The Phylum Chordata The Chordates • Phylum Chordata • Subphyla: Urochordata, Cephalochordata, & Vertebrata The Vertebrate Classes • agnatha -jawless fish, • chondrichthyes-cartilage fish, • osteichtheyes- bony fish • amphibia, • reptilia, • aves -birds, • mammals Characteristics shared by all chordates at some stage in their lives: • 1) notochord -stiff rod of specialized cells, encased by a firm sheath along the length of the dorsal side. It becomes the vertebrate backbone notochord development in a zebra fish embryo notochord in a lancet • 2) gill slits or pharyngeal pouches: seen in embryos, develop into gills in fish & other similar structures • 3) dorsal hollow nerve cord : ( bundle of nerves runs down the back) above the notochord, the anterior end develops into the brain, & the remaining part becomes the spinal cord (part of the Central Nervous System) • 4) post anal tail - Extension of the body past the anal opening - -amphioxus propels aquatic forms Phylum Hemichordata - Protochordates – Lack some key chordate characteristics • Body in three parts: proboscis, colar and trunk Enteropneusta – Acorn worms (solitary, live in mud or vegetation)– filter feeders . • Pterobranchia – (pterobranchs) – tiny deep-sea colonial moss like animals – No trace of a notochord. • They live in secreted tubes. • Larval stage – “Planktonic snails – Sea Angels” The larva settle on substrate and reproduce asexually by budding to create a new colony. Sub phylum Urochordata: “Sea Squirts” (Tunicates) • Larval Form = tunicate larva possess all four characteristics • Larvae are tadpole-like and free-living, and have an endostyle, gill slits, dorsal nerve cord, and notochord. • The larval stage lasts only a few days, and ends when the larva attaches to a substrate and metamorphoses into an adult. • Adult is sessile (and sometimes colonial), and must obtain food by siphoning sea water through its body and trapping food particles in the endostyle. • Their outer covering is called the tunic_ barrelshaped animals that live on the ocean bottom. • Adapted for filter-feeding. • They are hermaphrodites. Fertilization is _external_. They have a pouch-like _pharynx_ but no notochord, nerve cord, or post anal tail. Eat Them! Subphylum - Cephalochordata: Lancelets • The Origin of the free-swimming vertebrates (Led to Fish - actively feeding & predatory) • Structures shows cephalzation. 1) blade-shaped animals they retain: notochord, dorsal nerve cord, pharyngeal pouches, & post anal tail, (They are weak swimmers, and burrow into sand.) 2) Live in shallow warm water. 3) Use cilia to draw in water and trap food, water leaves body through the atriopore.