Mesozoic Life Life of the Mesozoic Era • Age of Reptiles – most diverse and abundant land dwellers • Mammals appear • Birds appear • Flowering plants appear • Some marine invertebrate groups recover Pm\Tr • Another big extinction at end K Mesozoic Marine Life • Permo-Triassic extinctions left voids that were rapidly filled. • Several important invertebrate phyla radiated, as did certain fishes • Predator/prey relationships changed • Appearance of marine reptiles at top of marine food chain Just before the Mesozoic: Permian\Triassic Extinctions • Many major Paleozoic invertebrate groups extinct • a) Trilobites • b) Rugose and tabulate corals • c) Blastoids (echinoderms) • d) Fusulinid foraminifera • e) Brachiopods and crinoids severely reduced Replacement radiations • New marine groups assumed ecological roles of extinct organisms • a) New corals and planktonic foraminifera • b) Several classes of mollusks Ammonites radiate Clams replace many brachiopod groups– Rudistid clams become reef formers • Marine Reptiles replace fish as top marine predators The diversification of Mollusks • Bivalves replaced most brachiopods in benthic community • Oysters were successful, abundant in shallow Atlantic and Gulf ocean. • Ammonites underwent a tremendous radiation Rudists • Bizarre rudistid bivalves (clams, oysters) • Shells had large cone-shaped valve, smaller lid valve • Shells up to 1meter long grew in masses, formed reefs • They replaced corals as dominant reefformers by Middle Cretaceous Ammonites • Greatest mollusk diversification was among cephalopods • Nautiloids and ammonoids had appeared in Paleozoic • Ammonites were ammonoid cephalopods that underwent tremendous radiation • They were differentiated on the basis of their complex suture patterns • The evolutionary purpose of the intricately folded septa – Complex infolding sustained shell against great water pressure at depth Cephalopod shell morphology (living today) (extinct end K) Ammonites • Ammonites were rapidly evolving, freeswimming predators • A favorite food of mosasaurs (huge marine lizards) • Among most important Jurassic and Cretaceous index fossils – Rapid evolution of sutures – Widespread, many facies Scleractinian Corals • 1) Scleractinian corals replaced rugose, tabulate corals • (a) First appeared in Middle Triassic • (b) Soft-bodied sea anemones possible ancestors • (c) Scleractinian corals formed large reefs by end of Triassic Success of Scleractinian corals Skeletons faster growing – Crowd out other benthic organisms • Symbiotic relationship with algae (zooxanthelae) responsible (found in fossils) • Algae supply food, oxygen to coral • Coral furnishes home to algae in shallow, sunlit waters Echinoids • Stalked echinoderms largely gone at Paleozoic end. A few crinoids still survive. • Varieties of echinoids flourished in the benthic environment • Sea urchins • Sand Dollars Microfossils Foraminifera • Foraminifera greatly reduced after Perm-Triassic extinctions, radiated again • Benthic until Jurassic Period • Planktonic foraminifera then inhabited surface waters • Thinner-walled, globular–shaped tests • Important index fossils Coccolithophorids • Unicellular gold-brown algae • Calcareous shells consist of microscopic disk-shaped plates • First appearance in Jurassic • Major component of Cretaceous chalk deposits, along with forams • Chalk deposits represent the original reference section that defined the Cretaceous System both sides of the English Channel Mesozoic Fishes • Bony Fishes continued to dominate • Especially Teleosts • Sarcopterygians reduced to a few species Marine Reptiles • Several groups of Triassic diapsid and euryapsid reptiles became successful marine predators • They developed streamlined bodies • Flipperlike modifications of limbs • Large size • Ichthyosaurs • Mosasaurs • Plesiosaurs Mosasaurs - Marine Lizards Diapsids A really big ”Goanna” or Monitor Lizard Lived offshore near here Euryapsid Marine reptiles Mosasaurs (Cretaceous) 1) Giant marine lizards up to 15 meters long 2) Flattened tails and flipperlike limbs 3) Ate fish and cephalopods Plesiosaurs 1) Found in Jurassic and Cretaceous rocks 2) Short bodies, flipperlike limbs, and long necks 3) Ate fish. Up to 12 meters in length Nothosaurs (Tr-J) 1) Contemporary with placodonts 2) Streamlined bodies elongated necks 3) Ancestral to plesiosaurs Icthyosaurs Tr to Late K, long snouts, fishlike bodies like sharks and dolphins; convergent evol. Rapid swimmers Placodonts (Triassic) 1) Short-necked, body < 2m 2) sea floor dwellers 3) Crushed shellfish for food Plus Marine Crocodiles Placodonts Ichthyosaurus Plesiosaurus Mosasaurs marine predators in Cretaceousc, mostly ate fish and ammonites Land Plants Gymnosperms • Cycads – – Cylindrical trunks and large-fernlike leaves Dominant during Jurassic, common until Cretaceous • Conifers – most modern conifer types – dominated Cretaceous forests as cycads declined • Ginkgoes – Common in