UNIT 2--F98: ECOLOGY • • • • ALPINE TUNDRA CHAPTERS 46-50 IN TEXT READ CHAPTERS 48, 49 FIRST CHAPTERS 46, 47, 50 FOLLOW HAND-OUT NOTES ALPINE BOREAL FOREST--CLINGMANS DOME • WHAT IS IT??? • DEFINE DISCIPLINE • INTERRELATIONSHIPS OF “OLOGIES” HISTORICAL PERSPECTIVE • ARISTOTLE--4 th CENT B.C. – ATTEMPTS TO EXPLAIN PLAGUES OF MICE & LOCUSTS – CYCLES OF POPULATIONS HIGH/LOWS TIED TO WEATHER CONDITIONS • NATURAL HISTORY APPROACH – VARIOUS WORKERS: 17-1800’s TO PRESENT ECOLOGY • SCIENCE WHICH FORMULATES AND TESTS HYPOTHESES ABOUT ENVIRONMENTS • NOT SYNONYMOUS WITH ENVIRONMENTAL CONCERNS, I.E., CONSERVATION – EARTH DAY--APRIL 22, 1970 1 ECOLOGY: THE STUDY OF RELATIONSHIPS • ERNST HAECKEL--1870 (1866) • THOREAU (1858)-USED WORDS IN VARIOUS LETTERS, NOT DEFINITION • OTHER PUBLICAITONS RE: ISSUES IN 1600’s, 1700’s CHAS. KREBS • • • • • • THE SCIENTIFIC STUDY OF THE… …ABUNDANCE …DISTRIBUTION …OF ORGANISMS LEAVES OUT RELATIONSHIPS--…INTERACTIONS THAT DETERMINE THE DIST/ABUND OF ORGANISMS BASIC APPROACHES TO ECOLOGICAL STUDY • DESCRIPTIVE – NATURAL HISTORY APPROACH • FUNCTIONAL – RELATIONSHIPS, IDENTIFICATION & ANALYSIS OF PROBLEMS COMMON TO ALL AREAS • POPULATION & COMMUNITY STUDY – EVALUATE CAUSE/EFFECT RESPONSES OF POPULATIONS/COMMUNITIES TO ENVIRONMENTAL CHANGE – ALLOWS US TO ASK--HOW DOES THE SYSTEM OPERATE??? ECOLOGY: DEFINITION • THE STUDY OF RELATIONSHIPS OF ORGANISMS... • ...TO THEIR ENVIRONMENT • ODUM: – THE STUDY OF THE STRUCTURE AND FUNCTION OF NATURE – THE SCIENCE OF THE LIVING ENVIRONMENT REAL ISSUES IN NATURE • WHERE ARE THE ORGANISMS??? • HOW MANY ARE THERE?? • WHY??? • WHAT ARE THE FACTORS THAT AFFECT DISTRIBUTION??? BASIC APPROACHES (CONT’D) • EVOLUTIONARY • ORGANSISMS ARE HISTORICAL PRODUCT OF EVOLUTIONARY PORCESS & INTERACTION • ALLOWS FOR STUDY OF ULTIMATE CAUSES – WHY NAT. SELECTION FAVORS A PARTICULAR ADAPTATION – WHY DOES NAT. SEL. FAVOR A PARTICULAR ECOLOGICAL SOLUTION?? 2 PORTION OF EARTH’S SURFACE AND ATMOSPHERE WHICH CONTAIN LIVING SYSTEMS LIKENED TO THE “SKIN”OF AN APPLE TO TOTAL VOLUME OF EARTH BIOLOGICAL HIERARCHY • ECOLOGY PRIMARILY CONCRNED WITH 3-4 LEVELS • ECOSYSTEM • COMMUNITY • POPULATION • GOOD ECOLOGISTS HAVE FOUNDATIONAL UNDERSTANDING OF ALL LEVELS BIOLOGICAL HIERARCHY • • • • • • BIOSPHERE… ECOSYSTEM COMMUNITY POPULATION INDIVIDUAL…. ATOM CONCEPT OF ECOSYSTEM • AN AREA IN NATURE… • ...WHERE THERE IS INTERACTION... • ...AND EXCHANGE OF MATERIALS – ENERGY & NUTRIENTS • …BETWEEN BIOTIC (LIVING) AND ABIOTIC (NON-LIVING) COMPONETS ENERGY FLOW IN AN ECOSYSTEM 3 ECOSYSTEMS: STRUCTURE & FUNCTION • EXCHANGE OF MATERIALS –ENERGY –NUTRIENTS TROPHIC STRUCTURE • AUTOTROPHS – PHOTOAUTOTROPHS--SUNLIGHT ENERGY; GREEN PLANTS – CHEMOAUTOTROPHS--CHEMICAL En; CERTAIN BACTERIA • HETEROTROPHS – “ANIMALS”; GET ENERGY FROM EXTERNAL (OTHER ORGANISM) SOURCES TROPHIC STRUCTURE • AUTOTROPHS – PRODUCERS – PRIMARY PRODUCERS--CAPTURE ENERGY FROM SUNLIGHT – TRANSFORM SUNLIGHT En TO SOME OTHER FORM OF