Unit 4.2 Phylum Cnidaria 1 Phylum Cnidaria • Radial symmetry • Mouth at oral end surrounded by tentacles. • One opening into and out of gastrovascular cavity. • Cnidocytes that discharge nematocysts • Two body forms – Polyp – Medusa • Two germ layers • Nerve net • Still no circulatory or respiratory systems 2 Radial symmetry 3 Mouth and Tentacles 4 Fig. 13.2 5 Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. Generalized Cnidarian Life Cycle 6 Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. Cnidocyte Structure and Nematocyst Discharge 7 Diploblastic - 2 germ layers – Epidermis - outer covering (ectoderm) – Mesoglea - middle non-living jelly-like layer – Gastrodermis - lines gastrovascular cavity (endoderm) 8 Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. Cnidarian Body Wall 9 Nervous System 10 Class Hydrozoa • Freshwater & marine. • Cnidocytes present only on epidermis. • Medusa (if present) with a muscular velum that aids in propulsion. • Asexual reproduction by budding. • Sexual reproduction via gametes produced by epidermis & released into water. • Hydra, Ctenophores, Obelia, Man-of-War 11 Class Hydrozoa Hydra Encapsulated embryo Ovaries Budding Spermaries Sexual cycle Asexual cycle 12 Hydra Body Wall Gastrovascular cavity Gastrodermis Mesoglea Epidermis 13 14 15 Obelia • Freshwater or marine. • Live in colonies rather than free living like hydra. • Colonies contain two types of polyps, feeding and reproductive. • Dual stage life cycle like other Cnidarians. 16 Obelia 17 Obelia 18 Obelia Gonangium Medusa bud Medusae 19 Class Hydrozoa Gonionemus 20 Class Hydrozoa Gonionemus Velum 21 Class Hydrozoa Physalia 22 Class Hydrozoa Physalia • Commonly called the Portuguese Man-ofWar. • Not a jellyfish or even a single organism. • Made of a colony of hydrozoans that perform specialized functions. 23 Class Hydrozoa Physalia • Gastrozooids – Feeding & digestion. • Dactylozooids – Prey capture. • Gonozooids – Reproduction. • Pneumatophore – Gas-filled float. 24 Class Hydrozoa Physalia 25 Class Scyphozoa 26 Class Scyphozoa • True Jellyfish • Tentacles up to 70 meters in length • Cnidocytes present in gastrodermis & epidermis • Thick mesoglea contains amoeboid cells • Gametes produced by gastrodermis • All marine 27 Class Scyphozoa Aurelia – Moon Jellies 28 Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. Aurelia Life History 29 Fig. 13.18 30 Class Cubozoa • Medusa dominant & cuboid • Tentacles arise at four corners from blade-like pendalium. • All marine • Strong swimmers which prey primarily on fish • Stings of some may be fatal within minutes to humans. 31 Class Cubozoa Gonad Pedalium Tentacle 32 Class Cubozoa The Box Jellyfish 33 Class Cubozoa The Box Jellyfish • The most venomous poison known to man. • One brush against the skin will inject enough venom to put a full grown adult in the hospital for over a week. • Even a full morphine drip can not relieve the pain. 34 Class Cubozoa The Box Jellyfish 35 Class Cubozoa The Box Jellyfish 36 Class Anthozoa • Sea Anemones & Coral • Medusa stage virtually absent • Solitary or colonial • Some produce protective skeletons • Cnidocytes on tentacles and lining the gastrovascular cavity • Mesoglea contains amoeboid cells like all other Cnidarians • All Marine 37 Class Anthozoa Sea Anemone 38 Class Anthozoa Metridium Tentacles Mouth Pharynx Septum Gastrovascular cavity 39 Symbiosis 40 Class Anthozoa Corals • Protective skeleton of calcium carbonate • Polyp retracts when not feeding 41 Zooxanthellae • Photosynthetic dinoflagellates (brown) • Live in corals • Provide nutrients for coral by photosynthesis while gaining a home 42 • Mutualism Corals Colony of interconnected polyps 43 Class Anthozoa Meandrina Brain Coral 44 Class Anthozoa Gorgonia Sea Fan 45 Class Anthozoa Tubipora Pipe Organ Coral 46 Class Anthozoa Actinodiscus Mushroom Coral 47 Class Anthozoa Acropora Staghorn Coral 48 Coral Reefs • • • Formed over thousands of years from successive layers of coral skeleton deposits (calcium carbonate forms underwater mountains of coral animal skeletons) The underwater equivalent of the Amazon jungle- very high species diversity and biomass Reefs contain sponges, colonial hydrozoans, anemones, many varieties of coral, fish, many types of worms we’ve not discussed, not to mention bryozoans, ctenophores, protists, bacteria, etc etc.. 49 Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. Coral Reef Ecosystem Photo © McGraw-Hill Higher Education, Barry Barker, Photographer 50 Cladogram of Cnidaria Anthozoa Scyphozoa Hydrozoa Cubozoa Medusa cuboid Loss of medusa Polyp stage reduced All marine Radial symmetry & cnidocytes 51 The End 52