UNIT VII - CLASSIFICATION • Formation of Organic Molecules o Oparin/Haldane Hypothesis Primitive Earth’s atmosphere favored chemical rxns that made _____________ compounds from inorganic precursors Reducing/electron adding atmosphere would enhance joining of molecules No _________ because no plants to do photosynthesis Early oceans were an organic “soup” __________________ & UV radiation provided energy for complex organic molecule formation o Miller/Urey Experiment Tested Oparin/Haldane hypothesis Simulated atmosphere composed of water, hydrogen, methane, & ammonia All ____________________________, nitrogen bases, _______________ formed Hypothesis was supported I. EARLY EARTH I. EARLY EARTH • Three Eons 1st two described as ________________ o Archaean Eon & Proterozoic Eon Present day = Phanerozoic eon o Paleozoic era, Mesozoic era, & Cenozic era • Continental Drift Pangaea – “______________” Gondwana & Laurasia • Mass Extinctions • Biogeography Study of past & present distribution of species I. EARLY EARTH II. PHYLOGENY • Phylogeny - _______________ history of an organism • ______________________ - the tracing of evolutionary relationships; includes taxonomy • Linnaean System – ________________ nomenclature – Hierarchical classification • __________________ – taxonomic unit II. PHYLOGENY • ____________________ Diagram showing patterns of __________ characteristics among organisms Homology vs. analogy... o ________________ - likenesses attributed to common ancestry o USED IN CLADOGRAMS o ________________ - likenesses attributed to similar ecological roles and natural selection • Convergent evolution Species from __________________ evolutionary branches that resemble one another due to _________________ ecological roles II. PHYLOGENY III. PROKARYOTES III. PROKARYOTES Classification Methods Domain Archaea – Kingdom Archaebacteria Domain Bacteria – Kingdom Eubacteria Shape ___________ - spherical/round ___________- rod ___________ - helical/spiral Gram Stain Reaction Positive Negative Cell Wall IV. PROKARYOTES - EUBACTERIA _________________________ - consists of polymers of modified sugars cross linked by short polypeptides that vary ________________________ have no peptidoglycan Gram _________________ Simpler cell walls _____ of peptidoglycan Peptidoglycan traps ___________ dye of Gram stain Gram __________________ More complex cell walls _______ peptidoglycan Violet dye of Gram stain is easily rinsed; bacteria will turn _______________ Usually more threatening Capsule Adherence & Protection Pili Adherence & Conjugation IV. PROKARYOTES – EUBACTERIA Motility • Flagella • Helical shape Spirochetes • Slime • Taxis Stimulus IV. PROKARYOTES – EUBACTERIA • Have a ___________ region that contains the DNA • May contain _______________, small rings of DNA consisting of a few genes which are separate from the bacterial chromosome • _______________________ - bacteria asexual reproduction • “Sexual reproduction” – not really sexual reproduction……more like modes of genetic transfer _____________________ _____________________ _____________________ • Endospore Bacterial “hibernation” IV. PROKARYOTES – EUBACTERIA Nutrition – 4 Groups • __________________________ Photosynthetic Use light to drive the synthesis of organic compounds Cyanobacteria • __________________________ Oxidize inorganic chemicals for energy Obtain carbon from CO2 • __________________________ Use light to generate ATP Must obtain carbon in an organic form • __________________________ Consume organic molecules for both energy and carbon Saprobes - decomposers Parasites IV. PROKARYOTES – EUBACTERIA • Metabolism o Nitrogen fixation Conversion of atmospheric nitrogen ______ to ammonium ____________ o Oxygen relationships ________________________________ – must use O2 for cellular respiration and cannot grow w/o it ________________________________ – use O2 if it is present but can also grow via fermentation in an anaerobic environment ________________________________ – are poisoned by O2; some live exclusively by fermentation or some extract chemical energy by anaerobic respiration IV. PROKARYOTES – EUBACTERIA Prokaryotic Ecology • Decomposers • Nitrogen Fixation • Symbiosis ___________________________ – both organisms benefit _______________________ – one organism benefits while neither harming or helping another organism _______________________ – one organism benefits at the expense of the host IV. PROKARYOTES – EUBACTERIA Bacterial Pathogenesis • Koch’s Postulates – Criteria for bacterial disease confirmation 1) 2) 3) 4) • Opportunistic Normal residents of host; cause illness