The Late Lower Paleolithic Site of Holon, Israel: Subsistence, Technology, and Chronology Liora Kolska Horwitz1, Michael Chazan2, Adrian Lister3, Hervé Monchot4 and Naomi Porat3 1. Department of Evolution, Systematics and Ecology, The Hebrew University, Jerusalem 91004, Israel. 2. Department of Anthropology, 100 George Street, University of Toronto, Ontario M5S IA1, Canada. 3. Dept. of Palaeontology, Museum of Natural History, London UK 4. Department of Prehistory, Museum National d'Histoire Naturelle, Paris, France 5. Geological Survey of Israel, 30 Malkhe Israel Street, Jerusalem 95501, Israel. Abstract The open-air, Late Lower Paleolithic site of Holon is located on the southern coastal plain of Israel. Excavated between 1963 and 1970 by Tamar Noy, this site has produced one of the largest horizontal exposures of a Lower Paleolithic site with preserved fauna in the Near East. The publication of a monograph on the site in the spring of 2007 marked the end of a collaborative research program aimed at elucidating the timing and nature of hominin occupation at the site. This paper presents an overview of the results of this project focusing on evidence relevant to subsistence, technology, and chronology. Subistence: Holon is a palimpsest of kill/scavenge/butchery events in a swamp margin environment. Faunal analysis indicates multiple agents leading to bone accumulation at this locality including hunting and scavenging by hominins as well as carnivores. Technology: The Holon lithic assemblage includes handaxes, choppers, cores, flakes, and retouched flakes. The retouched flakes are dominated by sidescrapers with a high frequency of Nahr Ibrahim truncations. Technological analysis indicates an absence of the Levallois method and that handaxes were produced off-site. Chronology: OSL and ESR dating converge on a date in the latter part of OIS 7. We also explore the implications of these results for our understanding of Lower Paleolithic landscape use in the region, by comparing the data for Holon, an open-air kill/scavenge butchery site to preliminary results from the nearby site of Qesem Cave, which may have served as a Late Lower Paleolithic Levantine base camp, Key words: Late Lower Paleolithic, Holon, Qesem Cave, Near East, palimpsest site, kill/scavenge location, base camp Introduction Analyses of handaxe manufacture, morphology, and function have added significantly to our understanding of these enigmatic tools and the variability that underlies the seeming uniformity of the Lower Paleolithic (see articles in Goren-Inbar and Sharon 2006 and in the current volume). However, handaxes are best understood within the context of the relationship between hominins and the ecosystems they inhabited. Therefore, integrated interdisciplinary studies of sites where handaxes occur have the potential to provide important insights into the role played by these tools in Lower Paleolithic societies. In the first part of this paper we present a short overview of the Late Lower Paleolithic site of Holon, Israel, which has yielded an assemblage of 100 handaxes, in addition to over 1,000 other lithic artefacts and 500 identified faunal remains (Fig. 1). In the second part, we focus on a comparison of Holon with Qesem Cave, a Levantine Late Lower Paleolithic site which, to date, has yielded few handaxes (Barkai et al. 2003, Gopher et al. 2005, Karkanas et al. 2007). This comparison suggests a linkage between handaxes and sites like Holon which were used for butchery of a wide range of hunted/scavenged fauna, especially very large mammals (many well over 1 ton in weight). In contrast, the near absence of handaxes at Qesem Cave appears to be associated with a home-base economy focused on the exploitation of medium-sized animals. Taking this concept further, we propose that the generalized flake-based toolkits, characteristic of the Middle Paleolithic in the Levant (Shea 2003), emerged during Late Lower Paleolithic OIS 9-7 alongside more diversified toolkits that included handaxes. The comparison of Holon and Qesem Cave suggests that during the Late Lower Paleolithic of the Levant generalized flakebased industries are associated with sites best understood as base camps where food preparation and consumption took place (Table 1). Conversely, diversified toolkits with handaxes are often found in open-air sites associated with butchery of scavenged/hunted remains of very large animals. Consideration of a wider range of sites indicates that this dichotomy in toolkit may reflect differences