Month Day Topic Nov. 8 Individuals to populations 10 Holiday! 13 Populations to communities 15 Community patterns 17 Ecosystems 20 Film-1st showing 22 Film-2nd showing 24 Holiday! 27 Ecosystem services A population is: A population is: A group of individuals of the same species living in a given area at a given time. What controls where a population lives? What controls where a population lives? • It has to get there – Evolution – Dispersal What controls where a population lives? • It has to get there – Evolution – Dispersal • Its ecological niche Key concept in ecology: the niche Key concept in ecology: the niche • Fundamental niche – Abiotic conditions that the species can live within Key concept in ecology: the niche • Fundamental niche – Abiotic conditions that the species can live within • Realized niche – Biotic interactions that reduce or enhance the fundamental niche Barnacle story Balanus’ fundamental niche: upper, middle and lower zone Balanus’ realized niche: middle zone, limited from upper by competition, lower by predation Interactions among populations determine a species’ realized niche... A community is: A community is: Populations of different species that live together in a given area at a given time. Communities are... • Defined by abiotic factors – Populations in a community have similar fundamental niches Communities are... • Defined by abiotic factors – Populations in a community have similar fundamental niches • Defined by biotic interactions – Some are weak – Some are strong Types of community interactions POP. 2 POP. 1 Harm No effect Benefit Harm Competition Amensalism Predation or Parasitism No effect Amensalism Benefit Predation or Parasitism Commensalism Commensalism Mutualism Types of community interactions POP. 2 Harm POP. 1 No effect Benefit Harm Competition Amensalism Predation or Parasitism No effect Amensalism Commensalism Benefit Predation or Commensalism Mutualism Parasitism Types of community interactions POP. 2 Harm POP. 1 No effect Benefit Harm Competition Amensalism Predation or Parasitism No effect Amensalism Commensalism Benefit Predation or Commensalism Mutualism Parasitism Keys to competition (-/-) • Organisms compete for limited resources Keys to competition (-/-) • Organisms compete for limited resources • Can be: – Intraspecific Time Population size Keys to competition (-/-) • Organisms compete for limited resources • Can be: – Intraspecific – Interspecific Competition can: • Restrict species ranges Competition can: • Restrict species ranges • Reduce species abundances Competition can: • Restrict species ranges • Reduce species abundances • Cause the local extinction or competitive exclusion of species from an area The ghost of competition past... Predator-prey interactions (+/-) • Predators are generally larger than their prey (but many exceptions...) Predator-prey interactions (+/-) • Predators are generally larger than their prey (but many exceptions...) • Predators live outside of the body of their prey Predator-prey interactions (+/-) • Predators are generally larger than their prey (but many exceptions...) • Predators live outside of the body of their prey • Predators generally kill their prey Predator-prey interactions • Dynamics of predator and prey populations may be: Predator-prey interactions • Dynamics of predator and prey populations may be: • Loosely coupled – predator “switches” when prey is scarce • Tightly coupled – predator starves when prey is scarce Tightly coupled predator-prey interactions change over time • An evolutionary “arms race” Tightly coupled predator-prey interactions change over time • An evolutionary “arms race” • Mimicry Batesian mimicry Batesian mimicry Predator-prey interactions change over time • An evolutionary “arms race” • Mimicry • Plant defenses against herbivores Host-parasite interactions (+/-) • Parasites are generally smaller than their hosts • Parasites live inside or on the body of their host • Parasites generally kill their hosts more slowly than predators Amensalism (0/-) Commensalism (0/+) Mutualisms (+/+) Mutualisms can be loose or tight Acacia with ants Acacia without ants Communities are... • Defined by abiotic factors – Populations in a community have similar fundamental niches • Defined by biotic interactions – Weak and strong interactions determine a population’s realized niche • Dynamic – Vary in time and in space Why do communities vary in time? Communities are... Comprised of populations that respond individualistically to change Communities vary in time Ecological succession: a sequence of change in the species composition of a community Succession 1. Primary – establishment of communities on new sites that previously had no organisms 1949 2001 Succession 1. Primary – establishment of communities on new sites that previously had no organisms 2. Secondary – re-establishment of communities following disturbance Why do communities vary in space? Communities vary in space • • • • • Time Relief/aspect Parent material Climate Organisms Terrestrial biome Terrestrial biome • A major community type that differs from other types in the structure of its dominant vegetation. Terrestrial biome • A major community type that differs from other types in the structure of its dominant vegetation. • Primarily controlled by climate. – Relationship between rainfall and temperature Biomes • The vegetation of a biome has a similar appearance wherever on Earth that biome is found. • In many cases, this is due to convergent evolution. Euphorbiaceae Cactaceae Tundra Tundra -severe winters -short growing season, cool summer -arctic or alpine Tundra Temperate Forest Temperate forest Temperate Forest -distinct winter season, frost a defining feature -summer season usually moist Temperate Forest Desert Desert -hot or cold deserts exist -low precipitation Desert Tropical Rain Forest -found near the equator -temperature varies little from approximately 23°C -Over 2 m of rainfall Tropical Rain Forest Tropical Rain Forest -canopy trees up to 55 m tall -soils are generally unfertil -largest biome, on an area basis -nutrients and carbon store in plant biomass, not soils