Speciation Biology 2 Thursday, January 24, 2013 Evolution • Change in the genetic composition of a group of organisms over time. • Descent with modification Thursday, January 24, 2013 Microevolution • • Changes within species Short Time Period • e.g. Tuskless Elephants • e.g. Peppered Moths/Coloration • e.g. Bacteria/Antibiotic Resistance Thursday, January 24, 2013 Macroevolution • Changes over longer time periods • Speciation - evolution of 2 or more species from a single species Species B Species C Species A Thursday, January 24, 2013 Adaptive Radiation • One species evolves into many. Multiple Descendent Species Extinction Time Ancestral Species Thursday, January 24, 2013 Activity 4 Speciation of Caminalcules Thursday, January 24, 2013 Speciation of Caminalcules Gradualism Punctuated Equilibrium Time Morphology Thursday, January 24, 2013 Morphology Species • Biological Concept • Group of similarly constructed organisms that interbreed and produce fertile offspring Thursday, January 24, 2013 Reproductive Isolating Mechanisms • Prezygotic - prevents fertilization • Postzygotic - prevents development Good Summary Text Figure 17.4 Thursday, January 24, 2013 Prezygotic Isolating Mechanisms 1. Habitat Isolation - live different habitats Tree Canopy Ground Thursday, January 24, 2013 Prezygotic Isolating Mechanisms 2. Temporal Isolation - breed different times Squirrel Corn Thursday, January 24, 2013 Dutchman’s Britches Prezygotic Isolating Mechanisms 3. Behavioral Isolation - different courtships Blue-Footed Bobbies Dancing Bird Songs Fireflies Flashing Thursday, January 24, 2013 Prezygotic Isolating Mechanisms 4. Mechanical isolation - incompatible reproductive parts birch and passion flower pollen on passion flower Thursday, January 24, 2013 Postzygotic Isolating Mechanisms • Zygote Mortality • Hybrid Sterility Horse • Hybrid Fitness Mule Thursday, January 24, 2013 Donkey Species • Evolutionary Concept • Members of same species have similar structure • Can be applied to sexual and asexual reproduction. • Can be applied to fossils Thursday, January 24, 2013 Speciation • Evolution of 2 or more species from a single species Species B Species C Species A Thursday, January 24, 2013 Types of Speciation • Allopatric Speciation - geographical separation • Sympatric Speciation - same geographical area Thursday, January 24, 2013 Allopatric Speciation Textbook Figure 17.8 California Salamanders separated by large central valley Thursday, January 24, 2013 Allopatric Speciation Darwin’s Finches - Different Islands http://www.pbs.org/wgbh/evolution/library/01/6/ image_pop/l_016_02.html http://people.rit.edu/rhrsbi/GalapagosPages/SlideShow/ SlidePages/Darwin's%20Finches.html Thursday, January 24, 2013 Types of Speciation • Allopatric Speciation - geographical separation • Sympatric Speciation - same geographical area • Hybridization • Polyploidy Thursday, January 24, 2013 Hybridization • Crossing 2 different strains or related species. • Result in new species Wheat Type 1 Wheat Type 2 hybrid wheat Thursday, January 24, 2013 Polyploidy • Having more than 2 pairs of chromosomes 2N Diploid Thursday, January 24, 2013 3N 4N Polyploids Normal Meiosis Fertilization Egg/Sperm + Egg Sperm Thursday, January 24, 2013 Zygote Polyploidy • Chromosomes fail to separate in meiosis Egg Sperm + Thursday, January 24, 2013 Zygote Sympatric Speciation Polyploidy • Chromosome doubling due to no meiosis after replication Thursday, January 24, 2013 Wheat Evolution 7 14 Triticum gamete momococcum 2N = 14 Hybrid (Infertile) gamete Triticum turgidum 2N = 28 21 chromosome doubling Modern Wheat Triticum fauschi 2N = 42 Thursday, January 24, 2013 42