Chapter 29-33 Plant Unit Importance Evolution to land plants-classification Plant structure and function Plant Reproduction Plant Responses Plants and People Tree Man -Actually, this is believed to be caused by the Human Papilloma Virus (HPV) -Creepy though! Plants and People (really) Plants are used as food Plants are used for medicine Plants are used for clothing and fabric dyes Plants are used as fuels Plants are also used for ornamental purposes, improving erosion, reducing noise, providing habitats for wild animals, acting as windbreaks, providing shade, and moderating temperatures See page 566, Figure 29-3…good “Nonfood” list of plant uses Plants and the Environment Plant Ecology – plant interaction in environment Photosynthesis Provide organisms with inorganic nutrients Formation/maintenance of soil Plant and animal interactions Pollination Size, shape, color attract pollinators Plant and microbe interactions Fungi – Mycorrhizae = symbiotic relationship betw. Plants and fungi Bacteria – nitrogen fixing = take gas and fix so plants can use Plant protection – prevents new plant pests Harmful plants – can cause illness, rashes, allergies, and sometimes death Evolution of Land Plants Changes in Nutrient Absorption Vascular Tissue – transports water and dissolved substances Xylem-water and inorganic nutrients in one direction Phloem- organic compounds in any direction based on need Reproduction by spores and seeds – keep reproductive cells from drying out; can send structures via air Prevention of Water Loss – cuticle is waxy; prevents water loss but also keeps out carbon dioxide FYI: *Buoyancy of water provides physical support for algae Lignin is an evolutionary trait that hardens the cell walls of plants allowing them to have strength to stand upright Classification – Page 580, Figure 30-1 Nonvascular – no true vascular tissue nor true roots, stems, or leaves Vascular, seedless – vascular tissue and true roots, stems, and leaves (ferns) Vascular, seed – vascular tissue and true roots, stems, and leaves; also produce seeds for reproduction (4 phyla of gymnosperms, 1 phyla of angiosperms) Gymnosperms Include pine trees Produce seeds that are not enclosed in fruits Angiosperms Flowering plants Produce seeds within a protective fruit Alternating Life Cycles 2 phases 1st = diploid (2n); sporophyte plant; produces spores 2nd = haploid (1n); gametophyte plant; produces egg and sperm Alternation of generations = alternates between the gametophyte phase and sporophyte phase Nonvascular Plants “Bryophytes” Do not form true roots, stems, and leaves Usually grow on land near streams and rivers Seedless; produce spores Mostly terrestrial Have alternation-of-generations life cycle Very small Need water to produce sexually (sperm must swim) Phyla: Bryophyta, Hepatophyta, and Antherocerophyta Phylum Bryophyta Pioneer plants = mosses; often first species to inhabit a barren area Carpet moss Moss gametophytes anchored by non-vascular rhizoids Rhizoids anchor moss and absorb water and inorganic nutrients Sphagnum (peat moss) – partially decomposed plant matter; some countries used as fuel; widely used to enhance the water-retaining ability of potting and gardening soils. Phyla Hepatophyta and Anthocerophyta Hepatophyta – liverworts; grow in moist, shady areas Lie close to ground Most have thin, transparent leaflike structures arranged along a stemlike axis. Anthocerophyta – hornworts; resemble liverworts; grow in moist, shady areas. Similar characteristic to algae: each cell usually has a single large chloroplast rather than numerous small ones Changes in Nutrient Absorption Land Plants Algae Absorb resources from surrounding water Vascular Tissue transportation Air Light and carbon dioxide Shoots/leaves Soil Water and mineral nutrients Roots Vascular Plants Vascular Seed Plants Phylum Cycadophyta Cycads Seedless Vascular Plants Phylum Psilotophyta Whisk Ferns Phylum Lycophyta Club Mosses Phylum Sphenophyta Horsetails Phylum Pterophyta Ferns Phylum Ginkgophyta Ginkgoes Phylum Coniferophyta Conifers Phylum Gnetophyta Gnetophytes like Ephedra Phylum Anthophyta Largest phylum of plants Over 240,000 of flowering plant Monocots vs Dicots Specialized Plant Cells – See pg 599 Parenchyma – loosely packed; cube-shaped Involved in many metabolic functions (photosynthesis, storage of water and nutrients, and healing) Bulk of nonwoody plants (ex. Flesh of apple) Collenchyma – irregular shape; thick walls Usually grouped in strands Support regions of plant that are still