Chapter 35—Plant Structure, Growth, and Development Concept 35.1: The plant body has a hierarchy of organs, tissues, and cells. A. Plant cells—equip plants for specialized functions 1. Parenchyma cells a. least specialized b. thin walled c. storage 2. Collenchyma cells a. usually lack secondary walls b. uneven thickness c. support—elongates to continue support in young plant parts 3. Sclerenchyma cells a. functions in support b. rigid and thick c. do not elongate—may even be dead d. 2 types fibers (long) and sclerids (short) B. Organization Organs Systems Leaves Dermal Stems Vascular Roots Ground tissue 1. Dermal—epidermis a. aids in protection b. waxy cuticle helps plant retain water 2. Vascular tissue system a. xylem and phloem b. function in support and transport 3. Ground tissue system a. found between dermal and vascular tissue systems b. functions in photosynthesis, storage and support 1 4. Water conducting cells of xylem a. Trachiedslong and thin with parts to allow water to pass to next cell b. Vessel elementswider and shorter and arranged end to end Ends perforated to allow water to flow through vessel. c. Both dead at maturity 5. Food conducting cells of phloem a. Sieve tube members Alive at maturity Transport sucrose, compounds and minerals Arranged end to end with sieve tube plates Concept 35.2: Meristems generate cells for new organs 1. Plants have indeterminate growth 2. Parts of the plant have determinate growth (ex: flowers) a. apical meristemplant length primary growth (1º growth)—elongation secondary growth (2º growth)—increased girth b. lateral meristemscylinders of dividing cells extending along the lengths of roots and shoots secondary dermal tissue—replaces the epidermis add new layers to vascular tissue 2 Concept 35.3: Primary growth lengthens roots and shoots I. Angiosperm Body A. Roots—subterranean 1. Structured to anchor plants, absorb water and nutrients and store food 2. 2 Types of roots a. taproot (dicots) Large vertical root that produces smaller secondary roots Firm anchorage Maybe modified to store large amounts of food (carrot, turnip, sweet potato) b. fiberous root system (monocots) Mat of threadlike roots Provides extreme exposure to soil and water Prevent soil erosion Root hairs—help in water absorption—found in both systems B. Shoots—aerial (vegetative or floral shoots) 1. Stems a. nodes b. internodes c. axillary buds—dormant d. terminal bud—growth Axial buds may grow if damage occurs to terminal bud 3 II. Primary growth—extends roots and shoots by giving rise to the primary plant body. A. Roots 1. Root cap a. zone of cell division—closest to tip of root b. zone of elongation c. zone of maturation—furthest from tip 2. Tissues of roots a. protodermoutermost part of meristem contains root hairs b. procambiuma stele (central cylinder) phloem develops in stele c. ground meristemground tissue system stores food B. Shoots 1. Apical meristem is a dome-shaped mass of dividing cells at the tip of the terminal bud 2. Most shoot elongation occurs due to the growth of slightly older internodes 3. Axillary buds may form branches later in life 4. Tissue of stems a. Vascular bundles run length of stem Xylem and phloem 5. Modified stems a. stolonshorizontal stems (strawberries) “runners” b. rhizomehorizontal stems underground (irises) c. bulbsunderground shoots with modified leaves for food storage 4 C. Leaves 1. Main photosynthetic organ of plant 2. Monocotsveins run parallel to leaf 3. Dicotsveins can be pinnate or palmate 4. Leaves can be used to classify plants 5. Tissue organization of leaves a. epidermistightly locked cells protect prevents water loss b. specialized cells—stomata contained in epidermis are surrounded by guard cells control gas exchange for photosynthesis control transpiration—water loss c. ground tissue of leaf is mesophyll mainly parenchyma cells equipped with chloroplast d. leaf vascular tissue is continuous with stem vascular tissue and continuous from roots 5 Concept 35.4: Secondary growth adds girth to stems and roots in woody plants A. Lateral meristems create girth 1. Secondary vascular tissue a. vascular cambiumxylem and phloem b. fasicular cambiumwithin the vascular bundle c. interfasicular cambiumin the rays between vascular bundles d. ray initialsradial files of cells which permit lateral transport of water and nutrients as well as storage of starch and other reserves e. fusiform initialsproduce new vascular tissues; secondary xylem to the inside of the vascular cambium and secondary phloem to the outside 2. Periderm a. cork cambiumtough thick cover that replaces epidermis protects the plant body cylinder of fixed size and does not grow in diameter—creates rings in stems becomes woody B. Secondary growth of roots 1. Vascular cambium 2. Cork cambium Older roots become woody 6 Concept 35.5: Growth, morphogenesis and differentiation produce the plant body Development is the sum of all changes that progressively elaborate an organism’s body Growth is an irreversible increase in size resulting from cell division and cell enlargement Morphogenesis is the development of body shape and organization Cellular differentiation is the divergence in structure and function of cells as they become specialized during the plant’s development The cytoskeleton guides cell division and expansion by: 1. Orienting the plane of cell division 2. Orienting the direction of cell expansion—elongation is the primary direction of cell expansion, very little plant cell width occurs. A. Cell differention depends on gene regulation All cells in a plant posses a common genome This has been proven by cloning whole plants from single somatic cells. 1. All the genes necessary are present since these cells dedifferentiate in tissue cultures and then redifferentiate to produce the diversity of cells found in the plant 2. This ability that cellular differentiation is controlled by gene expression leading to the production of specific proteins 3. Different cells types selectively express certain genes at different developmental pathways that give rise to the diverse cell types 7 B. Pattern formation determines the location and tissue organization of plant organs pattern formation is the development of specific structures in specific locations pattern formation depends on positional information positional information are signals indicating a cell’s location relative to other cells in an embryonic mass C. The genetic basis of pattern formation in flower development The shoot tip in flowering plants shifts from indeterminate growth to determinate growth when the flower is produced -the meristem is consumed during the formation of primordial for sepals, petals, stamens, and carpels D. Positional information commits each primordium to develop into an organ of specific structure and function organ identity genes that function in development of the floral pattern are regulated by positional information mutations in these organ-identity genes can cause abnormal floral patterns 8