in activities carried out at a site. The Site of Holon The Excavation The Late Lower Paleolithic site of Holon is an open-air site, located some 2km south of the city of Tel Aviv (Fig. 1). Following industrial development, salvage excavations were undertaken at the site by Dr. Tamar [Yizraeli] Noy, during 1963 and 1964, followed by a third season in 1970 (Yizraeli 1963, 1967, Noy and Issar 1971) (Fig. 2). The total area excavated was 120m2. In the site section, Noy (Yizraeli 1967), recognized five strata (Fig. 3): Stratum A a thick red sandy loam soil (hamra) Stratum B a dark clay layer. Stratum C a light grey clay representing the archaeological horizon. It reaches a maximum a thickness of 1.7 meters but is usually much thinner. Within this stratum, the excavator identified three levels (see section below). Stratum D is again a red sandy loam soil (hamra), reaching a maximum thickness of 50cm. Stratum E lies at the base of the sequence and represents a cemented sand (kurkar) Reconstruction of the site location (Netser and Chazan 2007) has shown that it formed on the edge of a marsh which developed following blocking of the outlet of the paleo-Ayalon River by incursive dunes (Fig. 4). Details concerning the excavation, site stratigraphy and site formation processes are given in the comprehensive final report of the site that has recently been published by Chazan and Horwitz (2007). Lithics The Holon lithic assemblage numbered 1,415 artefacts including 100 handaxes, 39 choppers, 160 cores, and 1,116 flakes - with retouch on ca. 50% of them including both sidescrapers and truncated-faceted pieces termed Nahr Ibrahim truncations (Fig. 5). Following the system developed by Roe (1968) 53 percent of the handaxes belong to the pointed category and 47 percent belong to the ovate category (Chazan 2007a). Mean length for the handaxes is 10.5 cm and mean breadth is 7.0 cm. Very few small flakes were found, probably due to selective retrieval during excavation. While the presence of cores indicate that flake knapping took place on-site, technological and raw material analyses demonstrate that the handaxes and choppers were produced off-site (Chazan 2000a,b, 2007a). Correspondence Analysis showed that with the exception of bifaces, all lithic artefacts are highly associated (Chazan et al. 2007, Monchot et al. in press). Tool class range in Levantine sites dating to this time-span are broadly similar, but differ in detail such as proportions of different artifact types, as well as the exclusion of some tool classes in certain assemblages. This may be due to the exploitation of different environments and/or raw materials. The variable nature of the Late Lower Acheulean tool kits has been discussed by Chazan (2007a,b) and others (Bar-Yosef 1994, Jelinek 1990, Gisis and Ronen 2006) and may reflect the increasing complexity of hominin behavior, perhaps related to specialized tools being used for certain functions. Fauna Of the 1,569 bones recovered during excavations at Holon, less than half i.e. 573 (36.5%), could be identified to species. Of these, typical Holarctic taxa were the most common species: fallow deer, Dama dama cf. mesopotamica (N = 247) (Lister 2007), aurochsen, Bos primigenius (N=162) (Horwitz and Monchot 2007) and straight-tusked elephant, Palaeoloxodon antiquus (N=120) (Davies and Lister 2007) (Fig. 6). Small numbers of remains of. red deer, Cervus elaphus (N=3), wild boar, Sus cf. scrofa (N=2) and freshwater turtle, Mauremys caspica (N=3) were also found. Similarly, few remains of Afro-Arabian species were identified, and comprised hippopotamus, Hippopotamus cf. amphibius (N=29) and gazelle, Gazella gazella (N=7). Compared to the number of identified bones (NISP counts), there is a relatively high minimum number of animals (MNI counts), comprising at least 6 elephants, 5 fallow deer, 3 aurochsen, 2 hippopotamus and individuals of red deer, gazelle, wild boar and freshwater turtle. Due to the small number of identified bones, little data on age profiles or sex ratios are available. It is however noteworthy that the straight-tusked elephant was represented by young and very mature animals, a profile that matches natural mortalities such as found today at water holes in Africa (Haynes 1988). For ungulates, all age groups are represented, including prime adults. Surface modifications to the bones were few but included signs of both hominin and animal activity. In both instances this was of low intensity with only 3.6% of bones in the assemblage exhibiting