lengthening (ex. Celery stalk) Sclerenchyma – thick, even, rigid cell walls Support and strengthen plant Usually dies at maturity, providing support (ex. Gritty texture of pear fruit) Tissue Systems – Review Table 31-1 Dermal – outside covering of plants; surrounds ground tissue Ground – consists of all 3 types of cells; surrounds vascular tissue system; storage, metabolism, and support Vascular – transport and support; xylem and phloem; Growth in Meristems Growth where cells continuously divide Apical meristems – located at tips of stems and roots (remember the onion root tip?) Some monocots also have intercalary meristems = above bases of leaves and stems; allow rapid regrowth from damage Lateral meristems – gymnosperms and most dicots have this which allows roots and stems to increase in diameter Vascular cambium – betw. xylem and phloem…produces additional vascular tissue Cork cambium – located outside phloem…produces cork; cork cells replace epidermis in woody stems and roots; cork cells are dead cells that provide protection and prevent water loss Primary Growth – in length; apical and intercalary Secondary Growth - in diameter; lateral Roots Taproot – when first root becomes the largest root; carrots Fibrous root system – numerous small roots; many monocots…grass Adventitious – specialized roots that grow from stems and leaves; prop roots of corn (helps keep stems upright) Root Structure Root cap – covers apical meristem Root hairs - extensions of epidermal cells (increase surface area = increase plants ability to absorb water) Most form partnerships with mycorrhizal fungi = also increases surface area for absorption Just another FYI Cross section of monocot and dicot roots Stems See page 609… Shoot system and root system Segmented stems = internodes At each end of internode = node; contain 1+ leave and lateral bud (capable of developing into a new shoot) *Read over page 610 to review primary and secondary growth in stems Cross section of mature woody stem Bark – protective outer covering Heartwood – darker wood in center of tree Sapwood – lighter wood near outside Springwood/Earlywood – When water is plentiful; vascular cam. forms new xylem tissue (wide and thin-walled Summerwood/Latewood – When water is more limited; vasc. cam. forms small, thick cells. Annual Ring – abrupt change between small summerwood cells and following year’s large springwood cells. *Annual rings often do not occur in tropical trees because of their uniform year-round environment Stem functions Source to sink – CHO are “translocated” through the plant Source = where CHO are made or stored Sink = where the CHO are transported to be stored EX. In most plants…CHO from leaves to roots …to the shoot apical meristems, and to the developing flowers or fruits PressureFlow Hypothesis Page 612 -Movement in phloem -CHO are actively transported into the sieve tubes -water is also transported in by osmosis -+ pressure builds up -At “sink”, process reverses Cohesiontension theory -Water is pulled up the Stem xylem b/c water molecules are attracted to one another (cohesion) How roots absorb water and minerals and then move to leaves Trace the path of water and minerals from the roots to the leaves with the use of xylem Trace the path using a simple concept map, isolating key points Types of Leaves Leaf Tissue Another Crosssection Tissue Systems Dermal – epidermis Single layer Coated by cuticle Water, oxygen, and carbon dioxide enter and exit through stomata Mesophyll layers – Leaf = ground tissue made of chloroplast-rich parenchyma cells Palisade medophyll Spongy mesophyll Vascular Tissue Venation Moisture Maintenance Must maintain a watery internal environment Developed a cuticle – waxy coat that does NOT permit gas exchange Stomata permit gas exchange during certain times of the day Plants acquire nutrients from soil and air Three sources of a plant’s mass: Air – carbon and oxygen = used in photosynthesis = generates sugar which are the building blocks of other organic molecules Water – absorbed by plant – supplies the hydrogen for photosynthesis; solvent for transport of other molecules; makes up about 80-85% of a nonwoody plant’s mass Soil – source of inorganic nutrients (minerals) Mineral requirements of plants Most plants need 17 chemical elements to complete their life cycles: 3 are not obtained as minerals: C, O, and H Mineral nutrients absorbed in ionic form from the soil 6 of mineral nutrients plants require in greatest abundance: Nitrogen, Sulfur, Phosphorus, Potassium, Calcium, Magnesium