DNA Structure & Replication DNA Structure The discovery of the structure Francis Crick and James Watson • The deoxyribonucleic acid (DNA) molecule is the genetic blueprint for each cell and ultimately the blueprint that determines every characteristic of a living organism. The DNA molecule was discovered in 1951 by Francis Crick, James Watson, and Maurice Wilkins using X-ray diffraction. Photo Researchers, Inc. • "Francis Crick and James Watson," Microsoft® Encarta® 98 Encyclopedia. © 1993-1997 Microsoft Corporation. All rights reserved. In 1953 Crick, left, and Watson, right, described the structure of the DNA molecule as a double helix, somewhat like a spiral staircase with many individual steps. In 1962 Crick, Watson, and Wilkins received the Nobel Prize for their pioneering work on the structure of the DNA molecule. A little extra info • Although Maurice Wilkins from Cambridge is credited as a key player in the discovery, it was actually Rosalind Franklin, who was working in his lab, that used a technique called Xray diffraction to her determine the structure of DNA – she got no credit for the discovery! Watson/Crick Model of DNA • 1. 2 chains of nucleotides coiled around each other to form a double helix. • 2. The nitrogen bases of the 2 chains are joined together by weak hydrogen bonds. (easily broken) Watson/Crick Model of DNA • 3. A specific purine base is paired with a specific pyrimidine base. Adenine with Thymine (A-T) Guanine with Cytosine (G-C) • 4. The sequence of base pairs along the DNA molecule determine the genetic code. Structure Structure • Looks like a twisted ladder • The “handrails” or backbone is made of the phosphate and 5-C (pentose) sugar called deoxyribose • The “rungs” are made of the joined nitrogen bases • The nucleotides are joined together by covalent bonds into a single strand Nitrogen Bases There are 4 different nitrogen bases • Adinine,Thymine,Guanine,Cytosine • As Chargaff’s rule indicates: • Adinine bonds with Thymine • Guanine bonds with Cytosine • Therefore there will always be equal amounts of A and T , G&C. Purines • Nitrogen bases with a double ring structure • Adenine and Guanine Pyrimidines • Nitrogen bases that have a single ring structure • Thymine and Cytosine (and Uracil of RNA) Complementary Base Pairs • A two ring base will bind with a one ring base so that there are always three rings that separate the backbone • Three hydrogen bonds attach cytosine to guanine • Two hydrogen bonds attach thymine to adenine. Antiparallel • The two strands run opposite to each other Hydrogen bonds One end of the chain is 3’ (sugar end) the other end is 5’ (phosphate end) Draw and label a simple diagram of the molecular structure of DNA Un-seeable Biology What is a “genome”? • the genome is the entirety of an organism's hereditary information • The genome includes both the genes and the noncoding sequences of the DNA/RNA. RNA • Not the same as DNA because: • The sugar component of RNA is ribose rather than deoxyribose • Uracil instead of Thymine • Remains single stranded, though it can fold back on itself to produce regions of complementary base pairs THE CENTRAL DOGMA Table 12.01 12.08 DNA Replication Interphase of Mitosis/Meiosis • Semi conservative • Meaning one old strand combines with a new strand to produce two new double strands of DNA Parent Molecule Separation of strands “Daughter” DNA molecules each consisting of one parent strand and one new strand Animation • http://www.mcgrawhill.ca/school/applets/abbio/ch18/dna_replicatio n.swf DNA Replication • semi-conservative replication-new DNA molecule made of one parent and one newly replicated strand. • In general a DNA molecule ‘unzips’ down the middle of the paired bases, 2 individual strands are made that will become the ‘templates’ for new complete DNA stands The Steps for DNA Replication (during S-phase of Interphase): 1. Initiation starts at a specific nucleotide sequence, a group of enzymes called DNA helicases breaks hydrogen bonds between bases to unzip the double helix 2. Proteins bind to keep strands apart 3) RNA primers attach to a spot on the original DNA stand 4) DNA polymerase III – starts at where the primer attached to the DNA and makes new strand in 5’ to 3’ direction (always) 5) DNA polymerase 1 – removes primers and replaces with nucleotide 6) DNA ligase – joins DNA fragments • DNA Replication Remember—Replication 3’-5’ Build a DNA DNA Replication DNA makes DNA Un-seeable Biology 2:52