Habitat and Niche Habitat Type of environment in which a population or species regularly lives. Includes abiotic and biotic factors Examples of vegetative habitats Forest, meadow, pasture, coniferous forest, etc. Habitats Habitats can be subdivided into layers or zones Microhabitats Forest canopy: leaves, branches Shrub layer: leaves, branches, trunks Herb layer: leaves, stems, mosses, ferns Litter: dead fallen leaves, logs, Varies in depth, quality Soil: topsoil, subsoil Occupying Space A species may occupy a specific part of a microhabitat The limitations are often very distinct Example Hypogymnia imshaugii (a tube lichen) Lives on the upper part of branches of certain tree species Limited to a certain latitude Limited by rainfall and humidity Example 2 Western Rivers Rainbow trout can be found in highly oxygenated, faster-moving water and feeding on specific insects High concentration of dissolved oxygen Mountain whitefish can be found in less oxygenated, slowermoving water and feeding on different insects Low concentration of dissolved oxygen Fast current Slow current Niche (Pronounced “neesh”, but “nitch” is OK) Species’ or population’s role in its community Characteristics of a niche: Habitat & microhabitat (Space occupied) Food “spectrum,” essential nutrients Reproductive requirements Nutrition, nest/den sites Seasonality: When are resources required, used. Climate requirements: Humidity, rainfall, sunlight Niche Example Oregon little brown bat (Myotis lucifugus) Sleeping / reproduction space: rock crevices, caves, dense trees near water Seasonality / temporal aspects: Actively feeds at night – hibernates during the winter months Diet: Insectivore – prefer insects with aquatic life stages Range: Temperate coniferous forests Another species of bat may have a similar niche, but not exactly the same Two kinds of niche Fundamental niche All of the potential niche components for a given species if there were no competition for those resources Realized niche What the population actually gets while in competition with other species Niche diagram Here are niches for two species before competing for two resources Fundamental niches Niche diagram When the two species actually compete, there may be some overlap… Realized niches overlap leads to competition Niche No two species can occupy the same niche at the same time without intense competition one species will be removed Vacant niche: a niche that is not occupied by a species (not all scientists agree that this is real) example: rainbow trout in eastern Oregon have no gill parasites like some other fish – is this a vacant niche for some parasitic organism? Competition Organisms continuously compete for resources (space, food, mates, etc) Intraspecies competition: competition between organisms of a different species ie- bats and birds competing for a species of insect Interspecies competition; competition between individuals of the same species ie- two bats competing for a swarm of mosquitoes Competition Competition drives natural selection Co-evolution: in a natural setting, species continuously compete and evolve with each other and the evolutionary change or one is based on the evolutionary change of another Humans and our parasites Forest trees and fungi Invasive species are successful because they have not coevolved with the native species and can take over their niche New diseases Invasive plants Invasive insects Rabbits in New Zealand Reducing competition Different species often partition resources to reduce intraspecies competition Different species of birds will occupy different areas within the canopy of a tree to reduce competition from other species