when defenses are weakened • Exotoxins Bacterial proteins that can produce disease w/o the prokaryote present (botulism) • Endotoxins Components of gram negative membranes (Salmonella) V. ORIGINS OF EUKARYOTIC CELLS • _______________________________ (AKA “Endosymbiont Theory”) • Margulis • _______________________ and ________________________ were formerly from small prokaryotes living within larger cells V. ORIGINS OF EUKARYOTIC CELLS Support for Endosymbiotic Theory • Similarities between _______________ and the ________________ & ________________ of eukaryotes – Mitochondria & chloroplasts are appropriate ___________ to be descendents of bacteria – Inner membranes of chloroplasts & mitochondria have several _______________ & transport systems that resemble those found on plasma membranes of prokaryotes – Replication of Mitochondria/Chloroplasts • Split similar to ____________________________ – _______________________________ DNA molecule not associated with histones or other proteins – Ribosomes EUKARYOTES VI. KINGDOM PROTISTA • • • • Very diverse All are _____________________________ Mostly _____________________________ Protists are usually classified according to the eukaryotic kingdom the protist is most like and its nutrition mode ___________________ Ingestive Protozoa ___________________ Photosynthetic Algae ____________________ Absorptive Slime Molds VI. KINGDOM PROTISTA Protist Phylogeny VI. KINGDOM PROTISTA, cont Protist Systematics • Diplomonads Lack mitochondria, cell walls Giardia lamblia Trichomonas vaginalis • Euglenoids Most are heterotrophic; may be autotrophic Most lack cell wall, but have a structural protein layer under cell membrane Flagellated Many have “eyespot” Euglena Trypanosoma VI. KINGDOM PROTISTA - Systematics, cont • Alveolates Contain small sacs called alveoli; may help regulate water, ion concentration Dinoflagellates – phytoplankton, also known as “spinning algae” Red Tides Ciliates – Paramecium, Stentor Apicomplexa - all parasitic; lack cilia, flagella Plasmodium Toxoplasma VI. KINGDOM PROTISTA - Systematics, cont • Stramenopila • • Heterokonts – “different flagella” Water molds (decomposers), mildews (parasitic), algae (photosynthetic) Diatoms Photosynthetic; make up most of Earth’s plankton Have glass-like silicon shells Brown Algae Golden Algae VI. KINGDOM PROTISTA - Systematics, cont • Foraminiferans Ca Carbonate shells Dead foraminiferans settle on ocean floor; shells become chalk White Cliffs of Dover • Rhodopyhta Red algae Mostly multicellular • Chlorophyta Green Algae Volvox Spirogyra Chlamydomonas Unicellular; may be colonial Chloroplasts, cell walls of cellulose Gave rise to land plants VI. KINGDOM PROTISTA - Systematics, cont • Amoebozoa All have pseudopods Amoeba May be parasitic Entamoeba histolytica Plasmodial Slime Molds Feeding stage is a multinucleate mass known as a plasmodium Cellular Slime Molds Feeding stage is single amoeboid cell VII. KINGDOM FUNGI VII. KINGDOM FUNGI Characteristics of Fungi • Multicellular (except for yeast) • ___________________ by absorption; releases ___________________that helps the fungus digest the food outside its body • Ecological roles – Decomposers (saprobes) – Parasites – Mutualistic symbionts (lichens) • Hyphae - body filaments – __________________ - network of hyphae that the fungus uses for feeding • ___________ cell walls • Reproduce by ___________ VII. KINGDOM FUNGI, cont Life Cycle QuickTime™ and a Cinepak decompressor are needed to see this picture. VII. KINGDOM FUNGI, cont Classification • Phylum Chytridiomycota Most closely related to protists; considered to be most primitive member of Kingdom Fungi Aquatic fungi • Phylum Zygomycota Food mold; Rhizopus Mycorrhizae Produce spores from zygote surrounded by thick covering called zygosporangium; may remain dormant for months; resistant to extreme conditons VII. KINGDOM FUNGI - Classification, cont • Phylum Ascomycota Sac fungi Yeasts, truffles, morels, Sordaria Produce sexual and asexual spores Ascus – sexual spore Conidia – asexual spore • Phylum Basidiomycota Club fungi Mushrooms, puffballs, bracket fungi Basidiocarp – haploid hyphae fuse, meiosis occurs, formation of basidospores VII. KINGDOM FUNGI, cont Specialized Life Styles • Molds Used to be classified as Deuteromycota or “Imperfect Fungi” No known sexual stage Penicillium • Yeasts Unicellular Reproduce asexually; budding Saccharomyces • Lichens Mutualistic relationship with algae, cyanobacterium Sensitive to air pollution • Mycorrhizae Mutualistic relationship found in 95% of all plants Increases absorptive surface of roots