Biodiversity, Species Interactions, and Population Control Southern Sea Otter Fig. 5-1a, p. 104 5-1 How Do Species Interact? • The interactions between species in an ecosystem affect: • the resource use in an ecosystem and • population sizes of the species in an ecosystem. Five types of Interactions • Interspecific Competition • Predation • Parasitism • Mutualism • Commensalism Five types of Interactions • Interspecific Competition • Predation • Parasitism • Mutualism • Commensalism Interspecific Competition • Species compete for limited resources like • • • • Water Food/Nutrients Light Habitat/Space Five types of Interactions • Interspecific Competition •Predation • Parasitism • Mutualism • Commensalism Predation • Predators may capture prey by 1. Walking 2. Swimming 3. Flying 4. Pursuit and ambush 5. Camouflage 6. Chemical warfare Predator-Prey Relationships Fig. 5-4, p. 107 Avoiding Predation • Prey may avoid capture by 1. Run, swim, fly 2. Protection: shells, bark, thorns 3. Camouflage 4. Chemical warfare 5. Warning coloration 6. Mimicry 7. Deceptive looks 8. Deceptive behavior (b) Wandering leaf insect Fig. 5-5b, p. 109 (c) Bombardier beetle Fig. 5-5c, p. 109 (d) Foul-tasting monarch butterfly Fig. 5-5d, p. 109 (e) Poison dart frog Fig. 5-5e, p. 109 (f) Viceroy butterfly mimics monarch butterfly Fig. 5-5f, p. 109 (g) Hind wings of Io moth resemble eyes of a much larger animal. Fig. 5-5g, p. 109 (h) When touched, snake caterpillar changes shape to look like head of snake. Fig. 5-5h, p. 109 Predator and Prey Interactions Can Drive Each Other’s Evolution • Intense natural selection pressures between predator and prey populations • Coevolution • Interact over a long period of time • Example - Bats and moths: echolocation of bats and sensitive hearing of moths Coevolution: A Langohrfledermaus Bat Hunting a Moth Fig. 5-6, p. 110 Five types of Interactions • Interspecific Competition • Predation •Parasitism • Mutualism • Commensalism Parasitism • Parasitism – species feed off other species or live on or in them • Parasite is usually much smaller than the host • Parasite rarely kills the host • Parasite-host interaction may lead to coevolution Parasitism: Trout with Blood-Sucking Sea Lamprey Fig. 5-7, p. 110 Parasitism: Tapeworm in Human Eye Five types of Interactions • Interspecific Competition • Predation • Parasitism •Mutualism • Commensalism Mutualism • Mutualism – both species benefit • Nutrition and protection relationship • Clownfish and anemone • Gut inhabitant mutualism • Bacteria in our intestines • Not cooperation: it’s mutual exploitation Mutualism: Hummingbird and Flower Fig. 5-8, p. 110 Mutualism: Oxpeckers Clean Rhinoceros; Anemones Protect and Feed Clownfish Fig. 5-9, p. 111 Five types of Interactions • Interspecific Competition • Predation • Parasitism • Mutualism •Commensalism Commensalism • Commensalism • One species benefits, one not harmed • Bromeliad (air plant) in tree • Birds nesting in trees What Limits the Growth of Populations? • No population can continue to grow indefinitely because of • limitations on resources and • competition among species for those resources. Most Populations Live Together in Clumps or Patches • Population: group of interbreeding individuals of the same species • Population distribution 1. Clumping (ex. Flocks of birds) 2. Uniform dispersion 3. Random dispersion Generalized Dispersion Patterns Fig. 5-12, p. 112 Most Populations Live Together in Clumps or Patches • Why clumping? 1. 2. 3. 4. Species cluster where resources are available Clumps have a better chance of finding resources Protects some animals from predators Packs allow some to get prey Population of Snow Geese Fig. 5-11, p. 112 Population Growth and Decline • Population size governed by • • • • Births Deaths Immigration Emigration • Population change = (births + immigration) – (deaths + emigration) Population Growth and Decline • Age structure • Pre-reproductive age “kids” • Reproductive age “adults” • Post-reproductive age “elderly” • Sizes of each group will drive the growth or decline of population in the future • Example: A population with 70% of kids will most likely have a population growth spike as those kids become adults Some Factors Can Limit Population Size • Range of tolerance • Variations in physical and chemical environment that impact a population’s ability to survive and reproduce • Temperature, Rain, Nutrients… Trout Tolerance of Temperature Fig. 5-13, p. 113 Some Factors Can Limit Population Size • Limiting factor principle • Too much or too little of any physical or chemical factor can limit or prevent growth of a population, even if all other factors are at or near the optimal range of tolerance “Population can only grow as much as its scarcest resource!” No Population Can Grow Indefinitely: J-Curves and S-Curves • Size of populations controlled by limiting factors: • • • • • Light Water Space Nutrients Exposure to too many competitors, predators or infectious diseases No Population Can Grow Indefinitely: J-Curves and S-Curves • Environmental resistance • All factors that act to limit the growth of a population • Carrying capacity (K) • Maximum population a given habitat can sustain • Calculated by looking at the limiting factors No Population Can Grow Indefinitely: J-Curves and S-Curves • Exponential growth • Starts slowly, then accelerates to carrying capacity when meets environmental resistance • Known as “J” curve due to shape • No limiting factors yet No Population Can Grow Indefinitely: J-Curves and S-Curves (3) • Logistic growth • Decreased population growth rate as population size reaches carrying capacity Yellow line = calculated theoretical carrying capacity Green = actual population Logistic Growth of Sheep in Tasmania Fig. 5-15, p. 115 When a Population Exceeds Its Habitat’s Carrying Capacity, Its Population Can Crash • A population can “briefly” exceed (overshoot) the area’s carrying capacity due to: • Reproductive time lag • If it greatly overshoots the carrying capacity, the result is: • Population crash and • Damage may reduce area’s new carrying capacity Exponential Growth, Overshoot, and Population Crash of a Reindeer Fig. 5-17, p. 116 Species Have Different Reproductive Patterns • Some species • • • • Many, usually small, offspring Little or no parental care Massive deaths of offspring Ex. Insects, bacteria, algae Species Have Different Reproductive Patterns • Other species • • • • • • Reproduce later in life Small number of offspring with long life spans Young offspring grow inside mother Long time to maturity Protected by parents, and potentially groups Ex. Humans, Elephants Under Some Circumstances Population Density Affects Population Size • Density-dependent population controls • • • • Predation Parasitism Infectious disease Competition for resources Several Different Types of Population Change Occur in Nature • Stable • Irruptive • Population surge, followed by crash • Cyclic fluctuations, boom-and-bust cycles • Top-down population regulation • Bottom-up population regulation • Irregular Humans Are Not Exempt from Nature’s Population Controls • Ireland • Potato crop in 1845 • Bubonic plague • Fourteenth century • AIDS • Global epidemic 5-3 How Do Communities and Ecosystems Respond to Changing Environmental Conditions? • Concept 5-3 The structure and species composition of communities and ecosystems change in response to changing environmental conditions through a process called ecological succession. Communities and Ecosystems Change over Time: Ecological Succession • Natural ecological restoration • Primary succession • Secondary succession Some Ecosystems Start from Scratch: • Primary Succession • No soil in a terrestrial system or no bottom sediment in an aquatic system • Takes hundreds to thousands of years to build up soils/sediments to provide necessary nutrients • Ex. – Bare rock after a glacier retreats Primary Ecological Succession Fig. 5-19, p. 119 Some Ecosystems Do Not Have to Start from Scratch • Secondary Succession • Some soil remains in a terrestrial system or bottom sediment remains in an aquatic system • Ecosystem has been • Disturbed • Removed • Destroyed • Ex – old farmland, forest fires Natural Ecological Restoration of Disturbed Land Fig. 5-20, p. 120 Secondary Ecological Succession in Yellowstone Following the 1998 Fire Fig. 5-21, p. 120 Some Ecosystems Do Not Have to Start from Scratch: Secondary Succession (2) • Primary and secondary succession • Tend to increase biodiversity • Increase species richness and interactions among species • Primary and secondary succession can be interrupted by • • • • • Fires Hurricanes Clear-cutting of forests Plowing of grasslands Invasion by nonnative species Science Focus: How Do Species Replace One Another in Ecological Succession? • Facilitation • Inhibition • Tolerance Succession Doesn’t Follow a Predictable Path • Traditional view • Balance of nature and a climax community • Current view • Ever-changing mosaic of patches of vegetation • Mature late-successional ecosystems • State of continual disturbance and change Living Systems Are Sustained through Constant Change • Inertia, persistence • Ability of a living system to survive moderate disturbances • Resilience • Ability of a living system to be restored through secondary succession after a moderate disturbance • Some systems have one property, but not the other: tropical rainforests Three Big Ideas 1. Certain interactions among species affect their use of resources and their population sizes. 2. There are always limits to population growth in nature. 3. Changes in environmental conditions cause communities and ecosystems to gradually alter their species composition and population sizes (ecological succession).