butchery damage resulting from hominin tool use - cut marks, flake scars and chop marks (Horwitz and Monchot 2002; Monchot and Horwitz 2007). Of the 63 cut marks, 15 were on bones of Dama dama cf. mesopotamica, 17 on bones of Bos primigenius, 1 on a Gazella gazella bone and 30 on unidentified bone splinters. A further 3% exhibited carnivore and rodent damage (gnaw marks, pits and puncture holes). Of the 6.6% bones with hominin or animal surface modifications, 2 exhibited both hominin and carnivore damage, 1 had both rodent damage and a cut mark, while another exhibited both rodent and carnivore damage (Monchot and Horwitz 2007). Attrition of the faunal assemblage due to diagenetic processes such as those related to bone mineral density, appears to have played only a minor role in the modification of the faunal assemblage from Holon. Consequently, the results of the Utility Indices, which could be calculated only for Bos and Dama, are probably reliable indicators. Both show a negative utility curve (Lyman 1994), with a high proportion of skeletal elements with moderate to low utility value, a picture that characterizes kill/scavenge sites where the high utility elements have been removed. The identification of Holon as a butchery/scavenge locality is corroborated by Correspondence Analysis which indicates that there is a low statistical association between different faunal taxa and skeletal elements such that bones of different taxa are not spatially associated (Chazan et al. 2007). These data suggest discrete scavenge/kill locations within the site, although no articulated bones were recovered. Late Lower Paleolithic Levantine sites which have yielded fauna are listed in Horwitz and Chazan (2007: Tables 13.2-13.3). Ungulates (especially fallow deer and aurochsen) are predominant in all assemblages, with medium-sized ungulates more common in caves as are carnivores. Large and extremely large terrestrial mammals (elephant, hippopotamus, rhinocerus and camel) are found in all sites, but their remains occur in far higher proportions in open sites. The poor representation of these very large-sized taxa in caves probably reflects selective transport strategies, with their meat but very few of their bones transported to the cave sites/base camps from the open air kill/scavenge sites. In contrast, carcasses of smaller-sized animals would have been introduced in a more complete state into the caves, although some transport selection against low utility elements would undoubtedly have been practiced. The Holon Ages - Are They Too Recent ? OSL ages on sediments from two pits excavated near the original excavation area at the site, and correlated on the basis of their geology with the archaeological section, gave the following sequence (from the top of section down to its base): 81±8 ka, (top paleosol); 150±13 ka (lower paleosol); 198±22 ka (archaeological level); and 240±17 ka (beachrock) (Fig. 3) (Porat 2007). ESR ages on two aurochsen teeth from Holon gave ages of 197±11 ka and 210±17 ka, respectively (Porat 2007). Thus both the ESR and OSL ages converge on ~200 ka for the occupation of Holon i.e. towards the end of marine Oxygen Isotope Stage (OIS) 7 (Porat et al. 2002, Porat 2007). The Holon ages, as well as those deriving from the Upper Acheulean sites of Oumm Qatafa D1 and Yabrud Cave I (Porat et al. 2002), have been questioned by several researchers as being too young (Bar-Yosef 1994, 1998b, Mercier et al. 2000, Rink et al. 2004). They have mistakenly claimed that dating was undertaken solely on material from museum collections and that no onsite dosimetry was practiced. In fact: the Holon ages were obtained using two separate and independent dating methods - OSL and ESR. for Holon, the OSL ages were obtained on fresh sediment samples recovered from newly excavated pits located next to the original excavation area. The sedimentary units in the pits was correlated with the original site stratigraphy using standard geological techniques i.e stratigraphy and sedimentology. ESR dating at Holon and Oumm Qatafa were based on on-site gamma dosimetry as well as using new sediments collected from the same strata as the teeth dated by ESR. For Yabrud I a sediment sample from the original excavation, originating from the same depth and general location as the dated tooth, was used to assess external dose rates and the ESR age is supported by TL on burned flint (Porat et al. 2002). the teeth used for ESR dating from all the sites derive from in situ deposits. the Holon ages