COMPARING INVERTEBRATES Chapter 29 Taxonomy • The system we use today to name and classify • all organisms was developed by Carl Linnaeus. It is known as the system of binomial nomenclature because every organism has a two part name. – Ex: Pantera leo – Ex: Homo sapiens • In addition, Linneaeus classified every organism into a hierarchy of taxa, or levels of organization. Taxonomy--Classification • King • Kingdom • Philip • Phylum • Came • Class • Over • Order • For • Family • Great • Genus • Spaghetti • Species Example Classification • Domain • Eukarya • Kingdom • Animalia • Phylum • Chordata • Class • Mammalia • Order • Primate • Family • Hominid • Genus • Homo • Species • sapiens Taxonomy • All life can be organized into three domains: Bacteria, Archaea, and Eukarya. Bacteria • Single celled • • • • • • prokaryotes Aerobes/anaerobes Decomposers Pathogens some are photosynthetic Have no introns Includes viruses Archaea • Single celled • Prokaryotes • Extremophiles – Methanogens – Halophiles – Thermophiles EUKARYA • All have nucleus and • • internal organelles Includes animal and plant cells Consists of 4 kingdoms • KINGDOMS – – – – Protista Fungi Plantae Animalia Kingdom: Protista Kingdom: Fungi Kingdom: Plantae • Multicellular • Nonmotile • Autotrophic • • • • (photosynthetic) Have cell walls Store sugars as starch Alternation of generations Some have vascular tissue Kingdom: Animalia • • • • Multicelluar Heterotrophic Most are motile Most reproduce sexually and are diploid Kingdom: Animalia Animal classification placed into related groups 1. Invertebrates— majority of animals which lack a backbone 2. Vertebrate-animals with a backbone Common Phyla: Porifera Cnidarians Platyhelminthes Nematodes Annelids Mollusks Arthropods Echinoderms Chordates Evolutionary Trends A. Specialized Cells, Tissues, and Organs • As larger and more complex animals evolved, specialized cells joined together to form tissues, organs, and organ systems that work together to carry out complex functions. Evolutionary Trends B. Body Symmetry • Radial symmetry – parts are arranged in a circle around a central point • Bilateral symmetry – parts are mirror images of each other (left and right sides) • Asymmetrical – no definite shape Assymmetrical—Porifera Evolutionary Trends • Whereas primitive animals exhibit radial symmetry, sophisticated animals exhibit bilateral symmetry. Evolutionary Trends C. Cephalization • Along with bilateral symmetry came the development of cephalization, which is the concentration of sense organs and nerve cells in the front (anterior part) of the body. • The digestive, excretory, and reproductive structuress are located at the back (posterior) end. • Invertebrates with cephalization can respond to the environment in more sophisticated ways that can simpler invertebrates. Evolutionary Trends—Cephalization Evolutionary Trends D. Segmentation • Many animals who exhibit bilateral symmetry also have segmented bodies. • Segments have often become specialized for specific functions. • Segmentation allows an animal to increase in size. Evolutionary Trends E. Coelom Formation • Germ layers formed early in embryonic development: – Ectoderm (outermost layer) – Mesoderm (middle layer) – Endoderm (innermost layer) • The coelom is a fluid-filled body cavity that is completely surrounded by mesoderm tissue. • It represents a significant advance in animal evolution because it provides space for elaborate organ systems. Evolutionary Trends—Coelom Formation Types of body cavities: • Acoelomates do not have a coelom (body cavity) between their body wall and digestive cavity. • Pseudocoelomates have body cavities that are partially lined with mesoderm. • Most complex animal phyla are coelomates, meaning they have a true coelom that is lined completely with tissues from mesoderm. Acoelomate Body covering (from ectoderm) Digestive sac (from endoderm) Tissue-filled region (from mesoderm) Pseudocoelomate Body covering (from ectoderm) Muscle layer (from mesoderm) Digestive tract (from endoderm) Pseudocoelom Coelomate Body covering (from ectoderm) Coelom Tissue layer lining coelom and suspending internal organs (from mesoderm) Digestive tract (from endoderm) Coelomate ` Evolutionary Trends F. Embryological Development Blastopore: first opening during the embryonic stages of an organism Evolutionary Trends F. Embryological Development • Protostome – blastopore becomes the mouth, and the anus forms secondarily • Deuterostome – blastopore becomes the anus, and the mouth forms secondarily Trends in Animal Development Echinoderms Chordates Arthropods Annelids Mollusks Radial Symmetry Roundworms Flatworms Cnidarians Radial Symmetry Pseudocoelom Protostome Development Three Germ Layers; Bilateral Symmetry Sponges Tissues Singlecelled ancestor Deuterostome Development Coelom Trends in Animal Development From the Primitive • No symmetry or radial symmetry • No cephalization • 2 germ layers • Acoelomate • No true tissues • Little specialization • Sessile To the Complex • Bilateral symmetry • Cephalization with sensory apparatus • 3 germ layers • Pseudocoelomate or coelomate • Tissues, organs, and organ systems • Much specialization • Motile Form and Function in Invertebrates Ch. 29-2 Feeding and Digestion • The simplest animals break down food primarily through intracellular digestion, but more complex animals use extracellular digestion. • In intracellualar digestion food is digested inside the cells. – The food size must then be smaller than the cells. • In extracelluar digestion, food is broken down outside the cells. – The food size is larger than the cells of the organism. • How does a starfish eat? Patterns of Extracelluar Digestion • Some animals such as cnidarians and most • • • flatworms ingest food and expel wastes through a single opening. Some cells of the gastrovascular cavity secrete enzymes and absorb digested food. Other cells surround food particles and digest them in vacuoles. More complex animals digest food in a tube called the digestive tract, which may have specialized regions such as stomach and intestines. Intestine Gizzard Crop Mouth/anus Pharynx Mouth Gastrovascular cavity Annelid Anus Gastrovascular cavity Cnidarian Pharynx Crop Arthropod Anus Pharynx Mouth Mouth/anus Flatworm Stomach and digestive glands Rectum Respiration: Gas exchange of O2 and CO2 Two key features of all respiratory systems: • Respiratory organs have large surface areas that are in contact with air or water • Have ways to keep the gas exchange surfaces moist to allow diffusion to occur Respiration Tracheal tubes Gill Siphons Movement of water Spiracles Insect Mollusk Airflow Spider Book lung Circulation • In an open circulatory system, blood is only partially contained within a system of blood vessels. Heart (s) Blood vessels Tissues Sinuses (spongy cavities) Circulation • In a closed circulatory system, a heart or a • • • heart-like organ forces blood through vessels that extend throughout the body. The blood stays within these blood vessels. Materials reach body tissues by diffusing across the walls of the blood vessels. Blood circulates more efficiently in a closed circulatory system. Heartlike structure Hearts Small vessels in tissues Blood vessels Heart Sinuses and organs Heartlike structures Insect: Open Circulatory System Blood vessels Annelid: Closed Circulatory System Excretion • The excretory system is responsible for removing waste material and conserving water. • Waste product is usually nitrogenous, meaning it contains nitrogen. • This waste is usually in the form of ammonia (NH3), which is very toxic! Excretion • In aquatic invertebrates, ammonia diffuses from their body tissues into the surrounding water • Terrestrial invertebrates convert: ammonia urea (less toxic) • Some insects and arachnids convert: Ammonia uric acid Excretion Flame cells Flatworm Excretory tubules Nephrostome Excretory pore Flame cell Excretory tubule Nephridia Digestive tract Annelid Arthropod Malpighian tubules Response—Nervous System • The nervous system gathers information from the environment. • The simplest nervous system, found in cnidarians, are nerve nets. Trends in the Evolution of the Nervous System • Centralization—nerve cells are more concentrated (ex: ganglia) • Cephalization—high concentration of nerve cells in the anterior region (head/front) • Specialization—more developed sensory organs – To detect light, sound, chemicals, movement, etc. Arthropod Brain Ganglia Ganglia Brain Nerve Cells Flatworm Cnidarian Mollusk Movement & Support • Most animals use Three main kinds: • skeletons • Exoskeletons • Endoskeletons specialized tissues called muscles to move, breathe, pump blood, and perform other life functions. In most animals, muscles work together with some sort of skeletal system that provides firm support. • Hydrostatic Movement & Support • Hydrostatic skeleton – No hard structures – Lacks muscles – Water filled cavity (gastrovascular cavity) • Exoskeleton or external skeleton – Outside the body – Hard body covering made of chitin – Has to be shed (molting) • Endoskeleton – Structural support inside the body – Muscles Reproduction • Sexual reproduction is the production of offspring from the fusion of gametes. – Maintains genetic diversity because it generates new combinations of genes • Asexual reproduction – – – – – Ex: Fragmentation Ex: Budding All offspring are genetically identical to parent (clones) Allows for organisms to produce offspring faster Genetic diversity decreases less able to deal with changes Reproduction • Some organisms are hermaphrodites, meaning that they produce both sperm and egg. Reproduction Fertilization: unification of sperm & egg • External fertilization – – – – Observed in less complex animals Eggs are fertilized outside the body Gametes are released in surroundings Aquatic environment • Internal fertilization – Observed in more complex animals – Eggs are fertilized inside the female’s body – Require specialized organs