Invertebrate Zoology Lecture 7: Phylum Cnidaria, Part 2 Class Hydrozoa Siphonophore Photo: K. Raskoff / Monterey Peninsula College Class Hydrozoa Order Hydroidea: the hydroids Polyp form often colonial, branched (Obelia) Includes feeding and reproductive polyps Coenosarc: common gastrovascular cavity Class Hydrozoa Order Hydroidea: life cycle (Obelia sp.) Planula larvae Some solid, others with g.v. cavity Move via cilia Class Hydrozoa Order Siphonophora Two medusa types Swimming Sexual Reproduction Polyps: feeding, defense, asexual reproduction Predators! Gas-filled float Many variations! Class Hydrozoa Order Siphonophora Cormidium “colony within a colony” Detachable Class Hydrozoa Order Chondrophora (Velella velella, by-thewind sailors): Free-floating polyp Secondary polyps (feeding, reproduction) Tiny, free-living medusae Class Hydrozoa Order Trachylina: life cycle No true polyp stage Planula develops into medusa Class Hydrozoa Order Hydrocorallina: hydrocoral, fire coral Not a “true” coral! (True coral = C. Anthozoa) Calcareous skeleton Some with symbiotic zooxanthellae Polyps within chambers Reduced medusa stage Why called “fire” coral? Class Scyphozoa Semaeostomae (most jellies) Key predators (impact?) Stauromedusae Sessile medusa or polyp? Develop from planula larva Class Scyphozoa Life cycle, Semaeostomae (Aurelia) Key: strobilation of polyp ephyra medusa Class Scyphozoa Pelagia Note trailing mouth lobes Class Scyphozoa Life cycle, Pelagia No polyp stage: planula ephyra medusa Class Cubozoa: box jellies Polyp medusae via direct metamorphosis Chironex: deadliest toxin known www.zoologie-online.de/.../Cubozoa/cubozoa.html Class Anthozoa Anemones Coral Sea pansies Gorgonians Etc… Class Anthozoa Generalized life history: no medusae Asexual reproduction also important Class Anthozoa: Subclass Hexacorallia Order Actiniaria: solitary anemones Class Anthozoa: Subclass Hexacorallia Order Corallomorpha Anemone-like, but with clubbed tentacles Clonal form shown; clone wars occur! Class Anthozoa: Subclass Hexacorallia Order Scleractina: stony (“true”) corals Deposit CaCO3 skeletons Contain mutualistic zooxanthellae Some are reef builders Clubbed tentacles; polyps open at night… Class Anthozoa: Subclass Hexacorallia Order Scleractina: stony (“true”) corals Gastrovascular cavity connects polyps Mutualistic zooxanthellae within gastrodermis = acontia Mesentery = Class Anthozoa: Subclass Hexacorallia Order Scleractinia: stony (“true”) corals Initial colony: from planula larva produced sexually (VIDEO!) Colony growth: asexual division Fragmentation “new” colony Class Anthozoa: Subclass Ceriantipatharia Order Ceriantharia: tube anemones Double whorl of tentacles Build sand tubes; capable of burrowing Class Anthozoa: Subclass Octocorallia Features of subclass Eight-tentacled, pinnate polyps Eight complete mesenteries Class Anthozoa: Subclass Octocorallia Order Gorgonacea: sea whips, sea fans skeleton of gornonin (protein), and/or calcareous spicules often highly branched in one plane Oriented perpendicular to prevailing current Class Anthozoa: Subclass Octocorallia Order Alcyonacea: soft corals fleshy, sometimes tree-like pump water throughout the colony Scubatravel.com www.freeinfosociety.com Class Anthozoa: Subclass Octocorallia Order Pennatulacea: sea pens, sea pansies Main body = polyp; has secondary, lateral polyps Gastrozooids (feeding) Siphonozooids (water flow) Calcareous spicules provide structure © Alvaro E. Migotto Class Anthozoa: Subclass Octocorallia Order Stolonifera: organ pipe coral Calcareous "spicules" provide structure Polyps within tubes or on stalks