Mesozoic forests – Single surviving species like Mesozoic ancestors Cycad: Sago palm Ginkgo biloba Cycads and cycadeoids in Jurassic Scenes Angiosperms - Flowering Plants • Most significant Mesozoic evolutionary event • Reproductive adaptations out compete gymnosperms • Flower uses color and scent to encourage insect pollination • Higher pollination success rates than gymnosperms that use wind • Manufacture of seeds with a food supply • Animals became important in distributing seeds from fruits Demo, cones and Magnolia seed sheath after flowers wilt. Flowering Plants - Primary Producers on land • Triassic and Jurassic land plants consisted of seedless vascular plants and gymnosperms with seeds. • Angiosperms (flowering plants) appear in Early Cretaceous, similar to magnolias SAYREVILLE • The evolution of flowers and an enclosed seed ensured their success • No more relying on wind Mammal-Like reptiles • Therapsid reptiles recovered from PermoTriassic extinctions • Dicynodonts were common until end of Triassic – Gondwana Lystrosaurus found on all southern continents • Cynodonts radiated in Early Triassic • Gondwana carnivorous Cynognathus found in Africa and South America • Herbivorous cynodonts lived until Middle Jurassic Triassic Therapsids Lystrosaurus Thrinaxodon Mesozoic Therapsids Origin of Mammals • Therapsids gave rise to cynodonts, which evolved into the mammal class – skeletal structure is used to identify mammals in the fossil record – differences in the lower jaw and ear in particularly distinguish mammals Late Triassic Morganucodon Mammaliaformes descended from Cynodonts ~ 200 mya • Mesozoic groups: multituberculates, • monotremes, • marsupials and placentals Was looking for early humans Roy Chapman Andrews Protoceratops In the Gobi desert of Mongolia, found the first dinosaur nests and evidence of parental care in the dinosaurs Protoceratops Modern studies of Dinosaurs still test his theories Discussion: Oviraptor Dinosaur Ancestry • Archosaurs, reptiles that radiated in Permian and especially Early Triassic • Large quadrupedal “galloping” crocodile-like forms. Hind limb much longer. • Small bipedal forms probably ancestral to dinosaurs and birds Basal Archosaurs Herrerasaurus Common fossil near campus Rutiodon Phytosaurs Archosaurs and the Origin of Dinosaurs • Archosaurs gave rise to crocodiles, pterosaurs, dinosaurs, and birds • Dinosaurs two groups distinguished by hips: Saurischian Hip – Saurischia – Ornithischia Ornithischian Hip First dinosaur found: Iguanodon Mary Anning •First dinosaur discovered in 1822 •1842 Richard Owen “dinosaur” •Dinosaurs are terrestrial, so they are rare fossils •Hypsilophodon Dinosaurs Saurischian Back of hip this side Ornithischian Front of hip this side Dinosaurs • Saurischians – theropods were bipedal carnivores – sauropods were the giant, quadrupedal herbivores • Ornithischians – Ornithopods (duckbill) – Pachycephalosaurs – Ankylosaurs – Stegosaurs – Ceratopsians Saurischian Dinosaurs • Theropods, the carnivorous dinosaurs • And Sauropods, the long necked giants Theropods Dilophosaurus Larger relative of Velociraptor Dennis Neary Trenton NH Museum Theropods Sauropods Sauropods Parental Care, Herds, Land Ornithischian Dinosaurs • Stegosaurs (plate-backed dinosaurs) • Ornithopods including Hadrosaurs (duckbills) • Ankylosaurs (armored dinosaurs) • Pachycephalosaurs (thick-headed dinosaurs) • Ceratopsians (horned dinosaurs) Ornithischians Social behavior in duckbill dinosaurs Colony nesting also known in Protoceratops Roy Chapman Andrews Maiasaura Jack Horner Parasaurolophus Crest dimorphism, function Armored dinosaurs Pachycephalosaur Ceratopsians Flying Reptiles • The pterosaurs were the first vertebrates to fly – common from Late Triassic to Cretaceous – wing membrane supported by an elongated fourth finger – light hollow bones – development of brain areas associated with coordination and sight – likely to have been endothermic Principle pterosaur groups Discussion: stability and steering in flight Pitch, Yaw and Roll Rhamphorynchoids Long keeled tail, teeth Pterodactyloids No tail, strong crest Some have fine “hair” Jurassic Bird Archaeopteryx Archeopteryx lithographica Late Jurassic, Solenhofen Germany Coelurus The K\T ash layer in Alberta Shocked Quartz Hell Creek Formation Luis and Walter Alvarez Cretaceous-Tertiary boundary The Chicxulub structure K-T Mass Extinction A Crisis in the History of Life Cretaceous-Tertiary extinction claimed dinosaurs, flying reptiles, marine reptiles, and many marine invertebrates Dust cloud reached stratosphere Blocked the sun Plants need light Herbivores eat plants Carnivores eat herbivores Hibernating survivors can sleep through it. Uh, oh. That can’ t be good. The End of the Age of Reptiles ZZZZZZZZZ ZZZZZZZZZ ZZZZZZZZZ Meanwhile, mammals able to hibernate could sleep through the cold dark crisis, getting up occasionally for a supper of stored seeds and underground insects and worms