INVERTEBRATE SURVEY LAB Spring 09 CHARACTERISTICS OF ANIMALS Representatives of the animal kingdom display a wide diversity in appearance. Regardless, they do share certain characteristics: ALL ANIMALS ARE HETEROTROPHIC ALL ANIMALS ARE MULTICELLULAR ALL ANIMALS ARE MADE OF EUKARYOTIC CELLS THAT LACK A CELL WALL The animal kingdom is divided into several phyla (categories). Each phylum contains animals which demonstrate common traits. These traits allow for a classification system that shows progressive change in the phyla from simple to more complex in forms. KEY TERMS TO KNOW Body Symmetry Asymmetry – no specific shape Radial Symmetry – body plan in which body parts repeat around the center of the body Bilateral Symmetry – a single imaginary plane could divide the organism in to 2 equal halves Asymmetry KEY TERMS TO KNOW • Cephalization – Cephalization is a concentration of sense organs and nerve cells in the front of the body (toward the head). KEY TERMS TO KNOW Coelom Coelom means body cavity (a tube within a tube). About the Coelom – Coelom means body cavity (a tube within a tube). – – – Acoelomates • Have NO TRUE body cavity! Pseudocoelomates • Have a fluid filled cavity BUT – it is not lined like that of higher-order animals. Coelomates • Have a body cavity that separates the body into an inner and outer tube (fully lined). KEY TERMS TO KNOW Early Development Protostomes are organisms that develop a mouth first and then an anus. Most invertebrates are protostomes Deuterostomes are organisms that develop an anus first and then a mouth. Echinoderms and all vertebrates are deuterostomes. KEY TERMS TO KNOW Early Development– During early development, the cells of most animal embryos separate into three layers called germ layers: 1. 2. 3. ENDODERM: innermost layer; develops into the lining of the digestive tract & much of the respiratory system MESODERM: middle layer; develops into muscles and much of the circulatory, reproductive, and excretory systems ECTODERM: outermost layer; develops into the sense organs, nerves, and outer layer of the skin KEY CONCEPT TO KNOW TRENDS IN ANIMAL EVOLUTION: As animals get more advanced, they have more specialized features than those animals found in the prior phyla. PHYLUM PORIFERA - SPONGES KEY CHARACTERISTICS Sponges are: Multicellular, but do not have tissues! Filter feeders that sift microscopic food particles from the water. Asymmetrical – they have no front or back end! Sessile – they do not move! No cephalization! Early development is protostome. Habitat is water. No segmentation! No germ layers! No coelom! ECOLOGY OF SPONGES Sponges play an important part in the ecology and survival of numerous aquatic organisms: They usually live attached to the sea floor. They form mutually beneficial relationships with bacteria, algae, and plantlike protists. They provide a habitat for snails, sea stars, sea cucumbers, and shrimp. PHYLUM PORIFERA - SPONGES PHYLUM CNIDARIA – JELLYFISH, SEA ANEMONES, & CORALS Epidermis Mesoglea Gastroderm Tentacles Mesoglea Gastrovascular cavity Mouth/anus Mouth/anus Gastrovascular cavity Tentacles Medusa Polyp KEY CHARACTERISTICS RADIAL SYMMETRY CARNIVOROUS NO CEPHALIZATION MOVEMENT: Polyps are sessile Medusas are free swimming EARLY DEVELOPMENT: protostome & nerve net SOFT/SAC-LIKE BODY PLAN 2 WAY DIGESTION (ONE OPENING) Eat & excrete through same hole! NO COELOM NO SEGMENTATION 2 GERM LAYERS: ectoderm & endoderm IMPORTANT FEATURES They typically have a life cycle that includes two different-looking stages: a polyp and a medusa: A polyp is a cylindrical body with arm-like tentacles. • In a polyp, the mouth points upward. • Polyps are usually sessile (do not move). A medusa has a motile, bell-shaped body with the mouth on the bottom. CNIDARIAN ACTIVITY Cnidarians are soft-bodied, carnivorous animals that have stinging tentacles arranged in circles around their mouths. – They are the simplest animals to have body symmetry and specialized tissues. – Cnidocytes are stinging cells that are located along their tentacles. – A nematocyst is a poisonfilled, stinging structure that contains a tiny coiled dart – used to paralyze and kill prey. After paralyzing its prey, a cnidarian pulls the prey through its mouth into its gastrovascular cavity. – Nutrients are then transported through the body by diffusion. – Cnidarians gather information from their environment using specialized sensory cells. – Most cnidarians reproduce both sexually and asexually • Most sexual reproduction takes place with external fertilization (outside the female’s body). PHYLUM PLATYHELMINTHES FLATWORMS KEY CHARACTERISTICS Three germ layers: ectoderm, mesoderm, endoderm Bilateral symmetry Cephalization is present Acoelomates - No coelom Use diffusion to transport materials through body No segmentation Movement w/ muscles or cilia Early development – protostomes Habitat water or moist dirt Basic body parts: mouth, pharynx, intestine, ganglia, flame