9. Almost Rosids to Rosids
Santalales






Lots of parasitism
Polyacetylenes
1-seeded ovules, pendulous and freecentral placentation
Molecules
In ours, flowers not prominent
Families not on required list
“Santalaceae”


Hemiparasites
Root parasites
Viscaceae








Epiphytic parasites
Haustoria
Stems jointed
Lvs opp, oft. Coriaceous
or succ.
Flrs unisex, inconspic,
radial
T3-4A3 (no fil),G3-4,
inf. Viscous berry
Passive gametophyte
Pantrop + some
temperate
Loranthaceae






Us. Epiphytes
Lv opp or almost so
Flrs bisex, rad or bilat
Ca rimCo(3)5-6(9),
distinct or conn, oft.
Red or yellow, A=Co,
opp, fil adnate
Aggr. gametophyte
Tropical
ROSIDS





Weakly supported group
Also some lack of clarity of relationships
within
Lots of representatives in our part of
the world
Hypanthium features in many families
Radial symmetry common
Saxifragales



Some families chiefly (only?) on
molecular data
Hypanthium, 2-united carpels, glandular
leaf teeth
Retained: 5-merous flowers with
separate parts
Saxifragaceae





Two interpretations Shaw – as in text; Welsh
– much broader (older)
Herbs, often basal leaves, alt. NO STIPULES
Flr rad to bilat, HYPANTH,
Ca4-5 sometimes connate, Co4-5, oft.
Clawed, St 3-10. G2(-5), connate, varying
position, axile, oft. nectar disk
Fr. Capsule or follicle
Crassulaceae
Geraniales


Five families, 1 in north Utah
Morphological features of order



Leaf margins with glandular teeth
Inflorescence cymose
Calyx persistent in fruit
 A few other characters


Anomocytic stomates
Vessel elements with simp[le perforations
Geraniaceae






Remarkable feature is fruit
Herbs to subshrubs; stems usually jointed at
nodes
Hairs glandular, with aromatic oils (I.e.,
smelly plants)
Stipules present, leaves with gland. Teeth
K5C5A(5)10,15, filaments often slightly connateG5conn
Fruit a schizocarp, persistent style column
Persistent style
Note the fruits in picture above
and on the right
Malpighiales


This order contains 7.8% of the
diversity of angiosperm diversity
There does not seem a whole lot tying
it together, see:
http://www.mobot.org/MOBOT/Research/APweb/welcome.html
The Angiosperm Phylogeny Group’s home
page
Euphorbiaceae