accord extremely well with the known chrono-stratigraphy of the Israeli coastal plain (see Horowitz 1979, Gvirtzman et al. 1997, Netser and Chazan 2007). at no time have we claimed that, based on the Holon, Oumm Qatafa and Yabrud I ages, that the Late Lower Paleolithic of the Levant dates to 200 ka. We have argued that the transition from the Late Lower Paleolithic to the Middle Paleolithic falls within OIS 7, which ranges from 247 kyr to 183 kyr (Rohling et al. 1988). most importantly, many more Upper Acheulean OIS 7 ages are now available. As illustrated in Fig. 7, several other sites, dated by different researchers and using different methods, have also obtained ages for the Late Lower Paleolithic or Early Mousterian in the 200 ka range (Henning and Hours 1982; summary of ages in Bar-Yosef 1998a; Porat et al. 2002; Rink et al. 2004; Mercier et al. 2007). Thus, while the earliest ages for the Achuelo-Yabrudian are currently circa. 400 kyr the uppermost ages lie in the 150-200 kyr range (Mercier et al. 1995, 2000, summary of ages in BarYosef 1998). Although several Early Mousterian sites are now dated to 220-200 kyr (Mercier et al. 2007) we think that it is highly unlikely that the Upper Acheulean overlaps with the Middle Paleolithic. When the standard deviations for the ages, as well as the lack of fine resolution inherent in these dating methods is taken into account, as previously proposed by us (Porat et al. 2002), it is evident that the overlap of ages obtained from the two periods is an expression of the rapidity of the transition between the Late Lower Paleolithic (Upper Acheulean) and Early Middle Paleolithic (Early Mousterian). Is Holon a multi-phase site ? Based on Noy's stratigraphic observations, Bar-Yosef (1994, 1998b) suggested that Holon contains more than one archaeological level and that only artifacts from the main horizon had been published by Yizraeli (1967). In order to examine this contention and to test whether Holon represents a single archaeological assemblage, faunal and lithic material from the three different excavation seasons (1963, 1964 and 1970) were tested for differences. None were found between excavation seasons in the range of species or tool types represented, their relative proportions or even in their size (length, breadth, thickness), or even in size ratios (Chazan 2007b, Monchot and Horwitz 2007). Moreover, statistical tests, such as a variance mean ratio test, have demonstrated that the lithic and bone remains are spatially associated rather than randomly distributed (Chazan et al. 2007, Monchot et al. in press). As such it was confidently concluded that Holon represented a single horizontal archaeological occurrence. However, what of the vertical stratigraphy? The main find horizon of the site, Stratum C, is horizontal and lies at a depth of slightly above 38.00 m.a.s.l. It is a light grey clay attaining a maximum thickness of 1.70 meters. The excavator (Yizraeli 1967), identified three levels within this main horizon - Top: many chalk incrustations; Middle: very clayey with fewer chalk incrustations but with dense archaeological material; Bottom: sandier with few archaeological remains, mostly turtle. Although, there is evidence for some post-depositional vertical dispersal of material, due to movement of water through the soils, this transport was limited in scope and of low energy (Chazan 2007, Monchot and Horwitz 2007). Figure 8 presents a vertical projection of artefact and faunal provenience of the central area of the 1970 excavation. As illustrated here, the vast majority of archaeological material derives from the middle layer of Stratum C, which represents a single and clearly constrained archaeological horizon. In this profile, artefacts and fauna are clearly associated and it is difficult to discern any evidence for more than a single find horizon. There is however clear vertical dispersion of both artefacts and fauna. For the fauna, 39% of bones (N=960) had no depth measurements assigned to them. Of the 61% that could be analysed, the vast majority (82%) were recovered from a horizon 0.63 metres thick which was constrained between 37.51 to 38.14 m.a.s.l. A further 14% were found below this height between 38.15 to 38.99 m.a.s.l, while only 2% of bones were found either above 37.51or below 38.99 m.a.s.l. These results clearly indicate that the archaeological find horizon is indeed well defined. Moreover, no differences were found by depth in taxon representation or in the relative proportions of identified