
F
S
F
F
F
F
F
S
S
F
F
F
F
S
F
F
F
F
S
S
S
F
F
F
F
F
F
S
S
S
S
F
F
F
F
F
F
S
F
F
F
S
F
3
Taxa
Gymnopilus cf. sapineus
Gymnopilus sapineus
Gyromitra esculenta
Gyromitra infula
Gyromitra montana
Hapalopilus nidulans
Helvella lacunosa
Hericeum corraloides
Hydnellum caeurleum
Hydnellum peckii
Hydnellum regium
Hydnellum suaveolens
Hydnum rapandum
(=Dentinum rapandum)
Hygrocybe conica
Hygrocybe psittacina
Hygrophorus agathosmus
Hygrophorus bakerensis
Hygrophorus borealis
Hygrohorus camarophyllus
Hygrophorus chrysodon
Hygrophorus erubescens
var. erubescens
Hygrophorus fuscoalbus
Hygrophorus goetzii
Hygrophorus hypothejus
Hydrophorus marzuolus
Hygrophorus monticola
Hygrophorus morrisii
Hygrophorus picea
Hygrophorus pratensis
Hygrophorus pudorinus
Hygrophorus purpurescens
Hypholoma capnoides
Hypsizygus tessellatus
Inocybe geophylla
Inocybe pudica
Inocybe sororia
Inonotus-tomentosus
Laccaria laccata
Lactarius affinis var.
viridilactis
Lactarius cf. deliciosus
Lactarius fallax var.
concolor
Lactarius pallescens
Bunchgrass
Meadow
Spring Fall
Lake Leo
Trail
Spring Fall
F
S
Sherman
Peak Trail
Spring Fall
Sullivan
Creek
Spring Fall
F
F
S
S
F
S
F
F
F
S
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
S
F
S
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
4
Taxa
Bunchgrass
Meadow
Spring Fall
F
Lactarius pseudomucidus
Lactarius pubescens var.
pubescens
Lactarius rufus
Lactarius scrobiculatus
Lactarius torminosus
Lactarius uvidus
Leccinum discolor
Lentinellus montanus
S
Lentinellus vulpinus
Lentinus lepideus
Lepiota acutesquamosa
Lepiota magnispora
Lepista subconnexa
Leratiomyces squamosus
var. squamosus (=Psilocybe
squamosa)
Leucopaxillus gentianeus
(=Leucopaxillus amarus)
Limacella glishra
Lycoperdon pyriforme
Lyophyllum cf. loricatum
Lyophyllum connatum
(=Clitocybe dilatata)
Lyophyllum decastes
Marasmius androsaceus
Melanoleuca angelesiana
Melanoleuca melaleuca
Morchella cf. elata
Mycena adonis
Mycena algeriensis
Mycena longiseta
Mycena overholtsii
Mycena pura
Neolecta vitellina
Nolanea holoconiota
S
Nolanea strictia
Nolanea verna
Oligoporus caesius
(=Tyromyces caesius)
Oligoporus guttulatus
Oligoporus obductus
Onnia tomentosa (=Inonotus
tomentosus)
Panaeolus papilionaceus
var. papilionaceus
Lake Leo
Trail
Spring Fall
Sherman
Peak Trail
Spring Fall
Sullivan
Creek
Spring Fall
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
S
F
S
S
S
F
S
F
S
F
F
S
S
F
F
F
F
F
5
Taxa
Panellus serotinus
Peziza violacea
Physarum sp.
Pithya vulgaris
Pholiota astragalina
Pholiota cf. ferruginea
Pholiota flavida
Pholiota populnea
(=Pholiota destruens)
Pholiota squarrosa
Phyllotopsis nidulans
Pluteus cervinus
Polyozellus multiplex
Polyporus arcularius
Polyporus badius
Pseudohydnum gelatinosum
Pycnoporellus alboluteus
Ramaria rasilispora
Ramaria stricta
Rhodocollybia maculata
Russula adusta
Russula aeruginea
Russula brevipes var. acrior
Russula brunneola
Russula fragilis
Russula laurocerasi
Russula murrillii
Russula nigricans
Russula occidentalis
Russula sanguinaria
Russula subloculata
Russula veternosa
Sarcodon imbricatum
Sarcodon rimosus
Sarcodon stereosarcinon
Sarcodon subincarnatus
Scutellinia umbrarum
Spathularia flavida
Strobiluris occidentalis
Stropharia ambigua
Stropharia hornmanii
Stropharia pseudocyanea
Suillus brevipes
Suillus cavipes
Suillus flavidus
Bunchgrass
Meadow
Spring Fall
Lake Leo
Trail
Spring Fall
Sherman
Peak Trail
Spring Fall
Sullivan
Creek
Spring Fall
F
S
S
S
F
F
F
F
F
F
F
F
F
F
F
F
F
S
F
S
F
F
F
F
F
F
F
F
F
F
F
F
S
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
F
S
F
6
Taxa
Suillus granulatus
Suillus gravellii
Suillus ochraceoroseus
Suillus punctatipes
Suillus tomentosus
Tapinella atrotomentosa
(=Paxillus atrotomentosus)
Thaxterogaster pinguis
Trichaptum abietinum
Tricholoma atroviolaceum
Tricholoma caligatum
Tricholoma flavovirens
Tricholoma inamoenum
Tricholoma moseri
Tricholoma portentosum
Tricholoma saponaceum
Tricholoma vaccinum
Tricholoma virgatum var.
