MOLLUSKS Chapter 16 Molluscs are triploblastic which means three tissue layers. II. Molluscs are coelomates: a fluid filled body cavity lined by mesoderm. Have a complete digestive system. 2 GROUPS OF COELOMATES: 1. Protostomes: Molluscs and Annelids Spiral, determinate cleavage The blastopore forms the mouth Deuterostomes: echinoderms, chordates v Radial, indeterminate cleavage. v The blastopore forms the anus. 2. The coelom may have formed from splitting of the mesoderm or from outpocketing of the gut. PHYLUM MOLLUSCA SHARES CHARACTERISTICS . 8 different classes Body composed of a head-foot and visceral mass (organ systems) A mantle – covers visceral mass and may secrete a calcareous shell III OTHER COMMON CHARACTERISTICS Bilateral symmetry Protostome characteristics A trochophore larva A reduced coelom Open circulatory system (exception: cephalopods which have a closed circulatory system) Many molluscs feed via a radula Class Examples Feeding/digestion Respiratory Gastropoda Snails and Slugs Herbivores, feeding with a radula Gills; mantle contractions circulate air; have a siphon which serves as an inhalant tube. Circulatory Open (blood flows from heart leaves the vessels and goes directly into sinuses.) Skeletory Hydraulic (blood confined to tissue spaces for support.) 6 ganglia in the head foot and a visceral mass a. eyes on tentacles b. statocysts on the foot for equilibrium c. ophradia in the anterior wall of the mantle that helps detect prey. Nervous system Excretory Nephridium – converts ammonia to uric acid Reproduction Locomotion Special Features External fertilization with some being monoecious and others being dioecious. A ciliated, flattened foot covered with glands used to creep across the substrate. Torsion – gives shell the coiled appearance Class Bivalvia Examples Feeding/digestion Respiratory Clams, oysters, mussels, and scallops Filter-feeders; loss of head and radula Cilia covered gills forming folded sheets called lamellae; cilia moves water over an incurrent siphon Open External shell 3 pairs of ganglia, sensory cells on the margin of the mantle, photoreceptors, statocysts and osphradium Nephridium Most dioecious; external fertilization Sedentary Two convex halves of shell called valves Circulatory Skeletory Nervous system Excretory Reproduction Locomotion Special Features Class Cephalopoda Examples Octopuses, squids, cuttlefish, and nautili Feeding/digestion Locate prey by sight and capture prey with tentacles that have adhesive cups; radula and beak for tearing food; peristalsis replaces ciliary action Respiratory Circulatory Skeletory Nervous system Excretory Reproduction Locomotion Special Features Gills Closed – blood confined to vessels; more efficient Shell is reduced or absent; except nautilus Large brain – memory and decision making; can form images, distinguish shape & some color; statocysts by brain; osphradia only in nautilus Have a kidney which allows waste to filter directly from the blood stream which is very efficient Dioecious; males have a hectocotylus for spermatophore transfer into the female near the opening of the oviduct; eggs deposited onto a substrate and hatchlings are miniature adults. Jet propulsion (water forced through funnel) Modified foot into tentacles or arms. They have chromatophores (pigment cells) which allow animal to change color and discharge ink for defense; confuses predator allowing