Seattle Urban Nature Project Habitat Delineation and Descriptions September 2002 This document contains information about how the data were collected, a description of the habitat categories, and a glossary of terms. Formal descriptions are provided within the document for italicized terms. Data Collection and Habitat Delineation Techniques These data were collected using vegetation field surveys and aerial photographs in which landscape features, such as trees and other vegetation, are visible. Areas that contain similar vegetation (a.k.a. habitat type) were mapped by drawing lines on the photos. Areas inside these lines are called polygons. Polygon shapes can be simple, as in a square drawn around a parking lot, or complicated and irregular, as in a winding line drawn around a grove of trees. Polygon borders were digitized and made into a GIS data layer of habitat types. Habitat classifications/categories were based on the Seattle Department of Parks and Recreation’s “Urban Wildlife and Habitat Management Plan” July, 1994. Developed Areas: Features that could be clearly seen on aerial photographs, such as parking lots, were first delineated. Nearby buildings and impervious surfaces were often included in the polygons. Other developed areas contained dense paved trail networks, cemeteries, long, thin areas with a trail or sidewalk in the center, playgrounds, and picnic areas. Depending on the proportion of impervious surface, the area is designated as a “light,” “medium,” or “heavy development” habitat type. Landscaped Areas: Next, landscaped areas (e.g. areas where shrubs and trees are pruned or mulched, lawns are mowed, and/or flowerbeds, paved sidewalks, baseball diamonds and playgrounds exist) were field mapped. Any area with these conditions and with less than 10% impervious surface qualified as a landscaped area (aesthetic criteria do not apply). © 2002 Seattle Urban Nature Project. All rights reserved. 1 Natural Areas: Finally, the more naturally vegetated areas of the parks were surveyed using the aerial photos. Areas with similar vegetation types (see Habitat Key) were identified. If the boundaries of these areas were easily identified on the aerial photos, boundaries were drawn on the photos at tree canopy edges. In many areas, boundaries between habitat types (e.g. representative of a change in size class of a forest without a change in the type of trees or wetlands under a tree canopy, etc.) were difficult to delineate on the photographs. When this occurred, the area was surveyed from a location identifiable on the photograph and used measuring tapes and compasses to determine the boundaries. Wetlands: To determine the presence of a palustrine (freshwater wetland) system, the prevalence of hydrophytic (“water-loving”) vegetation, surface water present in the late spring and summer, and/or the existence of deep, mucky soils were identified. These observations provided very conservative (smaller) estimates of wetlands areas. The criteria required by local, state, and federal agencies for jurisdictional delineations includes information about soil types, depth of ground water and the presence of hydrophytic vegetation. Using these criteria to determine the wetland area would have expanded the number and sizes of wetlands already identified. In many areas, the canopy over wetlands includes overhanging branches from trees rooted in uplands (non-wetland areas). These wetland areas are thought to function in many ways as “forested wetlands,” although some vegetation classifications require that the trees be hydrophytic for the area to be considered a “forested wetland.” However, although a canopy may not include hydrophytic vegetation, the shading, perching and foraging contributions of the trees in close association with water, wet soil, and wet plants can create many qualities of a “forested wetland.” Roads and Trails: In order to create a reasonably readable map, only the largest and most durable roads and trails were assigned their own polygons (see Roads and Trails Criteria). In the future, an independent road and trail data layer would be very useful for depicting the fragmentation of some habitats. Mapping Diverse and Transitional Areas: The intent of this survey was to show major habitat areas. In general, areas of similar “character” or ecology were grouped. Habitat boundary definitions were often difficult to draw and subject to interpretation. It is important to recognize that boundary definition determination is a subjective process; not only will classifications vary between staff members, but one person may define the same polygon in an alternative fashion on different days. The polygons are intended to serve as a tool to help study and inventory an area, not to act as barriers within ecosystems. However, the database design is able to accept further refinements as they come to light. The character of a forest seldom changes abruptly. Putting a discrete line between areas that have a subtle transition or layer transition from one habitat type to another often © 2002 Seattle Urban Nature Project. All rights reserved. 