Plant Physiology Roots, Stems, Leaves and Flowers Plant Physiology Physiology – understanding how parts of an organism work We will learn how roots, stems, leaves and flowers all work together to keep a plant alive Plant Identification There are two main categories of angiosperms. Monocots – grasses, grain crops, lilies, gladiolas, and palm trees Monocots Mono = Single Cot = Short for cotyledon Cotyledon = embryonic leaf • Embryo = developing plant offspring located in the seed Monocots Have four distinct features Veins in leaves are parallel to each other Flower parts are arranged in groups of three Xylem and phloem are arranged in bundles Single embryonic leaf Vascular System Xylem Phloem Transports water throughout the plant Transports food (sugars) throughout the plant Cambium Layer of cells that creates new xylem and phloem Monocot Vein arrangement Convallaria majalis (lily of the valley) The major veins of monocot leaves are generally arranged parallel to each other along the length of the leaf blade. Plant Identification Dicots - most of the other plants such as the shrubs, trees, and flowers. • Veins in leaves are branched • Flower parts are arranged in groups of four or five • Xylem and phloem are arranged in layers • Two embryonic leaves Dicot Leaf Venation Helianthus (sunflower) The major veins of dicot leaves are generally arranged in a netted (reticulate) pattern that extends across and down the leaf. Monocot and Dicot Leaf Venation Convallaria majalis (lily of the valley) Monocot Parallel Leaf Venation Helianthus (sunflower) Dicot Netted Leaf Venation Monocot Flower Parts sepal petal stigma stamen Trillium In most monocots, the flower parts are arranged in multiples of three. This trillium flower has three sepals, three petals, six stamens, and three stigmas on the pistil. Dicot Flower Parts petal stamen pistil Cydonia oblonga (quince) In most dicots, the flower parts are arranged in multiples of four or five. These quince flowers have five petals, twenty stamens, and five pistils. The stamens are too numerous to count in this image. Monocot and Dicot Flower Parts petal sepal petal stamen stigma stamen pistil Trillium Monocot Flower Parts in Multiples of Three Cydonia oblonga (quince) Dicot Flower Parts in Multiples of Four or Five Roots Functions of a root Water Absorption • Most water enters the plant through the roots Anchorage • Holds the plant in one place Reproduction • Some roots allow for asexual reproduction Food Storage • Store sugars for later use Roots The type of root will normally help you identify the plant. It will place the plant into a monocot or dicot category. Types of Roots Tap Root Have a main central root and may have some lateral branching E.g. Carrots e.g. = exempli gratia = for the sake of example Tap Roots Penetrate the soil to various depths - some only a few inches, others like the mesquite to as deep as 114 ft. Tap Roots Benefits of a tap root Access deep water Hold plant in more securely Store larger quantities of sugars Types of Roots Fibrous Have many roots of equal size and a lot of lateral branching Fibrous roots are generally much more dense and closer to the surface Types of Roots This root system can effectively prevent any other plant from becoming established – e.g.: grasses idea of a healthy lawn is to compete with weeds Types of Roots Prop Roots augment regular roots for anchorage aid ex: corn - roots come out above soil and help hold plant up Parts of Roots – Outer layer of cells, protecting inner cells Root Hairs – Small hairs that grow from the epidermis, helping water absorption and holding root in place Epidermis Parts of Roots – Area of storage in the root Cortex Cylinder – area that contains xylem and phloem tissues Vascular Parts of a root Root Cap – Layer of cells that protect the growing area of a root Apical Meristem – Meristem – point of new cell development Apical – Tip (end of a branch) Why Different Types of Roots All plants are in competition with each other for water and nutrients By having different types of roots, the plants can reach different depths in the soil and still live side by side with other plants Stems Stems For identification: type of stem (woody or herbaceous), monocot or dicot, Monocot stem phloem xylem Dicot stem cortex Primary vs Secondary growth Primary Growth Growing from a specific location • In woody stems, often at the ends of the branches (apical meristem) • In herbaceous stems, can be at end or at the surface of the soil (e.g. grasses) Secondary growth • Growing wider around a trunk Mature Structure of Woody vs Herbaceous Stems Herbaceous stems Lack secondary growth plants only live above ground during the growing season • Annuals – plants that live only one growing season Mature Structure of Woody vs Herbaceous Stems Stems remain soft and flexible. Buds lack protective scales (don’t need to survive harsh conditions) Mature Structure of Woody vs Herbaceous Stems Woody stems Plants living and growing over multiple seasons have secondary growth (xylem, phloem) increasing diameter of the stems Mature Structure of Woody vs Herbaceous Stems – Die at the end of the growing season Annuals • Fast, frequent reproduction, with many seeds – Live several growing seasons Perennials • Slower, less frequent reproduction Growing season - The time of the year in which most of the plant’s growth occurs - usually in the summer - can be other times • Wet season • Spring • Fall Specialized Stems Rhizomes - underground horizontal stems (ex: