81 LABORATORY EXERCISE VII PHYLUM ECHINODERMATA echinoderms These spiny animals have a radial symmetry that is secondary, i.e., it is derived from a bilateral ancestor. Echinoderm symmetry is also described as pentamerous—can you determine what this means? The generally hard endoskeleton is due to plates, called ossicles, which are joined together in various ways. Echinoderms are coelomate and possess a unique body system called the water vascular system. This system is involved in locomotion in many echinoderms. CLASS ASTEROIDEA sea stars or starfish Sea stars typically have five arms that grade into the central disk, but there is much variation, as seen in the specimens on demonstration. Asterias Starfish arm c.s. Slide (Box V slide tray 8; 4x) Be able to recognize the ampullae, podia and suckers of the tube feet, which are part of the water vascular system. Be able to distinguish them from the coelom. Also be able to find the pyloric ceca and the radial canal. Asterias Sea star specimen The common starfish of the New England coast may be used to illustrate the general echinoderm structure. Be able to label the tube feet, madreporite, central disk and the mouth. Starfish diversity Both wet and dry preserved specimens Be able to recognize the starfish on demonstration as members of the phylum Echinodermata, class Asteroidea. Sea stars have remarkable powers of Regeneration—with at least 1/5 of the central disk present, a sea star will regenerate. One of our specimens shows the regenerative process. CLASS OPHIUROIDEA serpent stars or brittle stars The five arms of serpent stars are usually distinct from the central disk. 82 Brittle stars specimens Notice that each arm is distinct from the central disk. Notice the articulated plates of the skeletal system of the arms that give them great flexibility and mobility. Basket starfish or Gorgon's head These highly modified ophiuroids are common in the cold ocean waters of northern Maine and Canada and is also found in the tropics. But they are not common in temperate waters. CLASS ECHINOIDEA Echinoids are without arms and are spherical or flattened in form, but they still retain the pentamerous symmetry characteristic of the phylum. Spines are very noticeable in most echinoids and vary in shape, size, and number from species to species. The class includes the spherical sea urchins and the flattened cake urchins, sea biscuits, and sand dollars. Sea urchins The numbers and shape of spines differ greatly among sea urchins, as demonstrated. Look for the tube feet. Sea urchin skeleton The bleached-out skeleton of sea urchins, common in many "shell shops," is made up of fused ossicles. In living animals, spines are located on the knobs, and tube feet project through the holes. Using this skeleton, can you discover evidence of pentamerous symmetry? Aristotle's lantern The spherical sea urchins possess a unique and intricate oral grinding device known as Aristotle's lantern. They use this structure to feed on algae growing on rocks or coral. 83 Cake urchin skeleton The cake urchin or sea bisquit comes from shallow waters in the subtropical and tropical Atlantic. Living specimens have spines similar to sea urchins. Can you see the pentamerous symmetry Sand dollars (preserved and bleached skeleltons) These sand dollars are common in waters along coastal Carolina. Actually, they can be found in virtually any shallow water marine habitat. Note the lunules—holes in their skeletons. The function of these structures is not known but may help the animal maintain stability in currents. CLASS HOLOTHUROIDEA sea cucumbers Sea cucumbers are usually elongated and soft creatures. Ossicles are reduced to individual microscopic plates imbedded in the skin; there are no spines. Feeding tentacles usually surround the mouth, which are highly modified tube feet. Sea Cucumbers What evidences of pentamerous symmetry do you see on these specimens? Dissected specimen This dissection is to allow you to examine the respiratory trees, a unique holothuroidean structure. Also be able to locate the gut and the gonads. CLASS CRINOIDEA sea lilies and feather stars Extant sea lilies are stalked and attached deep water forms. Feather stars are without stalks and can swim and walk short distances. Crinoid fossil Notice the stalk and the arms of this fossilized crinoid. Crinoids dominated during the Paleozoic but today there are only about 500 extant species. 84 PHYLUM HEMICHORDATA acorn worms Acorn worms are delicate, though usually large inhabitants of marine mud. They have pharyngeal gill slits and a dorsal, occasionally hollow nerve chord (thought by most scientists to be homologous to the chordate nerve chord). But they have no notochord (although early researchers mistakenly thought they did, hence they were once classified as a chordate). Thus we call them hemichordates. Hemi is Greek for half. Acorn worms Notice the collar, proboscis and trunk. These marine worms usually live in muddy bottom sediments.