Brian Villmoare, PhD.
Center for the Advanced Study of Hominid Paleobiology
2114 G St., NW, Washington, DC
20052
Office: Rm. 301 /202 994 7152
bav@gwu.edu
Current Position
Research Scientist, George Washington University
Other Affiliations
Senior Research Associate, University College London
Previous positions held

2008-2011 Visiting lecturer – University College London Department of Anthropology

2006, 2008
Instructor – Arizona State University, Tempe, AZ. Course: Human Origins
(ASM 246)

2006
Instructor – Mesa Community College, Mesa, AZ. Course: Stones, bones, and
human evolution (ASM 104)

2003-2004, 2007
Laboratory instructor – Arizona State University, Tempe, AZ.
Course: Stones, bones, and human evolution (ASM 104)
Education

May 2008 – PhD Arizona State University
Dissertation: A geometric morphometric analysis of the midface of fossil hominins,
great apes, and modern humans

Feb. 1999 – MA Anthropology, Arizona State University
Thesis: An analysis of the single species hypothesis for African and Asian Homo
erectus

June 1990 – BA Philosophy & English Literature, University of Virginia
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Research Interests
Hominid and primate systematics, morphological integration, character analysis,
homology, geometric morphometrics, evolutionary theory.
Grants
2010 AAPA Career Development Grant
2006 Leakey Foundation Pre-doctoral Research Grant
2006 National Science Foundation Dissertation Improvement Grant
2006 ASU GPSA Grant
Peer-reviewed articles

2005
Villmoare B. Metric and non-metric randomization methods, geographic
variation, and the single-species hypothesis for Asian and African Homo erectus.
Journal of Human Evolution. 49: 680-701.

2011
Villmoare B., Fish J., and Jungers, W. Selection, morphological integration, and
strepsirrhine locomotor adaptations. Evolutionary Biology, 38: 88-99.

2011
Villmoare, B. and Kimbel, W. CT-based study of internal structure of the
anterior pillar in extinct hominins and its implications for the phylogeny of robust
Australopithecus. Proceedings of the National Academy of Sciences 108: 1620016205.

2011
Fish, J., Villmoare, B., Köbernick, K., Compagnucci, C., Tarabykin, V., and
Depew, M., Satb2, modularity, and the evolvability of the vertebrate jaw. Evolution
and Development 13: 549-564.

2012
Villmoare, B. Morphological integration, evolutionary constraints, and
extinction: a computer simulation-based study. Evolutionary Biology, DOI
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10.1007/s11692-012-9186-3
Manuscripts in preparation/review

2012
Villmoare, B., Dunmore, C., Kilpatrick, S., Oerteltd, N., and Fish, J.
Craniofacial modularity and the evolution of the mid-face in early African hominins. In
review

2013
Villmoare, B, Kuykendal, K, Rae, T, Brimacombe, C. Continuous dental eruption
and the age at death of Sts 5. In prep

2013
Tocheri, M. , Roach, N., Orr, C., Villmoare, B., Jungers, W., McFarlin, S.,
Senck, S., Kalthoff, D., Groves, C. Gorillas in the Midst: Rare congenital defects in the
hands and feet provide new insights into the evolution of eastern gorillas and the origin
of the grauer gorilla. In prep
Book Reviews

2012
Review of Larsen, Clark (ed.) A Companion to Biological Anthropology. The
Journal of the Royal Anthropological Institute, 18(2): 472–473.
Invited talks

2012
Studying Paranthropus from the inside out: a CT-based analysis of the anterior
pillar, and its importance for the phylogeny of early hominins. Given at the
Smithsonian Museum of Natural History, May 16, 2012.

2011
Why Paranthropus matters: what we can learn from the evolution and
extinction of the robust hominins. Given at University College London, May 15, 2011

2010
Integration in the face of Paranthropus – a geometric morphometric analysis.
Given at Roehampton University, Feb. 10, 2010.
Meeting abstracts

2012
Fish, J., Villmoare, B., Dunmore, C., Kilpatrick, S., Depew, M,. and Marcucio, R.
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A Study of Character: developmental approaches to modularity, integration and
evolvability of the craniofacial skeleton. Podium presentation at the annual meeting of
American Association of Physical Anthropologists in Portland, OR, April 11-14.

