Light in Lakes Light is energy Major energy source to aquatic habitats Productivity controlled by energy used in photosynthesis Thermal character of lake determined by solar energy Light is energy Solar radiation Capacity to do work Can be transformed into other energy forms Light from the sun Pulsating field of force, endless series of waves Packets of energy photons Energy proportional to frequency (high-high), inversely to wavelength (highshort) Light from the sun Mixture of wavelengths, energies Most (50%) striking lake surface is infrared, visible (especially red part of spectrum) Light from the sun Amount striking lake surface dependent on: Latitude Season Time of day Altitude Meteorological conditions Light and atmosphere Light absorbed by particles in atmosphere Less atmosphere to pass through, more light makes it to earth - angle of incidence Shorter wavelengths selectively absorbed by O2, ozone, H2O vapor, CO2 Red sky at dawn, dusk Indirect Light Some solar radiation reaches lake indirectly Scattered light Light scattered as it passes through atmosphere (20%) Mostly UV and short wavelength visible (blue) Indirect Light Importance of indirect light changes with angle of incidence Contribution of indirect small when sun directly overhead Contribution significant (~20-40%) when sun low in sky Reflected Light Significant fraction of light striking lake surface may be reflected Amount increases with decreased angle of incidence Wave action increases reflection only at low angles of incidence Other Losses of Light Reflection comprises ~1/2 of light lost from water Remaining half lost by scattering Deflection by water molecules, dissolved substances, suspended particles Varies with depth, season, particle loading Lake Color Scattering and absorption of light give lake part of its characteristic color Clean water - blue color More and bigger particles scatter longer wavelengths and absorb shorter wavelengths Blue-green, green, yellow Light Attenuation Radiant energy diminished with depth Results from both scattering and absorption Absorption - loss of solar energy with depth by its transformation to heat Light Attenuation In distilled water lake, >1/2 of light energy transformed into heat with first 1 meter Light Attenuation Absorption not same for all wavelengths Longer wavelengths more readily absorbed than shorter wavelengths Light Attenuation Light Attenuation Few distilled water lakes Dissolved, suspended stuff affects absorption Less absorption, greater transmittance in clear, unproductive lakes than in productive, murky waters Light Attenuation Blues disappear, greens penetrate, reds change with productivity Transmission drastically affected by cover of cloudy ice, snow Shuts down photosynthesis, reduces O2 supply Euphotic Zone Region from surface to depth at which 99% of the surface light has disappeared Minimum intensity of subsurface light that permits photosynthesis is ~1% of incident surface light Water Transparency Measuring light penetration before instrumentation Secchi disk Depth at which disk disappears/reappears from/to sight Water Transparency Secchi disk transparency X 3 used as a “rule of thumb” estimate of depth of euphotic zone Highly variable (e.g., Lake Erie 5X) Heat & Density Layering Light to Heat Loss of light = gain in heat Should temperature profile parallel light profile? No Light to Heat Uniformly mixed layer of water near surface of same temperature Often extends below euphotic zone Mixing of upper layers of water by wind distributes heat downward Direct Thermal Stratification Lighter, warmer layer overlying denser, cooler layer Lake divided vertically into 3 regions Epilimnion Metalimnion Hypolimnion Direct Thermal Stratification Epilimnion - uniformly warm layer mixed by wind Direct Thermal Stratification Hypolimnion uniformly cool lower layer unaffected by wind Direct Thermal Stratification Metalimnion intermediate zone where temperature drops rapidly with increasing depth Also referred to as thermocline - plane between two depths between which temperature change is greatest A Thermally Stratified Lake Temperature (°C) 5 1 10 15 20 25 30 Epilimnion 2 Depth (m) 3 4 5 Metalimnion Thermocline 6 7 8 9 10 Hypolimnion Two separate water masses between which there is little mixing Epilimnion Upper Layer Warm Well mixed THERMOCLINE Hypolimnion Lower layer Cooler than epilimnion STABILITY OF THERMAL STRATIFICATION Stability—likelihood that a stratified lake will remain stratified. This depends on the density differences between the two layers. Examples: Epilimnion 8°C 22°C Hypolimnion 4°C 7°C 30°C 28°C Result Not much density difference Large density difference, Strong stratification Large density difference, Strong stratification (tropical lakes) Even a Hurricane Can’t Break Stratification Temperature (oC) 0 6 8 10 12 14 16 18 20 3 Depth (m) 5 8 10 13 15 After hurricane Before hurricane Thermal resistance to mixing Why do lakes stratify? (1) Density relationships of water Less dense water “floats” on deeper water (2) Effect of wind Molecular diffusion of heat is slow Wind must mix heat to deeper water How do lakes stratify? Temperature (°C) Example: 10 m deep lake in Lake County, IL 5 1 (1) Early Spring 2 3 No density difference Depth (m) No resistance to mixing 4 Heat absorbed in surface water is distributed throughout 5 6 7 8 9 10 10 15 20 25 30 Spring Turnover—time of year when entire water column is mixed by the wind Duration of spring turnover depends on the surface area to maximum depth In very deep lakes, the bottom water stays at 4°C, in more shallow lakes, can get up to > 