cells MOVEMENT & REPRODUCTION IN FLATWORMS Free-living flatworms typically move in 2 ways: Cilia help them glide through the water. Muscle cells allow them to twist and turn. Most free-living flatworms are hermaphrodites (have both male and female reproductive organs). Sexual reproduction involves joining in a pair and delivering sperm to each other. Asexual reproduction takes place by fission – where the organism splits in two and each half grows into a new organism. GROUPS OF FLATWORMS • Turbellarians (picture #1): free-living flatworms that live in marine or fresh water • Bottom dwellers • Flukes (picture #2): parasitic flatworms • Infect the internal organs of their hosts • Tapeworms (picture #3): long, flat, parasitic worms • adapted to life inside the intestines of their hosts 1 2 3 FLUKES – PARASITIC FLATWORMS PHYLUM NEMATODA ROUNDWORMS KEY CHARACTERISTICS Three germ layers present – ectoderm, mesoderm, endoderm Unsegmented Bilateral symmetry Cephalization present Pseudocoelom present Mouth forms first – protostome Basic body parts: mouth, anus, intestines Examples: Pinworms & Nematodes Roundworms • Roundworms are unsegmented worms that have pseudocoeloms and digestive systems with two openings – a mouth and an anus – A pseudocoelom is a “false coelom” – a body cavity lined only partially with mesoderm • Feeding: most are carnivorous with grasping mouthparts and spines to catch and eat other animals • Respiration, circulation, and excretion: occurs via diffusion – they have no internal transport system • Response: simple nervous systems consisting of several ganglia • Movement: muscles extend the length of their body and allow for movement • Reproduction: sexual, nonhermaphroditic ROUNDWORMS & HUMAN DISEASE PHYLUM ANNELIDA – EARTHWORMS & LEECHES KEY CHARACTERISTICS Three germ layers present Bilateral symmetry present Cephalization present True coelom present Segmentation present Movement Muscles Mouth develops first – protostome Basic body parts: mouth, anus, crop, gizzard, body segments – Ex: earthworm FORM & FUNCTION OF ANNELIDS • Feeding & Digestion: range from filter feeders to predators – They extend their pharynx to collect prey and the food moves through the crop where it is stored and then through the gizzard where it is ground into smaller pieces • Circulation: they have a closed circulatory system – Blood is contained within a network of blood vessels • Respiration: gills (aquatic annelids) and diffusion (land annelids) • Excretion: digestive wastes passes out through the anus and liquid waste is eliminated by nephridia (filter fluid) • Response: have well developed nervous systems that include a brain and several nerve cords • Movement: have 2 major groups of body muscles as part of a hydrostatic skeleton • Reproduction: most reproduce sexually (although some use external fertilization) and some are hermaphroditic GROUPS OF ANNELIDS • Oligochaetes: Typically have streamlined bodies and relatively few setae (hairs) compared to polychaetes (most live in soil or fresh water). • Leeches: external parasites that suck the blood and body fluids of their host. • Polychaetes: marine annelids that have paired, paddlelike appendages tipped with setae (hairs). Polychaetes Oligochaete Leech PHYLUM MOLLUSKA – SNAIL, CLAM, SQUID, OCTOPUS, ETC. KEY CHARACTERISTICS Ex. Clam, squid, octopus, snail, oyster, scallop... Three germ layers present: ectoderm, mesoderm, endoderm Have an external or internal shell for support Bilateral symmetry & Radial Symmetry (Squid) Cephalization present Segmentation present Movement with muscles Has a TRUE COELOM Mouth develops first – protostome Basic body parts: gills, specialized “foot” THE MOLLUSK BODY PLAN BODY PLAN & OTHER PROCESSES • Body Plan: contains 4 parts – foot, mantle, shell, visceral mass – The muscular foot has many forms, including flat structures for crawling, spade-shaped structures for burrowing, and tentacles for capturing prey – The mantle is a thin layer of tissue that covers most of the mollusk’s body – The shell is made by glands that secrete calcium carbonate and has been reduced or lost in snails and some other mollusks groups – The visceral mass lies just below the mantle and contains the internal organs • Respiration: breathe using gills (aquatic) or by diffusion (land) • Circulation: either open or closed circulatory system – Open - blood is pumped through vessels by a simple heart and then diffusion into other systems – Closed – blood is transported through the body much quicker than an open circulatory system and is always contained in vessels • Excretion: nephridia • Response: complexity of nervous systems varies • Movement: secretion of mucous or jet propulsion • Reproduction: sexually by external fertilization, sexually by internal fertilization, and some are hermaphrodites GROUPS OF MOLLUSKS GIANT CLAM VARIOUS MOLLUSKS PHYLUM ARTHROPODA – CRAYFISH, SPIDERS, LOBSTERS, ETC. KEY CHARACTERISTICS Three germ layers