Tissues Chapter 4 Tissue • a group of similar cells working together to perform a set of functions Plant Tissues • • • • • Meristems Simple (Ground) Tissue Vascular Tissue Dermal Tissue Secretory Tissue Meristems • Apical Meristems – located at the ends of shoots, roots and in buds; produce primary growth – protoderm, ground meristem and procambium • Lateral Meristems – located around the perimeter of shoots and roots; produce secondary growth Primary and Secondary Growth Your growth, typical of animals infant child You, today Plant Growth 1st year – all primary growth Primary growth – plant growth that produces an increase in length and new structures Secondary growth – plant growth that produces an increase in diameter in existing plant parts in the second and any subsequent growing seasons 2nd year – only secondary growth, in blue, is shown below 2nd year primary (in green) and secondary (in blue) growth 3rd year’s growth, showing both primary (black) and secondary (red) growth Woody dicot stem cross section showing growth rings Apparent increase in limb height on many trees is due to loss of lower limbs Lateral Meristems (cont.) • Vascular Cambium – produce vascular tissue • Cork Cambium – produces “bark” Intercalary Meristems • located near grass nodes • produces an increase in stem and leaf length • contributes to the grazing and cutting tolerance of grasses Simple (Ground) Tissues • Parenchyma • Collenchyma • Sclerenchyma Parenchyma • relatively spherical and thin walled • alive at functional maturity • perform a variety of functions – photosynthesis (chlorenchyma) – food and water storage – pith – wound healing Collenchyma • elongate with irregularly thickened cell walls • provide flexible support; celery strings • alive at functional maturity Sclerenchyma • elongate (fibers) to irregular shape (sclerids) • very thick, lignified cell walls • dead at functional maturity Vascular Tissue • Complex tissues since they include 2 or more kinds of cells • Xylem • Phloem Xylem • similar to straws • dead at functional maturity • vessels – composed of vessel elements (cells); uniform tubes, open at both ends, relatively large diameter, may be relatively short • tracheids – tapered at both ends, numerous pits rather than large openings between adjacent cells • conducts water and minerals upwards Xylem Phloem • alive at functional maturity • consist of sieve tube members and companion cells Sieve tube members – cylindrical, conduct organic molecules up and down through plant Companion cells – regulate activity of sieve tube members Phloem Phloem Xylem Dermal Tissue • Epidermis – covers primary growth, single layer of cells; often secretes cutin (lipid) forming cuticle (reduces transpiration and protection from pathogens) • Periderm – covers secondary growth, many cell layers; becomes impregnated with lipids (suberin) Epidermis (cont.) • Root hairs - single cell extensions from the epidermis near root tips • greatly increase surface area in contact with soil and thereby enhance water and mineral absorption Stomata • openings in epidermis • surrounded by Guard Cells that regulate opening Stomata from a epidermal peel of Commelina communis by Dr. J. Weyers Stomate Guard Cells Lenticels • regions of loosely packed cells in the periderm • promote gas exchange with tissues below Secretory Tissue • • • • • • • produce a variety of products nectar in flowers oils in citrus and mints latex resins opium mucilage – e.g. for trapping insects