CNBH, PDN, University of Cambridge Part II: Lent Term 2015: ( 1 of 4) Central Auditory Processing Roy Patterson Centre for the Neural Basis of Hearing Department of Physiology, Development and Neuroscience University of Cambridge email rdp1@cam.ac.uk Lecture slides on CamTools: Lecture slides, sounds, videos and background papers on http://www.pdn.cam.ac.uk/groups/cnbh/teaching/lectures/ CNBH, PDN, University of Cambridge Contents/Progress Act I: the information in communication sounds (animal calls, speech, musical notes) Act II: the perception of communication sounds (the robustness of perception) Act III: the processing of communication sounds: How [Tune7nCPHtone.mov] (auditory signal processing) axial axial Act IV: the processing of communication sounds: Where (auditory anatomy, physiology, brain imaging) CNBH, PDN, University of Cambridge Act I The form of communication sounds including speech and musical notes http://www.pdn.cam.ac.uk/groups/cnbh/teaching/lectures/PSDWshar08.pdf http://www.pdn.cam.ac.uk/groups/cnbh/teaching/lectures/PGW_SHAR_2010.pdf CNBH, PDN, University of Cambridge Sounds used to communicate at a distance, to declare territories and attract mates, are typically Pulse Amplitude Pulse-Resonance Sounds Time The pulse marks the start of the communication. The resonance provides distinctive information about the shape and size of resonators in the sender’s body. CNBH, PDN, University of Cambridge The pulses are produced in streams, each with a resonance source-filter source filter systems : pulse-resonance sounds The streams have distinctive onsets and offsets, and a stream with its onset and offset is referred to as a syllable. http://www.pdn.cam.ac.uk/groups/cnbh/teaching/lectures/PSDWshar08.pdf CNBH, PDN, University of Cambridge Fish Frog 400 ms Macaque Human Patterson, Smith, van Dinther and Walters (2008). Communication ‘syllables’ of four different animals mammalian vocal apparatus CNBH, PDN, University of Cambridge nose lips filter chin tongue glottal folds vocal tract source CNBH, PDN, University of Cambridge waveform and spectrum of a child’s /a/ pulse rate resonance shape envelope shape fine structure CNBH, PDN, University of Cambridge What happens as a child grows up into an adult? At age 3, average height is about 85 cm When adult, average height is about 170 cm As we double in height, our vocal tracts double in length http://www.pdn.cam.ac.uk/groups/cnbh/teaching/lectures/PSDWshar08.pdf CNBH, PDN, University of Cambridge Patterson, Smith, van Dinther and Walters (2008). Low Long Pitch VTL High Short Time Time CNBH, PDN, University of Cambridge waveform and spectrum of a child’s /a/ pulse rate resonance rate position of the envelope Sf Ss position of the fine-structure Frequency on a logarithmic axis (octaves) CNBH, PDN, University of Cambridge Decreasing VTL (1/Sf ) The role of Ss and Sf in speaker perception Increasing GPR ( Ss ) Kawahara and Irino (2004). Principles of speech manipulation system STRAIGHT. In Speech separation by humans and machines, P. Divenyi (Ed.), Kluwer Academic, 167-179. Rana catesbeiana Decreasing VTL (1/Sf ) CNBH, PDN, University of Cambridge Increasing GPR ( Ss ) CNBH, PDN, University of Cambridge The role of Ss and Sf in instrument perception Instrument Register Instrument Family Ss and Sf Envelope Shape http://www.pdn.cam.ac.uk/groups/cnbh/teaching/lectures/PGW_SHAR_2010.pdf CNBH, PDN, University of Cambridge Acoustic scale in instrument perception viola cello violin Instruments with same shape and construction, sound similar. Envelope Shape CNBH, PDN, University of Cambridge The perception of musical notes and instruments Instrument Register Instrument Family Ss and Sf Envelope Shape CNBH, PDN, University of Cambridge pulses van Dinther and Patterson (2004) Waveforms for trumpet and trombone resonance Time CNBH, PDN, University of Cambridge The effect of Ss and Sf on instrument perception Decreasing Size (1/Sf ) French Horn Pitch (Ss) Kawahara and Irino (2004). Principles of speech manipulation system STRAIGHT. In Speech separation by humans and machines, P. Divenyi (Ed.), Kluwer Academic, 167-179. CNBH, PDN, University of Cambridge The perception of musical notes and instruments Instrument Register Instrument Family Ss and Sf Envelope Shape CNBH, PDN, University of Cambridge General conclusions: If we divide the information in the magnitude spectrum into (a) envelope shape, (b) position of the envelope, Sf, and (c) position of the fine structure, Ss, it provides us with a good way of understanding the information in communication sounds. http://www.pdn.cam.ac.uk/groups/cnbh/teaching/lectures/PGW_SHAR_2010.pdf CNBH, PDN, University of Cambridge What and Who in vocal sounds I: The shape of the envelope largely determines vowel type. It determines the “What” aspect of the communication, in the sense of “What is being said” II: The acoustic scale variables, Ss and Sf , jointly determine the register of the voice, and whether the speaker sounds normal or unusual. They jointly determine much of the “Who” aspect of the communication in the sense of “Who is speaking” CNBH, PDN, University of Cambridge Family and Register in musical sounds I: The shape of the envelope/resonance largely determines the sound of the instrument family, or the family aspect of instrument perception. II: The acoustic scale variables, Ss and Sf , jointly determine the register or size aspect of instrument perception. III: The relationship between the acoustic scale variables, Ss and Sf determines whether an instrument sounds good, or thin or muddy. CNBH, PDN, University of Cambridge End of Act I Thank you Patterson, R. D., Smith, D. R. R., van Dinther, R. and Walters, T. C. (2008). “Size Information in the Production and Perception of Communication Sounds,” In: Auditory Perception of Sound Sources. Yost, W. A., Popper, A. N., and Fay, R. R. (Eds). Springer, LLC, New York, 43-75. http://www.pdn.cam.ac.uk/groups/cnbh/teaching/lectures/PSDWshar08.pdf Patterson, R. D., Gaudrain, E. and Walters, T. C. (2010). “The perception of family and register in musical tones,” In: Music Perception. Jones, M.R., Fay, R.R. and Popper, A.N., (eds). New York, Springer-Verlag. 13-50. http://www.pdn.cam.ac.uk/groups/cnbh/teaching/lectures/PGW_SHAR_2010.pdf Patterson, R. D., van Dinther, R. and Irino, T. (2007). “The robustness of bioacoustic communication and the role of normalization,” Proc. 19th International Congress on Acoustics, Madrid, Sept, ppa-07-011. CNBH, PDN, University of S S Cambridge f s pi pe ko kuuu ni ne mo muuu CNBH, PDN, University of Cambridge pi pe ko kuuu ni ne mo muuu pi pe ko kuuu ni ne mo muuu pi pe ko kuuu ni ne mo muuu 1/VTL (1/Sf ) Position of the envelope CNBH, PDN, University of Cambridge Position of the fine structure 1/VTL (1/Sf ) CNBH, PDN, University of Cambridge