vinaceum
Tricholomopsis decora
Tricholomopsis rutilans
Tricholoma zelleri
Tyromyces chioneus
Volvariella smithii
Xeromphalina campanella
Xeromphalina cornui
Bunchgrass
Meadow
Spring Fall
F
Lake Leo
Trail
Spring Fall
Sherman
Peak Trail
Spring Fall
Sullivan
Creek
Spring Fall
F
F
F
F
F
S
S
F
F
F
F
F
S
F
F
F
F
F
F
F
F
F
F
F
F
F
F
7
S
Table 2. Macro-fungi Survey Locations on the Colville National Forest.
Number
Location and
Number Dates of Number Dates of
Surveys
of Acres
Cooperator
of
Surveys of
in Fall
Proposed
in Spring Survey
Survey
to Survey
2009
Days,
Days in 2009
Fall 2009
Spring
2009
Bunchgrass Meadows
1
June 26
2
September 30
Research Natural Area,
July 16
3 and 28
Drew Parker
Lake Leo, Dr. James
1
May 26
2
September 50
Groth
24 and
October 8
Sherman Pass, Dr.
1
July 5
2
September 40
James Groth
8 and
October 5
Sullivan Creek, Drew
1
June 1
2+
September 60
Parker
July 17
12, 15, 29,
30 and
October
21
Total 4
6
8
180
8
Number
of Acres
Surveyed
40
21
10
98
169
Table 4. 2009 Bunchgrass Meadow Surveys by Drew Parker.
Comments from Parker: “I did well on the fall surveys despite the dry September and early end of the
season from the cold snap at the beginning of October. It was an unusual year weatherwise and the fungal
fruiting was a little unusual as well. Bunchgrass peaked in late August but was still fair near the end of
September. It was a matter of keeping track of when and where it rained.”
Taxa
Albatrellus ovinus
Amanita muscaria var. alba
Amanita muscaria var. flavivolvata
Amanita pantherina
Auricularia auricula
Boletus coniferarum
Boletus edulis
Boletus smithii
Byssonectria terrestris
Calvatia fumosa
Catathelasma ventricosum
Chalciporus piperatus
Chroogomphus ochraceus
Chroogomphus vinicolor
Climacocystis borealis
Clitocybe glacialis
Collybia tuberosa
Coltricia perennis
Cortinarius ahsii
Cortinarius caperatus (=Rozites caperatus)
Cortinarius croceus
Cortinarius flavoroseus (nom. prov.)
Cortinarius phoeniceus
Cortinarius semisanguineus
Cortinarius cf. subolivascens
Cortinarius traganus
Dacromyces chrysospermus (=Dacromyces
palmatus)
Gautieria monticola
Gloeophyllum sepiarium
Gomphidius subroseus
Guipiniopsis alpina (=Heterotextus
alpinus)
Gymnopilus cf. sapineus
Gyromitra esculenta
Gyromitra infula
Hydnellum peckii
Hydnellum regium
Hydnum rapandum
Hygrocybe psittacina
Hygrophorus erubescens var. erubescens
June 26
July 16
September 3
X
September 28
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
9
Taxa
Hygrophorus hypothejus
Inocybe sororia
Laccaria laccata
Lactarius cf. deliciosus
Lactarius pseudomucidus
Lactarius rufus
Leccinum discolor
Lentinellus montanus
Leucopaxillus gentianeus (=Leucopaxillus
amarus)
Lyophyllum cf. loricatum
Oligoporus caesius (=Tyromyces caesius)
Nolanea holoconiota
Pholiota cf. ferruginea
Pholiota flavida
Pholiota squarrosa
Phyllotopsis nidulans
Pluteus cervinus
Leratiomyces squamosus var. squamosus
(=Psilocybe squamosa)
Rhodocollybia maculata
Russula adusta
Russula aeruginea
Russula fragilis
Russula murrillii
Sarcodon imbricatum
Scutellinia umbrarum
Stropharia ambigua
Stropharia hornmanii
Suillus brevipes
Suillus cavipes
Suillus flavidus
Suillus granulatus
Suillus ochraceoroseus
Suillus punctatipes
Suillus tomentosus
Thaxterogaster pinguis
Trichaptum abietinum
Tricholoma atroviolaceum
Tricholoma flavovirens
Tricholoma inamoenum
Tricholoma moseri
Tricholoma portentosum
Tricholoma saponaceum
Tricholoma vaccinum
Tricholoma virgatum var. vinaceum
Tricholomopsis rutilans
June 26
July 16
September 3
September 28
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
10
Taxa
Xeromphalina cornui
June 26
11
July 16
September 3
September 28
X
Table 5. 2009 Lake Leo Trail Surveys by Dr. Jim Groth.