2 requires a compromise in the boundaries of one of the habitats. For example, when tree canopies overhang the edges of parking lots, roads, lawns and shrubs, the edges of the polygons could be drawn at the edge of the pavement or grass or at the edge of the tree canopy. It was determined that where tree canopies overhang impervious surfaces, the impervious surface has a greater effect on the habitat of the polygon than the overhanging trees, so the boundary would be placed at the edge of the impervious surface. However, when tree canopies overhang other plants, the tree canopy has a greater effect on the habitat of the polygon, so the boundary was placed at the edge of the tree canopy. In areas where species or sizes were mixed, questions arose concerning whether or not, for example, an area with the character of an older forest should encompass only the groves of large trees or should include groves of smaller trees. Mixed stand polygons were created when coniferous, deciduous, and broadleaved evergreen trees were found within the same area. If a patch covered more than 30% of an area a new polygon was created (see Habitat Key). Features larger than one half acre were mapped, except wetlands which were mapped in smaller areas. Small groves of one forest type (e.g. a size class different than the surroundings or patches of conifers or Madrone in a mostly deciduous forest, etc.) were often grouped for inclusion into the surrounding polygon. Their presence was recorded in the habitat description or vegetation table on the data sheet for the polygon. Describing the Habitat: Once the perimeter of a polygon was established, it was given a unique identifying code that indicated where it was located (e.g. discpk1 is the first polygon in Discovery Park). Cover class was assigned according to the percentage of the polygon that is vegetated, based on data collected on plants and non-vegetated areas (e.g. “pavement”, “building”, “sand”, etc.) within polygons. In instances when polygons had multiple layers of canopies (trees, shrubs, herbaceous groundcovers, etc.), total coverage was often greater than 100% because each layer was assessed separately. The canopy layer(s) occupied by each species were indicated. Data on special site features were collected (see checkboxes) and distinctive characteristics were described (see Habitat Descriptions). Habitat Descriptions: Developed Landscapes: Heavy Development: Buildings, parking lots, etc. (>50% impervious surface). Medium Development: Housing areas with extensive vegetation, play areas, picnic areas, dense paved trail networks (30-50% impervious surface). Light Development: Small yet evident presence of pavement, small structures, etc. (1030% impervious surface). Landscaped Grassland: Mown grass. Landscaped Shrubland: Pruned or shaped shrubs (e.g. a rose garden). Landscaped Tree Savannah: A few trees scattered over mown grass or manicured shrubs. Landscaped Forest: More trees over mown grass or manicured shrubs. © 2002 Seattle Urban Nature Project. All rights reserved. 3 Herbaceous Row Crop: Any food garden, generally restricted to the City P-Patches. Developed Pasture: Uncommon. Large tracts of land that are grazed by horses (seen) or other animals (not seen). Orchards: Restricted to old farm remnants. Uncommon. Open Canopy: Beaches and Dunes: Restricted to sandy beachfronts. Sparsely Vegetated Area: In Seattle, restricted to areas of landslides and formally graded areas that have not revegetated. Uncommon. Grassland: Uncut grass, fields. Shrub Savannah: A few shrubs scattered over uncut grass. Tree Savannah: A few trees scattered over uncut grass, shrubs or a mixture. Shrubland: Shrubs dominate the upper canopy.. Wetland Types: Riverine Consolidated Substrate: Perennial or robustly intermittent streams with an artificial bed. Riverine Unconsolidated Substrate: Perennial or robustly intermittent streams with direct soil connection. Riverine Tidal: Wetlands along large rivers whose hydrology is linked to tidal cycles. Forest Types: Riparian Forests: Contains many size classes and potential many species. Common trees are Cottonwood (Populus trichocarpa), Willow (Salix spp.), and Red Alder (Alnus rubra). Western Red Cedar (Thuja plicata) and Spruce (Picea spp.) are present and much less common. Unlike riverine systems, streams in riparian forests are well connected to the flood plain and the surrounding hydrology is impacted by or contributes to the stream’s water table Deciduous Forests: Primarily comprised of Red Alder, Big-leaf Maple (Acer macrophyllum) and Cottonwood. Large size classes include pre-settlement remnants, stands with less valuable timber, areas in recovery, and sites occupied by rapidly growing species (particularly Cottonwood). Small size designations include areas recovering from repeated logging, regenerating old farm sites or developed areas, and stands clinging to sliding hillsides. Wet sites and wetland margins commonly have a long-term succession of Alder and Cottonwood species. Coniferous Forests: Often dominated by Douglas Fir (Pseudotsuga menziesii), Western Red Cedar (Thuja plicata), and occasionally Western Hemlock (Tsuga spp.). Grand Fir (Abies grandis) and Spruce (Picea spp.) are less common. Introduced species (Redwoods and many horticultural varieties of pine, spruce, fir, cypress, etc.) have grown large and are regenerating in some areas. Significant stands of these immigrants have been included in the assessment. Sizes of coniferous stands range from planted areas full of small trees, to small regeneration stands at the edges of larger stands, and to all forms of second and third growth natural © 2002 Seattle Urban Nature Project. All rights reserved. 