10°C. Can last a few days or a few weeks. How do lakes stratify? Temperature (°C) (2) Mid Spring 5 1 Longer and warmer days mean more heat is transferred to the surface water on a daily basis 3 4 Depth (m) Surface waters are heated more quickly than the heat can be distributed by mixing 2 5 6 7 8 9 10 10 15 20 25 30 This increase in surface waters relative to the rest of the water column often occurs during a warm, calm period Now have resistance to mixing. Hypolimnion water temperature will not change much for the rest of the year. How do lakes stratify? Temperature (°C) (3) Late Spring 1 2 3 4 Depth (m) With the density difference established, the epilimnion “floats” on the colder hypolimnion 5 5 6 7 8 9 10 10 15 20 25 30 How do lakes stratify? Temperature (°C) 5 (4) Late Summer 1 The epilimnion has continued to warm 2 3 Strong thermal stratification Depth (m) In very clear lakes, can get direct hypolimnetic heating 4 5 6 7 The decomposition of dead plankton may result in loss of oxygen from the hypolimnion 8 9 10 10 15 20 25 30 How do lakes stratify? Temperature (°C) (5) Early Autumn 5 Heat is lost from the surface water at night 1 2 3 4 Depth (m) Cool water sinks and causes convective mixing 5 6 7 Thermocline deepens and epilimnion temperature is reduced 8 9 10 10 15 20 25 30 How do lakes stratify? Temperature (°C) (5) Mid-late Autumn 5 As epilimnion cools, reduce density difference between layers 1 2 3 Eventually, get “Fall Turnover” Depth (m) 4 5 6 7 8 Turnover returns oxygen to the deep water and nutrients to the surface water 9 10 10 15 20 25 30 How do lakes stratify? Temperature (°C) (7) Winter 5 Surface water falls below 4°C and “floats” on 4°C water 1 2 3 Ice blocks the wind from mixing the cooler water deeper Depth (m) 4 5 6 7 8 Get “inverse stratification” 9 10 10 15 20 25 30 5 14 2 6 7 8 18 17 11 10 ice 16 9 4 7 11 3 15 8 0 2 4 6 8 10 12 14 10 13 4 Apr May Jun Jul Aug Sep Oct Temperature 0 0 5 10 15 20 2 Depth (m) Depth (m) Seasonal Stratification in a Temperate Lake 0 5 10 15 20 0 Nov Dec Feb Mar (0C) 5 10 15 20 Direct 4 Jan 0 5 10 15 20 Inverse 6 8 10 12 14 Spring mixing Summer stratification Fall mixing Winter stratification Dimictic Lakes Complete circulations (turnovers) in spring and fall separated by summer thermal stratification and winter inverse stratification Very common in temperate regions Many other types based on circulation patterns Mixing Patterns 1. Amictic—never mix because lake is frozen. Mostly in Antarctica. Some in very high mountains. 2. Holomictic—lakes mix completely (top to bottom) 3. Meromictic—Never fully mix due to an accumulation of salts in the deepest waters. Holomictic: lakes are classified by the frequency of mixing Monomictic lakes: one period of mixing - Cold - Warm Dimictic lakes: two periods of mixing and two periods of stratification Polymictic lakes: mix many times a year - Cold - Warm Holomictic: lakes mix completely Cold monomictic lakes — one period of mixing Frozen all winter (reverse stratification) Mix briefly at cold temperatures in summer Arctic and mountain lakes Kalff 2002 Meretta Lake, CA Holomictic: lakes mix completely Kalff 2002 Warm monomictic lakes — one period of mixing Thermal stratification in summer Does not freeze, so mixes all winter Lake Kinneret Holomictic: lakes mix completely Dimictic—two periods of mixing and two periods of stratification Freeze in winter (inverse stratification) Thermally stratify in summer Wetzel 2001 Holomictic: lakes mix completely Cold polymictic lakes — mix many times a year Ice covered in winter, ice free in summer May stratify for brief periods during the summer, but stratification is frequently interrupted Shallow temperate lakes (< ~20 m) with large surface area mountain or arctic lakes Holomictic: lakes mix completely Warm polymictic lakes — mix many times a year Never ice covered Tropical lakes May stratify for days or weeks at a time, but mixes more than once a year Mixing Patterns 1. Amictic—never mix because lake is frozen. Mostly in Antarctica. Some in very high mountains. 2. Holomictic—lakes mix completely (top to bottom) 3. Meromictic—Never fully mix due to an accumulation of salts in the deepest waters. Meromictic: lakes are chemically stratified Thermocline Chemocline Monimolimnion Meromictic: lakes are chemically stratified Recall that salinity increases density The water in the monimolimnion does not mix with the upper water The mixolimnion can have any mixing pattern (e.g., dimitic, monomictic) Can get interesting thermal profiles Warmer water below colder water above 4ºC Recall salinity increases density Mixing Patterns 1. Amictic—never mix because lake is frozen. Mostly in Antarctica. Some in very high mountains. 2. Holomictic—lakes mix completely (top to bottom) Monomictic lakes: Cold / Warm Dimictic lakes: Polymictic lakes: Cold / Warm 3. Meromictic—Never fully mix due to an accumulation of salts in the deepest waters. Geographic Distribution All of these classification patterns are for lakes that are deep enough to form a hypolimnion “Shallow” lakes do not form a hypolimnion and are therefore unstratified. They have similar temperatures top to bottom. What is meant by “shallow” and “deep enough” is determined by the fetch and depth A lake with a maximum depth of 4m can stratify if it is in a protected basin Bullhead Pond Surface Area = 0.02 km2 Maximum fetch < 300 m 22 August 1993 A lake with a maximum depth of 12m can be unstratified if the fetch is long enough Te mpe r a t ur e ( C) 0 0 2 Oneida Lake, NY Surface Area = 207 km2 Maximum fetch = 33 km Depth (m) 4 6 8 10 12 5 10 15 20 25 30