perennial grasses, bamboo) - will grow a plant and roots at a node. Node – Any growth point on a stem or root E.g. ‘eyes’ of a potato. Specialized Stems Stolons - runners - usually above ground, horizontal stems; will grow a plant and roots at a node - ex: strawberries Specialized Stems Tubers - several nodes at the end of a rhizome (ex: potatoes) - eyes are axillary buds – where the tuber will grow a plant Specialized Stems Bulbs - large bud with small stem at lower end - storage in the form of numerous, fleshy leaves - ex: onion, lily, tulip Corms - look like bulbs, but are mostly stem tissue with a few, papery leaves on the outside ex: gladiolus, crocus Leaves Leaves Leaves are where plants conduct photosynthesis to produce most of it’s food. Leaves come in many shapes and sizes Parts of a Dicot Leaf blade – expanded, usually flat portion of a leaf Leaf – connects the blade of a leaf to a stem or branch – holds leaf up for better air flow and to catch the light Petiole Parts of a Dicot Leaf – threads of vascular tissue (xylem & phloem) Node – place on a stem where leaves or branches normally attach Veins Parts of a Leaf Midrib– the primary vein that is seen from the petiole to the tip Spine– Large veins that branch from the midrib Margin– the edge of the leaf blade Base – The lower part of the blade, where the petiole attaches Midrib Tip Spines Blade Margin Veins Petiole Base Parts of a Monocot Leaf – part of leaf that holds leaf to stem – encases stem Ligule – membrane-like tissue extending up from the sheath (on inside) – keeps dirt and moisture out – clear membrane on leaf where attaches to stem Sheath Blade Sheath Monocot Leaf Node Collar Auricle Stem Ligule Picture showing parts of a grass plant. Two Types of Leaves leaves – composed of a single blade and a petiole Simple Two Types of Leaves leaves – are composed of a blade that includes several leaflets and a petiole Compound Two types: Two Types of Leaves Compound – (chestnut) – the lobes or divisions come together and are attached at one place at the base Palmately Leaf Blade Palmately Compound Leaf Petiole Two Types of Leaves Compound – compound leaf with the leaflets on two sides, usually along a central vein – ex: ferns, ash, hickory Pinnately Leaflets Leaf Blade Petiole Pinnately Compound Leaf Leaf Arrangement – have only one type of arrangement – leaf comes off of a node – ex: grasses and grain crops Monocots Leaf Arrangement – flowering plants Alternate – one leaf per node Opposite – two leaves per node Whorled – three or more leaves per node Dicots Leaf Arrangements Arrangement of Veins Four types of vein arrangements: Parallel veins – veins are small and run more or less parallel – most are long and narrow – ex: Buckhorn Plantain, grasses and Iris – mostly monocots Arrangement of Veins veins – are large and small – the small ones connecting to each other to form a net – mostly dicots Netted Arrangement of Veins veined – with one larger midvein and smaller veins coming off along its length – mostly dicots Pinnately Arrangement of Veins Palmately veined- with two or more large veins arising at or near the base of the leaf blade (palm) – leaves are usually broad or fat – mostly dicots Parallel Veins Netted Veins Pinnately Veined Palmately Veined Monocot Leaf -Vein Arrangement Vein Dicot Leaf – Vein Arrangement Smaller lateral vein Midvein Leaf Structure Leaf Structure Epidermis – Tough outside layer of cells that protect the inner cells Divided into Upper epidermis and Lower Epidermis Leaf Structure Upper Epidermis Lower Epidermis Leaf Structure Stomata – Openings in the leaf that allow the passage of CO2 and water Stomata can open and close based on the needs of the plant They will typically stay closed when the plant needs to retain moisture, and will open up when there is enough water available Leaf Structure Stomata open and close due to cells on the edge of the stomata called guard cells Guard cells open and close the stomata by allowing water to enter and leave, causing them to swell or shrink Leaf Structure Upper Epidermis Guard Cells Stomata Lower Epidermis Leaf Structure Palisade Cells Elongated cells where most photosynthesis takes place Why are they elongated vertically, and not horizontally? Leaf Structure Upper Epidermis Palisade Cells Guard Cells Stomata Lower Epidermis Leaf Structure Spongy Mesophyll Water storage area of the leaf Helps to transport wastes out of the leaf through the stomata Leaf Structure Upper Epidermis Palisade Cells Spongy Mesophyll Guard Cells Stomata Lower Epidermis Leaf Structure Vascular System Xylem – Transports water from roots to the leaves (water is necessary for photosynthesis) Phloem – Transports sugars from the leaves to the rest of the plant Leaf Structure Upper Epidermis Palisade Cells Spongy Mesophyll Guard Cells Stomata Vascular System Lower Epidermis Flowers – are highly colored portions of the flower. May contain perfume (rose) or nectar glands –to attract pollinators. Number of petals on a flower is often used in the identification of plant families and genera. Petals Flowers –have sepals and/or petals in multiples of four or five Monocots – have sepals in multiples of threes Dicots Parts of a Flower Sepals Petals Stamens (anther & filaments) Pedicel Pistil (stigma,style & ovaries) Seeds Monocot seeds Contain the embryo (infant plant) Cotyledon (part of the embryo) Seed Coat (protects the embryo) Endosperm (provides nutrients to the embryo ) Radicle – The embryonic root tip Monocot Seed Dicot Seed