CHEMICAL En – USEFUL TO ORGANISMS • SUGARS, STARCH, ATP • HETEROTROPHS – CONSUMERS TROPHIC STRUCTURE TROPHIC STRUCTURE • HETEROTROPHS • TERTIARY CONSUMERS – PRIMARY CONSUMES • HERBIVORES • “SECONDARY PRODUCERS” • En SOURCE: FROM PRIMARY PRODUCERS-GREEN PLANTS – SECONDARY CONSUMERS • CARNINVOIRS • En SOURCE: PRIMARY CONSUMERS – CARNIVORES – En SOURCE: PRIMARY/SECONDARY CARNIVORES, HERBIVORES • DECOMPOSERS – DETRITIVORES – BACTERIA, FUNGI – En SOURCE: DEAD ORGANISMS (DEAD ORGANIC MATTER) 4 TROPHIC STRUCTURE • PRODUCERS • CONSUMERS • DECOMPOSERS ENERGY FLOW • LOWER TROPHIC LEVEL • NEXT HIGHEST TROPHIC LEVEL • CROSS-FEEDING BETWEEN TROPHIC LEVELS BY CARNIVORES • FOOD CHAIN • FOOD WEB CONCEPT ECOLOGICAL PYRAMIDS • “TRIANGLE” CONCEPT • ONE TROPHIC LEVEL, WITH GREATER ENERGY, BELOW NEXT HIGHEST TROPHIC LEVEL • ENERGY PYRAMID: – ALWAYS LARGEST AT BOTTOM OLD FIELD FOOD WEB ENERGY PYRAMID ENERGY PYRMAMID DECOMPOSERS SECONDARY PRODUCERS HERBIVORS PRIMARY PRODUCERS 5 ECOLOGICAL PYRAMIDS • ENERGY • BIOMASS • NUMBERS NUMBERS PYRAMIDS • SIMILAR SHAPE TO ENERGY & BIOMASS PYRAMIDS • SHAPE DETERMINED BY SAME LAWS THAT GOVERN ENERGY & BIOMASS PYRAMIDS • ALL BASED UPON ENERGY TRANSFER & EFFICIENCY BIOMASS PYRAMID NUMBERS PYRAMID • “TRIANGULAR SHAPE” • LARGER BASE---WHY??? • BASED UPON LAWS OF ENERGY TRANSFER, UTILIZATION BY NEXT HIGHEST TROPHIC LEVEL BIOMASS PYRAMID BIOMASS AND (NUMBERS) PYRAMID 6 ENERGY TRANSFER EFFICIENCY ENERGY BUDGET • 10% EFFICIENT BETWEEN TROPHIC LEVELS • WHAT HAPPENS TO OTHER 90%??? –HOW IS IT DISPERSED –IS IT LOST??? –ACCOUNT FOR IT ENERGY PYRAMID DECOMPOSERS ENERGY FLOW: 10 % EFFICIENCY SECONDARY PRODUCERS HERBIVORS PRIMARY PRODUCERS NITROGEN CYCLE NUTRIENT TRANSFER & RECYCLING • NUTRIENTS, SUCH AS NITROGEN, PHOSPHORUS, ETC. FLOW THROUGH ECOSYSTEM AS PART OF BIOMASS • FOLLOW FLOW OF ENERGY • ARE RECYCLED (BACK INTO SOIL) BY DECOMPOSERS ECOSYSTEMS DYNAMICS • ENERGY TRANSFERED BETWEEN TROPHIC LEVELS – 10% EFFICIENCY – 90% DISPERSED (HEAT, DEAD ORGANIC MATTER) • NUTRIENTS TRANSFERED BETWEEN TROPHIC LEVELS – RECYCLED FROM DEAD ORGANIC MATTER BY DECOMPOSERS 7 NUTRIENT BUDGET & RECYCLING GRAZING FOOD WEB BIOACCUMULATION/ BIOCONCENTRATION • ENVIRONMENTAL CONTAMINANTS (CHEMICALS) CONCENTRATE IN BIOLOGICAL TISSUES – FATTY TISSUE OF ORGANISMS • NOT BROKEN DOWN BY METABOLISM OF ORGANISM • TEND TO REMAIN IN BODY TISSUE FOR A LONG TIME (BIOCONCENTRATE) DETRITAL FOOD WEB DDT (INSECTICIDE) • APPLIED IN SMALL AMOUNTS TO SWAMPS FOR CONTROL INSECTS (MOSQUITOES) • CONCENTRATIONS OF 0.01 PPM • ABSORBED BY GREEN ALGAE--PRIMARY PRODUCES OF AQUATIC ECOSYSTEM • EATEN BY “WATER BUGS”, WHO ARE FED UPON BY SMALL FISH • SMALL FISH EATEN BY LARGER FISH • LARGE FISH EATEN BY MAN • DDT BIOCONCENTRATES IN FATTY TISSUE THROUGH OUT FOOD CHAIN • HEALTH EFFECTS ON MAN??? 8 OSPREY: POPULATIONS HAVE SIGNIFICANTLY INCREASED IN NUMBER SINCE THE BANNING OF CERTAIN PESTISIDES, SUCH AS DDT & RELATED CHLORINATED HYDROCARBONS (I.E., PCBs) 9