to unidentified bone remains. The prominence of turtle bones in the bottom, sandy level of Stratum C, which was noted by Yizraeli (1967), is not corroborated by the faunal analysis since only 3 such bones were recovered in the site. There is no method to determine the time duration represented by the palimpsest of activities that created the association of faunal and lithic remains found at the site of Holon. This is not a problem unique to Holon but, as recognized by Stern (1993), is an intrinsic element of the Lower Pleistocene archaeological record. As noted above, the nature of the vertical distribution of both artefacts and fauna indicates that the material was deposited during a constrained period of time. The condition of the faunal remains (Monchot and Horwitz 2007) point to rapid burial. Finally, there is no indication for variability in lithic typology or technology that would support the identification of multiple periods of occupation (Chazan 2007a, b). Conclusions with Respect to Holon Holon should best be interpreted as a multiple mortality/scavenging open-air location rather than as a base camp given the: broad spectrum of animals (and habitats) represented the high minimum numbers of identified animals (MNI counts) relative to the number of identified bones (NISP counts) a similar low intensity of hominin damage on bones (3.6%) and damage caused by animals (3% of bones) presence of bones and teeth of large sized species - elephants, hippopotamus - species whose skeletal remains are not readily removed although their meat may be taken offsite lack of evidence for fire/hearths lack of evidence for redistribution of food since the redistribution model described by Isaac (1984) for base camps is likely to result in discard areas with mixing of portions of different species and carcasses. This was not the case at Holon since there was no statistically significant spatial association between bones of different taxa or skeletal elements (Chazan et al. 2007, Monchot et al. in press). This demonstrates that the individual remains entered the site as isolated carcasses and were butchered as such. Due to its proximity to the river, Holon would have offered a mosaic of environments with excellent opportunities for both hunting of a wide range of animals as well as scavenging natural mortalities on the river banks. Following repeated visits by hominins to this favoured marsh locality on the edge of the paleo-Ayalon river, over a circumscribed period of time the remains of animal carcasses and lithic artefacts would have accumulated. Thus, Holon did not function as a base camp (as defined by Isaac 1984) but rather served as a locality frequently revisited (perhaps even seasonally), where intercept hunting/ambushes, as well as scavenging took place ( see discussion in Chazan and Horwitz 2006), in the same manner as suggested by Rollefson (1985) for the Lower Paleolithic site of Fjaje, Jordan. As defined by Chazan and Horwitz (2006: 444), Holon was a palimpsest site "representing multiple discreet events i.e. numerous SCS [single carcass scatters], which have taken place in adjacent areas or were even super-imposed on earlier activities". A site such as this would be geared towards primary food acquisition with limited food preparation and consumption activities taking place. Consequently, it is plausible that Holon represents just one aspect of a more complex pattern of Late Lower Paleolithic hominin landscape use. Indeed, with reference to the Holon assemblage, the presence of other types of sites, including base camps, elsewhere in the Levantine landscape is hinted at by: (a) a predominance of medium to low utility skeletal elements suggesting that high utility, meatrich elements were transported off-site (Monchot and Horwitz 2007). (b) a difference in butchery patterns between the large sized Bos and smaller sized Dama demonstrating that Bos carcasses were cut up into smaller parcels of meat and bone, while more complete carcasses of Dama were transported off-site (Horwitz and Monchot 2006). (c) bifaces and choppers were not manufactured at Holon but introduced into the site readymade (Chazan 2007a). A possible example of a Late Lower Paleolithic base camp from Israel that may complement Holon, are the upper levels of the Acheulo-Yabrudian site of Qesem Cave (Table 1). Qesem Cave - A Probable Late Lower Paleolithic Base Camp Qesem Cave, located some 12km east of Tel Aviv, was discovered during the course of highway development, during which the cave roof