The History of Life Chapter 17 Fossils Preserved traces and remains of ancient life. The Fossil Record Paleontologists study ancient life through fossils. They make inferences about past life and group similar organisms together to create the fossil record. Fossils show that more than 99% of all living species that have ever existed have become extinct. How Fossils Form Is the fossil record complete? Many more organisms die without leaving traces than those that do. Most fossils form in sedimentary rock. Figure 17-2 Formation of a Fossil Section 17-1 Water carries small rock particles to lakes and seas. Dead organisms are buried by layers of sediment, which forms new rock. The preserved remains may later be discovered and studied. Interpreting Fossil Evidence Relative Dating – Estimates a fossil’s age by comparing to other fossils present in the same layer. Index fossils identify a particular era Absolute (Radioactive) Dating – A more exact method of determining a fossil’s age by determining the percent of a radioactive element left in the sample. Relative Dating Compare/Contrast Table Section 17-1 Comparing Relative and Absolute Dating of Fossils Relative Dating Absolute Dating Can determine Age of fossil with respect to another rock or fossil (that is, older or younger) Age of a fossil in years Is performed by Comparing depth of a fossil’s source stratum to the position of a reference fossil or rock Determining the relative amounts of a radioactive isotope and nonradioactive isotope in a specimen Imprecision and limitations of age data Difficulty of radioassay laboratory methods Drawbacks Geologic Time Scale Represents evolutionary time. Precambrian time - before complex life Eras – after Precambrian time. Paleozoic – fish and amphibiams Mesozoic – age of dinosaurs Cenozoic – age of mammals Geologic Time Scale with Key Events Section 17-3 Era Cenozoic Mesozoic Paleozoic Precambrian Time (millions of Period Time years ago) Quaternary Tertiary Cretaceous Jurassic Triassic Permian Carboniferous Devonian Silurian Ordovician Cambrian 1.8–present 65–1.8 145–65 208–145 245–208 290–245 363–290 410–363 440–410 505–440 544–505 650–544 Key Events Glaciations; mammals increased; humans Mammals diversified; grasses Aquatic reptiles diversified; flowering plants; mass extinction Dinosaurs diversified; birds Dinosaurs; small mammals; cone-bearing plants Reptiles diversified; seed plants; mass extinction Reptiles; winged insects diversified; coal swamps Fishes diversified; land vertebrates (primitive amphibians) Land plants; land animals (arthropods) Aquatic arthropods; mollusks; vertebrates (jawless fishes) Marine invertebrates diversified; most animal phyla evolved Anaerobic, then photosynthetic prokaryotes; eukaryotes, then multicellular life Earth’s Early History Ancient earth was hostile. Poisonous atmosphere ; no oxygen Oceans were a “hot thin soup” UV radiation, lightening, volcanos… So how did life begin? Figure 17-8 Miller-Urey Experiment Section 17-2 The first organic molecules… Mixture of gases simulating atmospheres of early Earth Stanley Miller recreated early earth’s atmosphere Made simple organic molecules (amino acids…) Spark simulating lightning storms Condensation chamber Water vapor Cold water cools chamber, causing droplets to form Liquid containing amino acids and other organic compounds Coacervates Proteinoid microspheres may have formed in shallow pools as precursors to cells. Evolution of RNA and DNA Stromatolites Ancient prokaryotes added oxygen to the atmosphere. The rise of oxygen caused drove some life forms to extinction while others evolved. Figure 17-12 Endosymbiotic Theory Endosymbiotic Theory – origin of eukaryotic cells Chloroplast Aerobic bacteria Ancient Prokaryotes Nuclear envelope evolving Plants and plantlike protists Photosynthetic bacteria Mitochondrion Primitive Photosynthetic Eukaryote Ancient Anaerobic Prokaryote Primitive Aerobic Eukaryote Animals, fungi, and non-plantlike protists This theory proposes that eukaryotic cells arose from symbiotic relationships between bacteria and cells. Sexual Reproduction and Multicellularity This ancient jellyfish was an early multicellular animal. With the advent of sexual reproduction the rate of evolution took off. Evolution of Life Early Earth was hot; atmosphere contained poisonous gases. Earth cooled and oceans condensed. Simple organic molecules may have formed in the oceans.. Small sequences of RNA may have formed and replicated. First prokaryotes may have formed when RNA or DNA was enclosed in microspheres. Later prokaryotes were photosynthetic and produced oxygen. An oxygenated atmosphere capped by the ozone layer protected Earth. First eukaryotes may have been communities of prokaryotes. Multicellular eukaryotes evolved. Sexual reproduction increased genetic variability, hastening evolution. Heterotroph Hypothesis The first cells were… Heterotrophs autotrophs Anaerobic aerobic Unicellular multicellular Asexual sexual Patterns of Evolution Extinction The last member of a species dies….(failure of a species to adapt) Patterns of Evolution Adaptive radiation An ancestral form evolves into diverse forms through natural selection Patterns of Evolution Convergent evolution Unrelated species evolve similar adaptations due to similar environments Patterns of Evolution Coevolution Evolution of two different species in response to each other: symbiosis Patterns of Evolution Gradualism – A slow and steady rate of evolution Patterns of Evolution Punctuated equilibrium – Periods of rapid evolution followed by long stretches of stability Flowchart Section 17-4 Species that are Unrelated form Related in under under in in Inter-relationshiops Similar environments Intense environmental pressure Small populations Different environments can undergo can undergo can undergo can undergo can undergo Coevolution Convergent evolution Extinction Punctuated equilibrium Adaptive radiation