Marine Invertebrate Zoology The Hydrostatic Skeleton Phylum Cnidaria 1 Phylum Cnidaria The Hydrostatic Skeleton 2 Phylum Cnidaria The Hydrostatic Skeleton 3 The hydrostatic skeleton requires: Phylum Cnidaria Phylum Cnidaria Overview Defining Characteristics – – 4 Formation of complex intracellular organelles called nematocysts Planula larvae in the life cycle The phylum is composed of anemones, corals, sea whips, hydroids, and jellyfishes Diploblastic Phylum Cnidaria Diploblastic Animals 5 Epidermis Gastrodermis Phylum Cnidaria Nematocysts Used for gathering food or defense Produced in special cells called cnidoblasts SEM micrograph of nematocysts penetrating skin 6 Phylum Cnidaria Nematocysts 8 Phylum Cnidaria Cnidarian Nutrition 9 A major cnidarian drawback is the absence of an anus Carnivorous feeders on zooplankton and small fish Reef building corals also depend on symbiotic photosynthetic dinoflagellates for energy Phylum Cnidaria Reproduction and Nervous System Reproduce asexually and sexually – In the classes Scyphozoa and Hydrozoan the sexual stage is the medusa – – 10 Larvae - mouthless, ciliated, swimming planula Alternate between the polyp and medusa The polyp asexually produces a medusa that sexually produces a planula larvae, which then settles into a polyp Polymorphism = more than one stage per species Phylum Cnidaria Polyp & Medusa Polyp and Medusa Medusa (jellyfish) Usually solitary and free swimming The mouth and tentacles are directed downwards Contains thick mesoglea 12 Polyp (tube shape) Solitary or colonial, usually sessile attached by its base Mouth and tentacles are pointed upward Mesoglea is usually thin May have a fluid skeleton or calcium carbonate exoskeleton Phylum Cnidaria Cnidarian Classification Phylum Cnidaria – – – Class Scyphozoa Class Cubozoa Class Hydrozoa – Class Anthozoa 13 Order Hydroida Order Siphonophora Subclass Alcyonaria (Octocorallia) Subclass Zoantharia (Hexacorallia) Phylum Cnidaria Class Scyphozoa Defining characteristic – – 14 Asexual replication by strobilation Includes the true jellies, moon jellies (Aurelia) and Sea Nettles (Chrysaora) Phylum Cnidaria Scyphomedusa Structure Contains a large swimming bell that has tentacles and sensory capsules – Rhopalia Statocysts Ocelli Rhopalia 15 Phylum Cnidaria Medusa Movement 16 Phylum Cnidaria Scyphomedusa Structure 17 Phylum Cnidaria Scyphozoa Lifecycle Class Cubozoa Defining characteristics – 20 Medusa with box-like body Chiropsalmus quadrumanus (Sea Wasp) Phylum Cnidaria Class Cubozoa Known as box jellies or sea wasps due to shape and painful sting Occur in tropical and warm temperate seas Cubozoa Eyes 21 Phylum Cnidaria Class Hydrozoa 22 Due too small sizes and plant-like appearance people are unaware of their existence Display both polyp and medusa in the life cycle Mesoglea is thin or absent Phylum Cnidaria Hydroid Structure 23 Polyps can be solitary or colonial Very drab in appearance, no external skeleton, colonies are polymorphic Hydra Phylum Cnidaria Polymorphism Hydroid colonies have polyps of different functions – – – 24 Feeding - gastrozooids Defense - dactylozooids Reproduction gonozooids Phylum Cnidaria Hydromedusa Structure 25 Phylum Cnidaria Hydrozoan Reproduction Hydroid stage can reproduce by asexual budding – Eventually the bud detaches and becomes independent Gonozooids will release medusa – 26 Also has considerable powers of regeneration All medusa reproduce sexually Phylum Cnidaria Hydrozoa Lifecycle Order Hydroida Most species of the class hydrozoa in our area are hydroids Form sessile colonies of polyps and are usually mistaken as seaweed – 28 Exceptions are Velella and Porpita = hydroids that float freely at the surface Velella Phylum Cnidaria Order Siphonophora 29 Swimming or floating hydrozoan colonies Portuguese mano-war (Physalia physalis) Phylum Cnidaria 30 Phylum Cnidaria Class Anthozoa Defining characteristics – 31 Absence of a medusa stage Anemones, corals, sea whips, sea pansies Polyps consists of a column, flattened oral disk surrounding the mouth which bears tentacles Some polyps are solitary (anemones) others are colonial (coral) Phylum Cnidaria Class Anthozoa 32 Feed using tentacles that move food to the mouth Water is used by some as a fluid skeleton Produce planula larvae that settle and form new colonies Can also reproduce using pedal laceration and fission Phylum Cnidaria Subclass Zoantharia (Hexacorallia) 33 Posses 6 septa that separate the gastrovascular cavity Many species are solitary and lack a protective covering (sea anemones) Separated into two major orders Phylum Cnidaria Sea Anemones 34 Solitary and lack a rigid skeleton Have a broad adherent pedal disk Can reproduce asexually (pedal laceration) and sexually Phylum Cnidaria Stony corals Colonial with calcium carbonate skeleton secreted by the epidermis May be reef-building (hermatypic) or non reef building (ahermatypic) Hermatypic corals live in warm clear waters and can form chains around islands – 35 Zooxanthellae forms a symbiotic relationship with coral Phylum Cnidaria Subclass Alcyonaria (Octocorallia) Distinguishing characteristics – – 38 Sea whips, sea fans, and sea pansies are in this group Can harbor zooxanthellae Phylum Cnidaria