Comments from Groth: “Spring counts, 26 May 2009. Weather: clear, cool; elevation 3150 feet; northfacing aspect. Some rain in last two weeks, but not a lot. A larch-lodgepole fire forest, approx. 60 years
old. Fall counts, 24 September and 8 October 2009. Weather both days cloudy, dry, cool. Moderate rain
for two to three weeks previous.”
Taxa
Comments
Armillaria mellea
One location, terrestrial, no scales, lower
stem yellowish
One location, many stems
Many, one location
Several locations
Single specimen
Common, host mushrooms not generally
visible
Two specimens, one location
Dry, light brown cap, several in one area.
At least four distinct species of nonpurple
Cortinarius in several locations.
Several in one mossy location
Several in two spots
One cluster
Several locations
Several on a single log
Three locations
Several on one log
One small old specimen
Two locations, several
One location, several
One
One large fruiting body
Several, one location
One
3 large in ca. 5 m cluster, delicious
Common
Yellow, resupinate, cracked, 5 x 8 cm, on
edge of 20 cm diameter subalpine fir
stump
Common, spruce and larch cones
Common
Single at two locations
Two clusters
Bondarzewia montana
Camarophyllus recurvatus
Clitocybe gibba
Clitocybe squamulosa
Collybia tuberosa
Cortinarius semisanguineus
Cortinarius sp.
Cortinarius spp.
Cystoderma granulosum
Discina perlatua
Flammulina velutipes
Fomitopsis pinicola
Galerina autumnalis
Gomphidius subroseus
Gymnopilus sapineus
Gyromitra esculenta
Hypholoma capnoides
Hygrophorus chrysodon
Hygrophorus purpurescens
Inonotus tomentosus
Laccaria laccata
Lentinus lepideus
Morchella elata
Mycena pura
Physarum sp. (slime mold)
Strobiluris occidentalis
Suillus cavipes
Suillus gravellii
Xeromphalina campanella
May 26,
2009
12
September
24 and
October 8,
2009
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
Table 6. 2009 Sherman Peak Trail Surveys by Dr. Jim Groth.
Comments from Groth: “Spring counts. 5 July 2009. Weather moist, but not wet; elevation 5500-5600
feet, N-facing aspect; mature, mixed forest of lodgepole pine, subalpine fir, and Douglas fir. Fall counts. 8
September and 5 October 2009. September date: moderately warm and dry. No evidence of recent rain.
October date: weather cool, high about 35, snow cover about 50% near highway to 80% on Sherman Peak
Loop Trail. Snow is not deep enough to cover larger musrooms. However, dry conditions on pass nearly
all summer meant that few mushrooms were present before cold snap, as was observed there in August
(brief visit) and September.”
Taxa
Notes
July 5,
2009
Agrocybe praecox
Clitocybe squamulosa
Cortinarius. sp.
Cortinarius malachius
Fuscoboletinus ochraceoroseus
Gomphidius subroseus
Morchella elata
Mycena algeriensis
Polyporus arcularius
Polyporus badius
Suillus brevipes
Tricholoma zelleri
Several spots, many
19 specimens in three spots
Dry, light brown, plain white stem
3 spots, singles
Two locations, old specimens
2 spots
2 nice specimens
X
X
X
X
September
8 and
October 5
2009
X
X
X
X
Two locations
Several locations
X
X
X
Five specimens in one location, spores
up to 5 microns long, broadly elliptical,
with oil drop
13
X
Table 7. 2009 Sullivan Creek Surveys by Drew Parker.
Comments from Parker: “I did well on the fall surveys despite the dry September and early end of the
season from the cold snap at the beginning of October. It was an unusual year weatherwise and the fungal
fruiting was a little unusual, as well. The lower stretch of Sullivan Creek never did produce very much, but
the upper end near Gypsy Meadows was very good for a while.”
