Brachiopods & Bivalves: Revision Brachiopods Bivalves This table summarises the similarities and differences between brachiopods and bivalves. Articulate brachiopods Bivalves 1. Shell shape Two valves, one usually bigger Two valves, usually similar shape 2. Shell composition Calcite Calcite or aragonite 3. Shell symmetry Bilateral symmetry. Plane of symmetry along mid-line of each valve Bilateral symmetry. Plane of symmetry along plane where 2 valves meet 4. Valves Brachial & pedicle valves. (Pedicle valve bigger than brachial valve) Left & right valves. (Usually mirror images of each other) 5. Shell ornament Ribs, spines, growth lines Ribs, spines, growth lines 6. Hinge line Curved or straight Teeth on pedicle valve, sockets on brachial valve Curved or straight Teeth & sockets (if present) on hinge line of each valve 7. Hinge mechanism Teeth fit into sockets in opposite valve No ligament Teeth & sockets interlock Ligaments (external or internal) 8. Valve operation Two sets of muscles (adductor & diductor muscles) work together to open/close shell Ligaments hold valves under tension & one set of muscles (adductor muscles) contract (to close shell) or relax (to open shell) 9. Muscle scars At least two sets of muscle scars in each valve One or two muscle scars in each valve 10. Soft body parts & internal shell features Small body (& no simple pallial line or pallial sinus) No foot Most of shell cavity filled by filter feeding mechanism (lophophore), so shows brachidium (supports lophophore) Usually attached by pedicle (thick stalk or a series of fine threads) so shows a foramen Large body fills most of shell cavity (marked by pallial line) Siphons (position of retractable siphons marked by pallial sinus) Foot (for locomotion) No brachidium No pedicle, so no foramen (If attached - used byssal threads or cement for attachment) 11. Environment Marine Marine, freshwater, brackish water 12. Mode of life Epifaunal (attached by pedicle), Epifaunal (unattached & lying on sediment surface) Infaunal (burrowing) Semi-infaunal (unattached & partly submerged in sediment) Epifaunal (attached by cementation or byssal threads) Epifaunal (unattached & lying on sediment surface) Infaunal (deep burrowing or shallow burrowing) Free swimming Boring 13. Geological history Early Cambrian to Recent More common than bivalves until the mid-Triassic. After the mid-Jurassic less common than bivalves. Lower Cambrian to Recent Only more common than brachiopods in mid-Triassic. More common than brachiopods since the mid-Jurassic