DNA Replication copyright cmassengale 1 Replication Facts • DNA has to be copied before a cell divides • DNA is copied during the S or synthesis phase of interphase • New cells will need identical DNA strands copyright cmassengale 2 Synthesis Phase (S phase) • S phase during interphase of the cell cycle • Nucleus of eukaryotes S DNA replication takes place in the S phase. phase G1 interphase G2 Mitosis copyright cmassengale -prophase -metaphase -anaphase -telophase 3 DNA Replication • DNA must be copied in order for cell reproduction • The DNA molecule produces 2 IDENTICAL new complementary strands following the rules of base pairing: •Each strand of the original DNA serves as a template for the new strand A-T, G-C DNA Replication Step One - DNA Helicase – Travels down the DNA molecule and “unzips” it by breaking hydrogen bonds between base pairs. The Y-shaped junction of the unwound molecules is a replication fork. . DNA Replication • • Step Two: DNA polymerase can then add the new nucleotides to form a new strand of complimentary DNA attached to the split apart strand DNA polymerase can only travel down DNA is a 5’ to 3’ direction, creating a leading strand of DNA and lagging strand 6 Remember DNA Strands are Antiparallel – So both DNA Polymerases will be travel in opposite directions 5 while adding nucleotides, one will have an easy path, the other will have to travel backwards. O 3 P 5 O C G 1 3 O P 5 3 2 4 4 P 5 P 2 3 1 O T A 3 O 3 5 O copyright cmassengale 5 P P7 DNA Replication Copyright Pearson Prentice Hall Synthesis of DNA Strands • The Leading Strand is synthesized as a single strand from the point of origin toward the opening replication fork, it only needs DNA Polymerase to add nucleotides. This is the easy path. 5’ 3’ 5’ Nucleotides DNA Polymerase copyright cmassengale 9 Synthesis of the New DNA • The Lagging Strand is synthesized in chunks, not smoothly like the leading strand, against overall direction of replication • This strand is made in MANY short segments 5’ 3’ 3’ 5’ Lagging Strand copyright cmassengale 10 Lagging Strand Segments • Okazaki Fragments - series of short segments on the lagging strand • Must be joined together by an enzyme DNA Okazaki Fragment RNA Primer 5’ 3’ Polymerase 3’ 5’ Lagging Strand copyright cmassengale 11 copyright cmassengale 12 DNA Lagging Strand • Before new DNA strands can form on the lagging strand, there must be RNA primers present RNA Primase is the enzyme that synthesizes the RNA Primers – gives DNA polymerase a path to follow • After, DNA polymerase adds nucleotides to the lagging strand just like the in the leading strand copyright cmassengale 13 Third Step: • The enzyme DNA Ligase joins the leading strand nucleotides or Okazaki fragments together to make one strand DNA ligase 5’ 3’ Okazaki Fragment 1 Okazaki Fragment 2 3’ 5’ Lagging Strand copyright cmassengale 14 DNA Replication • Semiconservative Model: • From Watson and Crick • New DNA consists of 1 PARENTAL (original) and 1 NEW strand of DNA DNA Template Parental DNA New DNA Short video on Replication • https://www.youtube.com/watch ?v=JZXT2uOcD2w copyright cmassengale 17