17.1 The Linnaean System of Classification KEY CONCEPT Organisms can be classified based on physical Pangolian ↓ similarities. 17.1 The Linnaean System of Classification Linnaeus developed the scientific naming system still used today. • Taxonomy is the science of naming and classifying organisms. White oak: Quercus alba • A taxon is a group of organisms in a classification system. 17.1 The Linnaean System of Classification Aristotle • Grouped the types of creatures according to their similarities: animals with blood and animals without blood, animals that live on water and animals that live on land. • Hierarchical classification. He assumed that creatures could be grouped in order from lowest to highest, with the human species being the highest. 17.1 The Linnaean System of Classification Aristotle 384 BC – 322 BC • Greek philosopher and a student of Plato. • His writings cover many subjects, including poetry, theater, music, logic, linguistics, politics, government, ethics, biology, and zoology. • In the zoological sciences, some of his observations were confirmed to be accurate only in the 19th century. • All aspects of Aristotle's philosophy continue to be the object of active academic study today. 17.1 The Linnaean System of Classification Carl Linnaeus 1707-1778 • was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. • He is known as the father of modern taxonomy, and is also considered one of the fathers of modern ecology. 17.1 The Linnaean System of Classification • Binomial nomenclature is a two-part scientific naming system. – uses Latin words – scientific names always written in italics or underlined – two parts are the genus name and species descriptor 17.1 The Linnaean System of Classification • A genus includes one or more physically similar species. – Species in the same genus are thought to be closely related. – Genus name is always capitalized. • A species descriptor is the second part of a scientific name. – always lowercase – May refer to a trait of the species, scientist who first Tyto alba described it or native location White barn owl 17.1 The Linnaean System of Classification Advantages • Scientific names help scientists to communicate. – Some species have very similar common names. – Some species have many common names. 17.1 The Linnaean System of Classification Linnaeus’ classification system has seven levels. • Each level is included in the level above it. • Levels get increasingly specific from kingdom to species. 17.1 The Linnaean System of Classification Seven levels of classification • • • • • • • Kingdom Phylum/Division Class Order Family Genus Species 17.1 The Linnaean System of Classification The Linnaean classification system has limitations. • Linnaeus taxonomy doesn’t account for molecular evidence. – The technology didn’t exist during Linneaus’ time. – Linnaean system based only on physical similarities. 17.1 The Linnaean System of Classification • Physical similarities are not always the result of close relationships. • Genetic similarities more accurately show evolutionary relationships. Red panda → More related to raccoons 17.1 The Linnaean System of Classification Cladistics is classification based on common ancestry. • Phylogeny is the evolutionary history for a group of species. – evidence from living species, fossil record, and molecular data – shown with branching tree diagrams 17.1 The Linnaean System of Classification 17.1 The Linnaean System of Classification • Cladistics is a common method to make evolutionary trees. – classification based on common ancestry – species placed in order that they descended from common ancestor 17.1 The Linnaean System of Classification • A cladogram is an evolutionary tree made using cladistics. – A clade is a group of species that shares a common ancestor. – Each species in a clade shares some traits with the ancestor. – Each species in a clade has traits that have changed. 17.1 The Linnaean System of Classification • Derived characters are traits shared in different degrees by clade members. 1 Tetrapoda clade – basis of arranging species in cladogram – more closely related species share more derived characters – represented on cladogram as hash marks 2 Amniota clade 3 Reptilia clade 4 Diapsida clade 5 Archosauria clade FEATHERS & TOOTHLESS BEAKS. SKULL OPENINGS IN FRONT OF THE EYE & IN THE JAW OPENING IN THE SIDE OF THE SKULL SKULL OPENINGS BEHIND THE EYE EMBRYO PROTECTED BY AMNIOTIC FLUID FOUR LIMBS WITH DIGITS DERIVED CHARACTER 17.1 The Linnaean System of Classification • Nodes represent the most recent common ancestor of a clade. CLADE 1 Tetrapoda clade 2 Amniota clade 3 Reptilia clade 4 Diapsida clade • Clades can be identified by snipping a branch under a node. 5 Archosauria clade FEATHERS AND TOOTHLESS BEAKS. SKULL OPENINGS IN FRONT OF THE EYE AND IN THE JAW OPENING IN THE SIDE OF THE SKULL SKULL OPENINGS BEHIND THE EYE EMBRYO PROTECTED BY AMNIOTIC FLUID NODE FOUR LIMBS WITH DIGITS DERIVED CHARACTER 17.1 The Linnaean System of Classification Clades can be identified by snipping a branch under a node. • Molecular data may confirm classification based on physical similarities. • Molecular data may lead scientists to propose a new classification. • DNA is usually given the last word by scientists. 