was removed. Several excavation seasons undertaken since 2000 have resulted in an area of some 12m2 being opened to a depth of 7.5 meters from bedrock to the uppermost layer (Barkai et al. 2005; Gopher et al. 2005; Lemorini et al. 2006). A series of U/Th series dates indicate that site occupation began before 382 kyr and ended before 152 kyr, possibly around 200 kyr (Barkai et al. 2003). The picture obtained from the fauna spectrum is remarkably dissimilar to that found at Holon (Gopher et al. 2005, Lemorini et al. 2006) (Table 1). Unlike Holon, the fauna present are solely Holarctic with no Afro-Arabian forms which are found at Holon or in Levantine Early Mousterian sites (e.g. Shea 2003, Stiner 2005, Tchernov 1988, 1998). The diet was more focused than at Holon, with >80% representing fallow deer (Dama mesopotamica), predominantly mature individuals that would have been of prime interest to prehistoric populations as a meat source. Smaller quantities of aurochsen (Bos primigenius), caballine equids, wild boar (Sus scrofa), tortoise (Testudo graeca) were found, while red deer (Cervus elaphus) is rare. Notably, no bones of very large mammals – elephant, rhinoceros or hippopotamus- were recovered in the upper levels of Qesem Cave. As illustrated in Table 2, preliminary results on the taphonomy of the upper layers at Qesem Cave (Gopher et al. 2005, Lemorini et al. 2006) indicates an inverse picture for the representation of skeletal elements of Dama to that found at Holon; with a predominance of limbs and cranial bones (no antlers) and absence of trunk elements, pelves and foot bones. Indeed, Gopher et al. (2005:86) note that "hominids were selective about the body parts they transported to the cave, presumably following field processing of the carcasses elsewhere." Moreover, there is a very high frequency of cut marks on the Qesem bones, numerous cone fractures associated with marrow extraction, evidence for burning, and no rodent damage while carnivore damage was observed on only 1 bone out of some 2000 bones examined. These data contrast markedly to the results for Holon, suggesting that a significantly different range of activities took place at Qesem Cave and included food preparation and consumption, features more characteristic of a base camp. With regard to the lithic assemblage (Table 1), Qesem Cave has yielded only 4 bifaces (teardropshaped hand axes with sharpened edges), compared to the 100 from Holon. Blades, the most common artefact type at Qesem appear to have been used predominantly in butchering activities - cutting and defleshing (Barkai and Gopher 2005; Gopher et al. 2005; Lemorini et al. 2006). Conclusions In many ways the Late Lower Paleolithic of Qesem Cave anticipates the characteristics of Early Middle Paleolithic sites such as Tabun D, Hayonim E, and Misiliya (Jelinek 1981, WeinsteinEvron et al. 2003, Bar-Yosef et al. 2005). In terms of lithic technologies all these sites share an emphasis on the production of elongated flake/blades, although the technological strategies employed appear to differ markedly. The associated faunal assemblages demonstrate an emphasis on fallow deer and gazelle with high proportions of burnt, cut and fractured bones attributed to hominin food extraction and preparation activities (Stiner 2005, Yeshurun et al. 2007). It is interesting that at Misilya the bovid assemblage shows clear evidence of differential butchery with the larger fallow deer, undergoing partial field butchery, while the small sized gazelle were introduced as more complete carcasses (Yeshurun et al. 2007). The comparison between Holon and the upper levels of Qesem Cave suggests a dichotomy between butchery sites and home base sites in the Late Lower Paleolithic of the Levant. It is interesting that in this comparison handaxes are associated with primary butchery at the kill/scavenge locality. Thus it is possible that by the Late Lower Paleolithic handaxes had come to be used for tasks associated particularly with processing of large and very large animals. The employment at this time of a generalized flake- based toolkit, which becomes the hallmark of the Middle Paleolithic, is most clearly associated with butchery of small and medium sized animals within the context of home base sites. Clark and Schick (2000:199) have suggested that "The association at some sites of full sized bifaces with the small flake and core component [such as found at Holon], may be an indication of site reuse or of new individuals joining the group, but in every case the flake component always indicates the later stages of the reduction...suggesting the cores were reduced initially at another locale where the cortical flakes were left behind". A consideration of the broader context suggests that some caution should be employed when drawing overly broad conclusions from this comparison since the cave sites of Yabrud I, Tabun, and Oumm Qatafa all yielded a diversified lithic assemblage that included an important component of handaxes (Neuville 1951, Gilead 1970, 1977, Jelinek 1981, 1990, Bar-Yosef. 