AP Biology Chapter 27: Prokaryotes 12 Concept 27.1: Structural, functional, and genetic adaptations contribute to prokaryotic success Cell-Surface Structures: Quintessential towards prokaryotic survival; different surface structure and structural arrangement will protect the unicellular organism’s organelles from the external environment. Gram Positive: The simplicity of the surface structure allows it to be easily targeted and destroyed. Gram Negative: The complexity of the cell wall and the lipopolysaccharides make Gram-Negative bacterial more resistant to resistant to drugs and toxic. Fimbriae and Capsules are structural formations on the surface of cells that allow them to adhere to certain surfaces. Motility: Prokaryotes use flagella for motility. Taxis – cellular movement towards or away from a given stimulus. Internal and Genomic Organization: The infolding of plasma membranes in prokaryotes is similar to the energy producing organelles of eukaryotic cells: chloroplasts and mitochondria. The unbound DNA of prokaryotes reside in the nucleoid region. Prokaryotic DNA also exists as plasmids, which may be transferred between prokaryotes. Reproduction and Adaptation: Because prokaryotes are able to divide every 1-3 hours, their accumulation of beneficial mutations are much more common than in eukaryotes. Endospores, are resilient, dehydrated shells that prokaryotes may store their genetic information into if they experience unsuitable conditions. Concept 27.2: Variety of nutritional and metabolic adaptations in Prokaryotes AP Biology Chapter 27: Prokaryotes 12 Obligate aerobes and anaerobes require O2 and CO2, respectively, to survive, while facultative anaerobes can survive in either conditions, but would prefer an O2 environment. Certain prokaryotes obtain food by fixing inorganic nitrogen through nitrogen fixation. Metabolic Cooperation occurs in single bacterium or entire colonies where more than one metabolic process occurs and is the reason that some prokaryotes exist in close proximities. Concept 27.3: Prokaryotic Phylogeny: Through comparing small-subunit ribosomal RNA, Woese classified a type of prokaryote as Archaea as they exhibit a combination of bacterial and eukaryotic traits. Archaea extremohiles are prokaryotic organisms that thrive in extreme conditions. Extreme thermophiles exist in very hot environments. Extreme halophiles exist in very saline environments. Methanogens are unique archaea that derive their energy by using CO2 to oxidize H2 and release methane as a waste produce. Major Groups of Bacteria: Proteobacteria: A diverse group of gram-negative bacteria. Chlamydias: Parasitic cell that lack peptidoglycan. Spirochetes: Obtain motility through rotating flagellum-like filaments. Gram-Positive Bacteria: An equally diverse group of bacteria. Cyanobacteria: The only prokaryote that carries out oxygen-generating photosynthesis. Concept 27.4: Prokaryotes play crucial roles in the biosphere Relationship between organisms: Symbiosis is the ecological relationship between organisms of different species. The larger of the two is called the host, while the smaller is called the symbiont. In mutualism, both organisms benefit. In commensalism, one organism benefits while not harming nor benefiting the other. In parasitism, the parasite organism benefits at the expense of the host. Concept 27.5: Impact of Prokaryotes on humans Negative Impacts: Exotoxins - Toxins secreted by prokaryotes Endotoxins - Toxins that coat the outer membrane of gram-negative cells, otherwise known as the lipopolysaccharide layer. These toxins are only released once the cell has been destroyed and its cell wall broken down. Positive Impacts: Bioremediation – The use of micro-organisms to remove pollutants from the air, water, and soil.