17.1 The Linnaean System of Classification Molecular clocks use mutations to estimate evolutionary time. • How they work Mutations add up at a constant rate in related species. – This rate is the ticking of the molecular clock. – As more time passes, there will be more mutations. Mutations add up at a fairly constant rate in the DNA of species that evolved from a common ancestor. DNA sequence from a hypothetical ancestor Ten million years later— one mutation in each lineage Another ten million years later— one more mutation in each lineage The DNA sequences from two descendant species show mutations that have accumulated (black). The mutation rate of this sequence equals one mutation per ten million years. 17.1 The Linnaean System of Classification • Scientists estimate mutation rates by linking molecular data and real time. – an event known to separate species – the first appearance of a species in fossil record 17.1 The Linnaean System of Classification Mitochondrial DNA and ribosomal RNA provide two types of molecular clocks. • Different molecules have different mutation rates. – higher rate, better for studying closely related species – lower rate, better for studying distantly related species 17.1 The Linnaean System of Classification • Mitochondrial DNA is used to study closely related species. – mutation rate ten times faster than nuclear DNA – passed down unshuffled from mother to offspring grandparents mitochondrial DNA nuclear DNA parents Mitochondrial DNA is passed down only from the mother of each generation,so it is not subject to recombination. child Nuclear DNA is inherited from both parents, making it more difficult to trace back through generations. 17.1 The Linnaean System of Classification • Ribosomal RNA is used to study distantly related species. – many conservative regions – lower mutation rate than most DNA 17.1 The Linnaean System of Classification Classification is always a work in progress. • The tree of life shows our most current understanding. • New discoveries can lead to changes in classification. – Until 1866: only two kingdoms, Plantae Animalia and Plantae Animalia 17.1 The Linnaean System of Classification Classification is always a work in progress. • The tree of life shows our most current understanding. • New discoveries can lead to changes in classification. – Until 1866: only two kingdoms, Plantae Animalia and Plantae Animalia – 1866: all single-celled Protista organisms moved to kingdom Protista 17.1 The Linnaean System of Classification Classification is always a work in progress. • The tree of life shows our most current understanding. • New discoveries can lead to changes in classification. – Until 1866: only two kingdoms, Plantae Animalia and Plantae Animalia – 1866: all single-celled Protista organisms moved to kingdom Protista – 1938: prokaryotes moved to kingdom Monera Monera 17.1 The Linnaean System of Classification Classification is always a work in progress. • The tree of life shows our most current understanding. • New discoveries can lead to changes in classification. – Until 1866: only two kingdoms, Plantae Animalia and Plantae Animalia – 1866: all single-celled Protista organisms moved to kingdom Protista – 1938: prokaryotes moved to kingdom Monera – 1959: Fungi moved to own kingdom Monera Fungi 17.1 The Linnaean System of Classification Classification is always a work in progress. • The tree of life shows our most current understanding. • New discoveries can lead to changes in classification. – Until 1866: only two kingdoms, Plantae Animalia and Plantae Animalia – 1866: all single-celled Protista organisms moved to kingdom Protista – 1938: prokaryotes moved to kingdom Monera Archea – 1959: fungi moved to Fungi own kingdom Eubacteria – 1977: kingdom Monera split into kingdoms Eubacteria and Archaebacteria 17.1 The Linnaean System of Classification 1977: Carl Woese discovered two genetically different groups of prokaryotes, based on rRNA studies of prokaryotes • He proposed creating 3 domains above the kingdom level. – domain model more clearly shows prokaryotic diversity