Taxa
Agrocybe praecox
Albatrellus flettii
Albatrellus ovinus
Amanita muscaria var.
flavivolvata
Amanita porphyria
Armillaria ostoyae
Armillaria sinapina
Baeospora myosura
Bisporella citrina
Boletus rubripes
Catathelasma ventricosum
Chrysomphalina
chrysophylla
Clavariadelphus truncatus
Clavulinopsis laeticolor
Clitocybe clavipes
Clitocybe fragrans
Clitocybe glacialis
Clitocybe nebularis
Clitocybe odora var.
pacifica
Conocybe lactea
Coprinus comatus
Cortinarius caperatus
(=Rozites caperata)
Cortinarius cf. calochrous
Cortinarius cf. glaucopus
Cortinarius clandestinus
Cortinarius cf. flavoroseus
Cortinarius mutabilis
Cortinarius vibratilis
Cortinarius violaceus
Cudonia circinans
Cudonia monticola
Cystoderma amianthinum
Cystoderma fallax
Discina perlata
Elaphomyces muricatus
Floccularia albolanaripes
Ganoderma applanatum
June
1
X
July
17
September
15
29
12
30
October
21
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
14
Taxa
Gautieria graveolens
Geoglosum umbratile
June
1
X
July
17
September
15
29
12
30
October
21
X
Gloeophyllum sepiarium
X
Gomphidius glutenosus
Gomphus floccosus
Guipinia helvelloides
(=Tremiscus helvelloides)
(=Phlogiotis helvelloides)
Gymnopilus cf. sapineus
Gymnopus acervatus
Gyromitra esculenta
Gyromitra montana
Hapalopilus nidulans
Helvella lacunosa
Hericeum corraloides
Hydnellum caeurleum
Hydnellum peckii
Hydnellum suaveolens
Hydnum rapandum
(=Dentinum rapandum)
Hygrocybe conica
Hygrophorus agathosmus
Hygrophorus bakerensis
Hygrophorus borealis
Hygrophorus
camarophyllus
Hygrophorus chrysodon
Hygrophorus erubescens
var. erubescens
Hygrophorus fuscoalbus
Hygrophorus goetzii
Hydrophorus marzuolus
Hygrophorus monticola
Hygrophorus morrisii
Hygrophorus picea
Hygrophorus pratensis
Hygrophorus pudorinus
Hypholoma capnoides
Hypsizygus tessellatus
Inocybe geophylla
Inocybe pudica
Inocybe sororia
Laccaria laccata
Lactarius affinis var.
viridilactis
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
15
X
X
Taxa
Lactarius cf. deliciosus
Lactarius fallax var.
concolor
Lactarius pallescens
Lactarius pubescens var.
pubescens
Lactarius scrobiculatus
Lactarius torminosus
Lactarius uvidus
Lentinellus vulpinus
Lepiota acutesquamosa
Lepiota magnispora
Lepista subconnexa
Limacella glishra
Lycoperdon pyriforme
Lyophyllum connatum
(=Clitocybe dilatata)
Lyophyllum decastes
Marasmius androsaceus
Melanoleuca angelesiana
Melanoleuca melaleuca
Morchella cf. elata
Mycena adonis
Mycena longiseta
Mycena pura
Melanoleuca angelesiana
Mycena overholtsii
Neolecta vitellina
Nolanea strictia
Nolanea verna
Oligoporus guttulatus
Oligoporus obductus
Onnia tomentosa
(=Inonotus tomentosus)
Panaeolus papilionaceus
var. papilionaceus
Panellus serotinus
Peziza violacea
Pholiota astragalina
Pholiota populnea
(=Pholiota destruens)
Pithya vulgaris
Polyozellus multiplex
Polyporus badius
Pseudohydnum
gelatinosum
June
1
July
17
September
15
29
12
30
October
21
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
16
Taxa
Pycnoporellus alboluteus
Ramaria rasilispora
Ramaria stricta
Russula adusta
Russula aeruginea
Russula brevipes var.
acrior
Russula brunneola
Russula laurocerasi
Russula nigricans
Russula occidentalis
Russula sanguinaria
Russula subloculata
Russula veternosa
Sarcodon rimosus
Sarcodon stereosarcinon
Sarcodon subincarnatus
Spathularia flavida
Stropharia ambigua
Stropharia hornmannii
Stropharia pseudocyanea
Tapinella atrotomentosa
(=Paxillus atrotomentosus)
Tricholoma caligatum
Tricholoma flavovirens
Tricholoma saponaceum
Tricholoma vaccinum
Tricholoma virgatum var.
vinaceum
Tricholomopsis decora
Tyromyces chioneus
Volvariella smithii
Xeromphalina campanella
June
1
July
17
September
15
29
12
30
October
21
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
X
17
18
19
20
21
22
23
24
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