2/7/2011 40 Physiology, Homeostasis, and Temperature Regulation KEY CONCEPTS 1. Four Tissue Types in Animals 2. Energy & Metabolism 3. Why/how do Animals Regulate Internal Environment? HOMEOSTASIS 4. How does temperature affect animals & how do they regulate their body temperature? Animal Form & Function • Anatomy – • Physiology – • STRUCTURE LINKED TO FUNCTION Structure and function 2x • Key theme: structure is linked to function • Ex. Compare appendages of vertebrates: elephant ears rabbit ears whale and dolphin flukes/dorsal fin X=1 2x x x 2x x Surface area = 6x2 Volume = x3 Surface area = 6(2x)2 = 24x2 Volume = (2x)3 = 8x2 Ratio of SA to V: 6:1 3:1 1 2/7/2011 Four types of tissue: • Epithelial • Connective • Muscle • Nervous Tissues Form Organs Organs: consist of multiple tissues Organ system: group of organs that function together Energy Sources & Allocation ORGAN: STOMACH Stomach wall arranged in layers: • Epithelial tissue (lines inner surface) • Connective tissue (support) • Muscle (Smooth) tissue (stomach contraction) • Nervous tissue (controls movement & influences secretions) Animals harvest chemical energy from food Energy-containing molecules from food used to make ATP (Adenosine Triphosphate; powers cellular work) ORGAN SYSTEM: DIGESTIVE SYSTEM 2 2/7/2011 Metabolic rate • Metabolic rate - Quantifying Energy Use • Direct calorimetry • Calorie (cal) – • Indirect calorimetry Metabolic Rates METABOLIC RATE INCREASES WITH BODY MASS Basal Metabolic Rate (BMR): Endotherm What does this mean? Standard Metabolic Rate (SMR): Ectotherm Fasted, Resting, No Growth, No Reproduction Slope of curve = 0.75: Slope < 1, so as body size increases energy required per g decreases 3 2/7/2011 Why does mass specific metabolic rate decrease with larger body size? Animals regulate their internal environment 1.Single-celled & simple multicellular (i.e. sponge) organisms: Short Answer – Don’t know!!! Long answer –several hypotheses: Animals regulate their internal environment Homeostasis 2. Multicellular organisms: Homeostasis • Conformers – • Regulators – 4 2/7/2011 What kind of variables are regulated? For Humans: How is homeostasis controlled? Homeostatic control mechanism Receptor – Control center – Effector – Regulation Controlled by Nervous & Endocrine Systems Information required: • Set point: • Feedback information: • Error signal: How is homeostasis controlled? Negative feedback: Positive feedback: Feedforward information: 5 2/7/2011 Homeostasis *Homeostasis does NOT mean that physiological variables do not change Temperature affects physiological processes (rate increases w/ increasing temperature) Q10 - Homeostasis is EXPENSIVE! Regulating Body Remperature: Model for homeostasis Energy Budget (for temperature regulators) Endotherms Ectotherms Poikilotherm Homeotherm 6 2/7/2011 Heat Production Heat Loss Heat Loss Metabolism Convection Radiation Non-shivering thermogenesis Evaporation Shivering thermogenesis Tiki Conduction Thermoregulatory Strategies Strategies for Staying Warm and Cool Circulatory Adaptations Counter Current Heat Exchangers (CCHE)– Scholander and Schevill, 1955 7 2/7/2011 Circulatory Adaptations Thermal Windows Torpor vs. Hibernation Behavior Behavior: Can increase or decrease heat loss Body temperature regulation reliant on feedback Hypothalamus: “thermostat” Negative Feedback Mechanism Set Point can Change 8