Invertebrates III and Vertebrates Phylum: Echinodermata Deuterostomes – radial and indeterminate cleavage – Enterocoelous – anus from blastopore Phylum: Echinodermata Secondary Radial Symmetry Water vascular system – Ambulacral groove – Madreporite All marine Water Vascular System Madreporite Stone Canal Ring Canal Radial Canal Lateral Canal Ampulla Tube Feet Classification Class: Asteroidea (Seastars) Class: Opiuroidea (Brittlestars) Class: Echinoidea (Sea Urchins, Sand Dollars) Class: Crinoidea (Sea Lilies) Class: Holothuroidea (Sea Cucumbers) Class: Asteroidea Five arms radiating from a central disc Open ambulacral groove Madreporite on the aboral side Contain pedicellariae or papulae Class: Ophiuroidea Five thin arms radiating from a central disc Closed ambulacral grooves Madreporite on the oral side No suckers on tube feet, pedicellariae or papulae Class: Echinoidea No arms but have five rows of tube feets Contain spines Closed ambulacral grooves Madreporite on the aboral side Contain pedicellariae or papulae Aristotle’s lantern Class: Crinoidea Attached to substrate with many branched arms Open ambulacral grooves No Madreporite No pedicellariae or papulae Class: Holothuroidea Soft bodied Ambulacral areas with tube feet Internal Madreporite No pedicellariae or papulae Phylum: Chordata Deuterostomes – radial and indeterminate cleavage – Enterocoelous – anus from blastopore Bilateral Symmetry Both invertebrates and vertebrates – Contain four anatomical features Phylum: Chordata Notochord Dorsal, Hollow Nerve Cord Pharyngeal Slits Muscular, Postanal Tail SubPhylum: Urochordata Tunicates Sessile Only contains Pharynx with slits as an adult SubPhylum: Cephalochordata Lancelates Contains all four chordate characters as an adult Closest relative to vertebrates (Amphioxus) Paedogenesis SubPhylum: Vertebrata Backbones Contains all four chordate characters as an adult with modification Neural Crest – bones and cartilage of the skull Vertebrate Adaptations Living Endoskeleton – better for larger animals Pharynx and Efficient Respiration – increased metabolic rate Advanced Nervous System – developed system for distance reception Paired Limbs – increased movement Chordate Evolution Vertebrae Jaws and two sets of paired appendages Teeth Lungs Legs Amniotic Egg Hair, feathers Key Fish Characteristics Vertebral Column Jaws and paired appendages Gills Single Circuit blood circulation Superclass: Agnatha Without Jaws and Most without paired appendages Class: Myxini Hagfishes (scavengers) Class: Cephalaspidomorphi - Lampreys (parasitic) Superclass: Gnathostomata With jaws Evolved from skeletal supports of the pharyngeal slits Fossil Gnathostomata Placoderms – Plate-skinned Acanthodians – Probably led to bony fish Class: Chondrichthyes Class: Chondrichthyes Placoid Scales (teeth-like) Several rows of teeth – (Not embedded in the jaw) Class: Chondrichthyes Spiral valve within intestine Large fatty liver Senses – Ampullae of Lorenzini – Lateral Line Class: Chondrichthyes Cartilaginous skeleton (not primitive) – Subclass: Elasmobranchi Sharks, Skates, Rays – Subclass: Holostei Ratfish Subclass: Elasmobranchi Order: Selachidae – Sharks – Cartilagenous skeleton – Streamlined body – 5-7 gill slits Subclass: Elasmobranchi Order: Batiformes – Skates, Rays – Cartilagenous skeleton – flattened body – 5-6 gill slits on the underside of the body Subclass: Holocephali – Cartilagenous skeleton – Lack dermal scales – Venomous spine – single gill opening with hard covering Osteichthyes - Bony Fish (Bony Skeleton) Class:Sarcopterygii – Lobe-finned Fish – Lungfish Class: Actinopterygii – Ray-finned Fish Osteichthyes Embedded dermal (ctenoid) scales Operculum Swim Bladder Lateral Line Osteichthyes Fins – – – – – Dorsal Pectoral Pelvic Caudal Anal Class:Sarcopterygii Lobe-finned Fish Subclass: Coelacanthiomorpha Coelocanth – Fleshy pectoral and anal fins which are supported by bones. Class:Sarcopterygii Lobe-finned Fish Subclass: Dipnoi Lungfish – Fleshy fins – True lungs Class Actinopterygii Infraclass: Holostei – (Primitive fish) – Order: Lepisoteriformes Bowfins – Order: Amiiformes - Gars Bowfins Gar They are found in brackish conditions. They can use their swim bladders to obtain extra oxygen. Class Actinopterygii Infraclass: Teleostei In this infraclass, all of the fish are considered to be the ray-finned fish. They have a movable maxilla and premaxilla and modified muscles that allow them to have a protrusable mouth. Class Actinopterygii Infraclass: Teleostei Superorder: Osteoglossomorpha Superorder: Elopomorpha Superorder: Clupeomorpha Superorder: Ostariphysi Superorder: Protacanthopterygii Superorder: Stenopterygii Superorder: Scopelomorpha Superorder: Acanthopterygii Superorder: Osteoglossomorpha Bony Tongued Fish The Bony tongue is used to bite against. They are also found in brackish conditions. They can use their swim bladders to obtain extra oxygen. They are snakelike with long bodies. Unlike land snakes, eels are usually scale less, although a few species can be found with tiny scales along their bodies. Over 100 vertebrae form the eel's spine, which makes the animal very flexible. Superorder: Elopomorpha Eels Superorder: Clupeomorpha Clupeiformes This is the order of rayfinned fish that includes the herring family and the anchovy family. Clupeiformes are physostomes, which means that the gas bladder has a pneumatic duct connecting it to the gut. They typically lack a lateral line. Superorder: Ostariphysii These fish release an alarm substance and their first few vertebrates are used to pass sound from the swim bladder to the inner ear for acute hearing. They also have a gas bladder. Superorder: Protacanthopterygii These are fish that lack specialization. They are important game fish like Salmon and Trout Superorder: Stenopterygii Dragonfish are deep water fish. Many deep sea fish are biouminescent Superorder: Scopelomorpha Lantern fish are deep water fish. Many deep sea fish are bioluminescent. They have large eyes and adipose fins. Superorder: Acanthopterygii Ray-finned Fish – they make up of a very diverse group of fish which usually have a Pelvic fin spine present. Evolution of the Tetrapods Evolution of the Tetrapods Deep Sea Fish Deep sea fish are among the most elusive and unusual looking creatures on Earth.