present Exoskeleton present that requires molting (shedding) Jointed appendages Bilateral symmetry Cephalization present True coelom present Segmentation present Movement Yes (muscles & skeleton) Habitat In or Near Water Early Development Protostome – Ex. Insects, spiders, crayfish, millipedes, and centipedes CRUSTACEAN PARTS Cephalothorax: formed by fusion of the head with the thorax (which houses most of the internal organs) Abdomen: posterior part of the body Carapace: part of the exoskeleton that covers the cephalothorax Mandible: mouthpart adapted for biting and grinding food DECAPODS The largest group of crustaceans are the decapods: – They have 5 pairs of legs – The 1st pair of legs are chelipeds and bear large claws – The walking legs are the next pair – Behind the walking legs are the swimmerets (flipper-like appendages for swimming) Examples: crayfish, lobsters, crabs HORSESHOE CRABS • Horseshoe crabs are among the oldest living arthropods! SPIDERS & INSECTS • Horseshoe crabs, spiders, ticks, and scorpions are chelicerates – They have 2 pairs of appendages attached near the mouth • One pair are the chelicerae that includes fangs • The other pair, the pedipalps, are modified to capture prey – They have two body sections, a cephalothorax and abdomen – nearly all have 4 pairs of walking legs – They lack antennae – They respire using book gills or lungs • The largest group of arachnids are spiders! – Spiders are arthropods that do not have jaws for chewing, so they must liquefy their food to swallow it. – All spiders produce silk which is stronger than steel! • To spin silk into webs, spiders force liquid silk through spinnerets, which are organs that contain silk glands ANATOMY OF SPIDER & GRASSHOPPER INSECT RESPONSES Insects use a multitude of sense organs to respond to stimuli: – Compound eyes are made of many lenses. – Chemical receptors are used for taste and smell. – Sensory hairs detect slight movements in surrounding air or water. – Well developed ears detect sounds far above the human range. INSECT METAMORPHOSIS • The growth and development of insects usually involve metamorphosis – Metamorphosis is the process of changing shape and form – Most insects undergo either incomplete or complete metamorphosis • Incomplete metamorphosis: immature nymphs look much like adult form • Complete metamorphosis: immature larvae look nothing like parents INSECT COMMUNICATION & SOCIETIES • Many insects communicate using chemical signals called pheromones. – Insects can also communicate using sound, visual, and other types of signals. – Insect communication usually involves finding a mate. • Ants, bees, termites, and some of their relatives form complex associations called societies. – A society is a group of closely related animals of the same species that work together for the benefit of the whole group. – Within a society, individuals may be specialized to perform particular tasks in groups called castes. PHYLUM ECHINODERMATA – STARFISH, SEA URCHINS, SAND DOLLARS & SEA CUCUMBERS Eyespot Endoskeletal plates Anus Stomach Digestive glands Ring canal Radial canal Madreporite Reproductive glands Tube foot Sucker KEY CHARACTERISTICS 3 germ layers present Bilateral & Radial symmetry (adults) No cephalization in adults True coelom present Segmentation present Movement Tube feet & water vascular system Anus forms first – deuterostome!!! (the only invertebrate deuterostome) Basic body parts: spiny skin, endoskeleton, water vascular system, tube feet – Ex: starfish, sea urchins, sand dollars, sea cucumbers ENDOSKELETON & WATER VASCULAR SYSTEM • Echinoderms are characterized • A unique feature of echinoderms is a system of by spiny skin, an internal tubes called a water endoskeleton (internal vascular system: skeleton), a water vascular – This system is filled with fluid system, and suction-cuplike and carries out many structures called tube feet. – Most adult echinoderms exhibit five-part radial symmetry – The body parts are arranged around a central body like the spokes of a wheel essential body functions (respiration, circulation, movement) – It opens to the outside through filter-like structures called madreporites • A tube foot is a structure that operates much like a suction cup CLASSES OF ECHINODERMS • • • • • Sea urchins and sand dollars – Unique echinoderms in having large, solid plates that form a box around their internal organs – Many detritivores & Grazers Brittle Stars – Common in coral reefs; detritivores & filter feeders – Can shed one or more arms if attacked; arm will distract their predators Sea Cucumbers – Look like pickles; detritus feeders that move across sea floor Sea Stars – Carnivorous & prey on bivalves – Can repair itself when pulled apart, each section can become a new sea star if a portion of the central cavity still remains. Sea Lilies and Feather Stars – Filter feeders with long, feathery arms – Common in tropical oceans around coral reefs ECHINODERMS