1994). Since these are caves, it is unlikely that any served as primary kill/scavenge sites in the same manner as Holon. It is of further interest that the distinction between the Yabrudian and Acheulo-Yabrudian is based upon a fluctuation in the frequency of handaxes.. The comparison of Holon and Qesem Cave raises the possibility that this fluctuation may not be the hallmark of a cultural tradition but simply a reflection of shifts in the activities carried out at the sites. Acknowledgements We wish to express our appreciation to: Prof. H. de Lumley and Eric Boëda for inviting LKH and MC to attend the Tautavel conference and to The French Embassies in Israel and Canada for facilitating the attendance of the Tautavel conference of LKH and MC respectively. Research on aspects of Holon was financed by grants from The Canadian Social Sciences and Humanities Research Council and The Halbert Academic Exchange Fund (University of Toronto) (to MC), and the TMR program (co-ordinated by Prof. Naama Goren) and a Fyssen Grant (to HM). References Barkai, R, Gopher, A, Lauritzen, SE, Frumkin, A., 2003. 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Weinstein-Evron 2007. Modern hunting behavior in the early Middle Paleolithic: Faunal remains from Misliya Cave, Mount Carmel, Israel. Journal of Human Evolution 53, 656-677. Yizraeli (Noy), T. 1963. Holon. Israel Exploration Journal 13, 137. Yizraeli (Noy), T. 1967. A Lower Paleolithic Site at Holon. Israel Exploration Journal 17, 144152. Figure Captions Figure 1: Map of the Levant showing the location of Holon and other sites discussed in the text. Figure 2: Location of the different excavation seasons at Holon and recent Pits A and B sampled for OSL dating. Figure 3: Correlation of stratigraphy and OSL dates between the excavation section and Pits A and B. Figure 4: Location of site relative to the reconstructed course of the paleo-Ayalon river. Figure 5: Lithics from Holon. 1: Simple concave sidescraper; 2: Double sidescraper with Nahr Ibrahim truncation; 3: Core; 4: Handaxe; 5: Chopper. (Drawings by A. Sumner and M. Chazan) Figure 6: Palaeoloxodon tusk recovered in situ at Holon Figure 7: Graph showing range of dates for Levantine Late Lower Paleolithic sites and Early Mousterian sites based on several different radiometric methods Figure 8: Vertical projection of lithic and faunal provenience for the central area of the 1970 excavation at Holon. Note the complete mixing of faunal and lithic components, as well as of bifaces and other artefact types. Table Captions Table 1: Comparison of Holon and the upper levels of Qesem Cave Table 1: Comparison of Holon and the upper levels of Qesem Cave FEATURE HOLON QESEM (Upper Levels) Area excavated Dating Site type SITE 120 m2 x 2m depth 200 Ka Open air, adjacent to a river 12m2 x 7.5m depth 207-152 Ka Cave FAUNA N= 1,569 N=1,780 Faunal sample studied Most common faunal species Very large mammals Skeletal element representation Dama 43% of total id; 16% of total fauna Present (Paleoloxodon, Hippopotamus) Dama- complete Bos – low utility elements Burnt Bones Absent Cut marks Cone fractures Few – on 3.6% of bones Questionable if present - only isolated instances Present – on 3% of bones Common on many bones Carnivore activity Weathering Lithic density Bifaces Biface manufacture Dominant artifact type LITHICS Sparse: N = 1,415 but deposit was not sieved Many: N = 100 No handaxe debitage suggesting off-site manufacture Flakes, sidescrapers, Nahr Ibrahim truncations Dama 89% of total id Absent Dama – only cranial (no antlers) and limb elements Bos - few bones Common - 36% of all bones; 84% of unidentified bone splinters Many - on 19% of bones Common - on 38% of bones Absent Rare - 2% of bones Very dense: N = 50,000 items Few: N = 4+1 preform Presence of large flakes for fashioning bifaces suggesting they were made onsite Blades CONCLUSION Disarticulation of carcasses = Exploitation of meat and bone marrow Hunting/Scavenge site and cooking = Base camp