Path of N from atmosphere to plant’s roots N is often in limited supply Used to produce proteins, nucleic acids, and hormones Makes up 80% of atmosphere, however atmospheric Nitrogen is a gaseous form of nitrogen that plants can’t use Plants must absorb from the soil in the form of mineral ions Path of N from the atmosphere to plant’s roots N must first be converted to ammonium ions or nitrate ions Certain species of soil bacteria convert atmospheric nitrogen to ammonia (nitrogen fixation) Ammonifying bacteria contribute ammonia to the soil by breaking down feces and dead leaves Nitrifying bacteria convert ammonium ions to nitrate ions Nitrogen-fixing bacteria live freely in the soil, however, some plants house their own (legumes) Bacteria found in lumps on the legumes’ roots (root nodules) Relationship benefits legumes and nitrogen-fixing bacteria Bacteria fix nitrogen Plant provides CHO and other organic nutrients to bacteria Fertilizer contents Comercially produced; contain minerals Usually enriched in nitrogen, phosphorus, and potassium (3 most deficient in farm and garden soils) “10-12-8” three-number code in fertilizer; 10% Nitrogen, 12% phosphorus, and 8 % potassium Not stored in soil for later use, though Usually wasted; taken from soil by rainwater and irrigation = may pollute groundwater, streams, and lakes Rotate crops reduces N fertilizer use Mulch and manure release minerals more gradually Plant Reproduction Life cycle of mosses Life cycle of ferns Life cycle of conifers Life cycle of mosses Production of one type of spore = homospory Life cycle of mosses = homosporous alternation of generations Male Reproductive structure Dominant generation = gametophyte Female Reproductive Structure Life cycle of ferns Dominant Generation = Sporophyte Similar to moss…most are homosporous Life cycle of conifers Microspores = male grow into male gametophytes Megaspores = female grow into female gametophytes Heterospory = production of different types of spores… Heterosporous alternation of generation Male cones of a pine release huge numbers of pollen grains Flower Parts Male reproductive structure = stamens…made up of an anther and filament Female reproductive structure = pistil…made up of a stigma, style, and ovary Flower pollination Fruit and seed dispersal Dispersed by animals, wind, water forcible discharge, and gravity Fruit = a matured ovary Simple = formed from one pistil of a single flower Aggregate = formed from several pistils of a single flower Multiple = formed from several flowers growing together Seed coat = protective coat surrounding embryo of seed Cotyledon = seed leaf (part of embryo)…dicot = 2 cotyledons Embryo Radicle = embryonic root Hypocotyl = stem between the cotyledons and radicle Epicotyl = stem above the cotyledons Structure of seeds Conditions for seed germination Water Oxygen Temperature Vegetative propagation Use of vegetative structures to produce new plants Rhizomes, bulbs, and tubers Plant Horomones – see Page 648, Table 33-1 Auxins = involved in plant-cell elongation, apical dominance, and rooting Gibberellins = stimulates elongation growth Ethylene = responsible for ripening of fruit (hang green bananas above apples on counter…bananas ripen; ripen peaches in brown paper bag b/c they release ethylene and stimulate ripening) Cytokinins = promote cell division in plants (remember “cytokinesis”) Abscisic Acid = inhibits other hormones (causes closure of plant’s stomata in response to drought) Plant Movement Also referred to as “gravitropism” Chemotropism = a plant’s growth response to a chemical change (growth of pollen tube) Nastic Movements Photoperiodism Plants response to changes in the length of days and nights Critical Night Length = plant’s specific requirement for darkness; regulates flowering SDP = short-day plants; flowers when the days are short and the nights are longer LDP = long-day plants; flowers when the days are long and the nights are shorter DNP = day-neutral plants; not affected by day length Phytochrome = bluish, light sensitive pigment that allows plants to monitor changes in day length Vernalization Low-temperature stimulation of flowering Important for fall-sewn grain crops Farmers often use this to grow and harvest their crops before a summer drought sets in Fall colors Caused by photoperiodic response and by temperature response Longer nights = leaves stop producing chlorophyll…chlorophyll degrades and carotenoids become more visible (hidden by more abundant chlorophyll