2010
Villmoare B, and Fish, J. Selection, morphological integration, and strepsirrhine
locomotor adaptations. (Invited) poster presented at session in honor of Elizabeth
Harmon at the annual meeting of American Association of Physical Anthropologists in
Albuquerque, NM, April 14-17.

2007
Villmoare B. Evaluating character independence for hominin systematics using
geometric morphometrics. Podium presentation at the annual meeting of American
Association of Physical Anthropologists in Philadelphia, PA, March 28-31.

2006
Villmoare B. Phylogenetic analysis using geometric morphometric data: the
infraorbital region of early African fossil hominins. Poster presented at the annual
meeting of American Association of Physical Anthropologists in Anchorage, AK, March
8-11.

2005
Villmoare, B. Geometric morphometric data of the infraorbital region as
discrete phylogentic data. Poster presented at the annual meeting of American
Association of Physical Anthropologists in Milwaukee, WI, April 6-10.

2001
Villmoare, B. Randomization methods to analyze variation in non-metric
characters, and the single-species hypothesis for Homo erectus. Poster presented at
the annual meeting of American Association of Physical Anthropologists in Kansas City,
MO, March 28-31.
Research and field projects
Currently Director of the Ledi-Geraru Project, Ethiopia.

2012
Senior Researcher – Koobi Fora Field School
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
2004
Researcher – Ledi-Geraru Survey Project, Ethiopia. Principal Investigators:
Prof. Charles Lockwood and Prof. Kaye Reed

2002
Researcher – Ledi-Geraru Research Project, Ethiopia. Principal Investigators:
Prof. Charles Lockwood and Prof. Kaye Reed

2001
Research Associate – Temporal bone morphology and hominid systematics: a
morphological approach. Principal Investigator: Prof. Charles Lockwood

2000
Instructor – IHO Field School at Makapansgat, South Africa. Principal
Investigator: Prof. Kaye Reed

1999
Instructor – IHO Field School at Makapansgat, South Africa. Principal
Investigator: Prof. Kaye Reed

1997
Crew Chief and Instructor – Cottonwood Wash Survey Project at Bluff, UT.
Principal Investigator: Nancy Mahoney
Professional Affiliations
American Association of Physical Anthropologists
Royal Geographic Society
Academic Service
Article referee – Journal of Human Evolution
Article referee – American Journal of Physical Anthropology
Article referee – Quaternary International
Grant reviewer – National Science Foundation
Graduate advisors
Dr. Charles Lockwood, Department of Anthropology, University College London.
Prof. William Kimbel, Department of Anthropology, Arizona State University.
Prof. Kaye Reed, Department of Anthropology, Arizona State University.
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Prof. David Strait, Department of Anthropology, SUNY Albany.
Collaborators and other affiliations:
Prof. Brian Richmond (CASHP and George Washington University), Prof. William Kimbel
(Institute of Human Origins and Arizona State University), Prof. William Jungers (SUNY
Stonybrook), Prof. David Strait (SUNY Albany), Dr. Jennifer Fish (Kings College), Dr.
Matthew Tocheri (Smithsonian Institute), Prof. Kaye Reed (Institute of Human Origins and
Arizona State University), Dr. Chris Campesano (Institute of Human Origins and Arizona
State University), Dr. Charles Lockwood, (University College London), Dr. Kristi Lewton
(Harvard University), Prof. Ramone Arrowsmith (Arizona State University), Dr. Kevin
Kuykendal (University of Sheffield), Dr. Anne Kandler (Santa Fe Institute).
Thesis Advisor and Post-graduate Scholars Supervised:
Major advisor:
Thomas O’Mahoney (MSc, 2009)
Lauren Wallis (MSc, 2009)
Nadja Oerteldt (MSc, 2010)
Filiz Altinolouk (MSc, 2010)
Christopher Dunsmore (MSc, 2011)
Shaun Kilpatrick (MSc, 2011)
Monica Nelson (MSc, 2011)
Hanna Enrith-Small (MSc, 2011)
Melanie Perritt (MSc, 2011)
Other
Certified Wilderness First Responder
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