4 regeneration. A few areas of very large trees (termed old-growth relicts) remain in Schmitz and Seward Parks. Extensive second growth stands carpet the Llandover area. Broadleaved Evergreen Forests: The Pacific Madrone (Arbutus menziesii) makes pure and nearly pure stands along many ridge and bluff tops with good drainage and sun exposure. Small trees are seen regenerating in these areas and in harsh environments (riprap along I-5, not-so-recently graded areas). Larger size trees in remnant stands dot the city. A dramatic association evident in Lincoln Park and other bluffs is an overstory of Madrone, a midstory of Dogwood (Cornus spp.), and Snowberry (Symphoricarpos alba) in the shrub layer. Mixed Forests: Combinations of Deciduous, Coniferous, and Broadleaved Evergreen Forests. Other: Cliffs: Slumping on steep slopes of glacial till can transport entire habitat areas downhill into Puget Sound or onto underlying structures. What remains are nearly vertical, unvegetated areas of substrate ranging from sand to clay. The cliff/bench/cliff formations in Discovery Park are good examples. Rock and Talus: Generally large erosion and slide control areas. Uncommon. Sensitive Species: The exact locations of five plant species have been masked in the database in the interest of protecting remnant native populations of locally rare herbaceous species. These species were determined to be sensitive without formal criteria. The following species (and associated codes) were found on Seattle’s Public Lands but have not been listed at the individual park or polygon level: Vanilla Leaf Pathfinder Maiden Hair Fern Wild Ginger Trillium ACTR ADBI ADPE ASCA3 TROV © 2002 Seattle Urban Nature Project. All rights reserved. 5 Annotated Glossary: Aerial photograph: Aerial photographs were the primary tool used in the field to assign polygon boundaries. Photos taken in 1995 were provided by Triathlon in Bellevue, WA (formerly NIES Mapping Group, Inc.). The large scale (1:2400, 1”=200’) digital photos clearly show tree canopies, roads, buildings, grassy areas, shrubs and other landscape features. The digital prints were corrected to overcome the “fisheye” distortion effect at the edges of most photos caused when using a circular lens. With this correction, distances could be accurately measured from all locations on the print. Broadleaved evergreen tree: Trees with wide flat leaves (not needles or scales) that lose leaves and grow new leaves slowly all year. The most common example in Seattle is the Madrone (Arbutus menziesii). Some Magnolias (Magnolia spp.) and Live Oaks (Quercus spp.) also fit the definition although they are rare in the public land holdings. Canopy: The upper layer of vegetation. In a forest this includes the leaves and branches of the trees but not the trunk. The canopy provides a protective cover over the forest floor. Canopies come in many forms. Deciduous forest canopies provide summer shade and winter sun (when available) and deep mulch in the fall. They are often wide and round. Coniferous and broadleaved evergreen forest canopies provide year round shade, a sheltered climate, and a slow trickle of needles, leaves, bark, and cones (see Mulch). Canopies can be wide and round as in Madrone, tall and pointy like many conifers, or broken and ragged like a bluff top stand of Douglas Fir. Animals at different times of year use canopies for nesting, feeding, resting, hiding, hunting, or shelter. Many individual plant canopies together can create distinctly different microclimates and soils and attract alternative wildlife populations. For example, a mowed area with trees has a different canopy and habitat potential than one with only grass. While providing habitat for many species, forest canopies will discourage animals that prefer open meadows. In addition, areas located between distinct habitats (e.g. on the edge of a forested area), can provide niches for some animals. Niches can be shade or shelter islands with hunting and hiding perches that provide direct access to open areas, or encompass a rich collection of woodland-dwelling or sun-loving plants. Conifers: Trees or shrubs that have needle- or scale-like leaves and often cones. They lose and grow leaves throughout the year. Puget Sound lowland native species include: Douglas Fir (Pseudotsuga menziesii), Grand Fir (Abies amabalis), Shore Pine (a.k.a. Lodge Pole Pine) (Pinus contorta), Sitka Spruce (Picea sitchensis), Western Hemlock (Tsuga heterophylla), Western Red Cedar (Thuja plicata), and Western White Pine (Pinus monticola). A related group is the Yews (Taxus spp.), which have needles and berries instead of cones. There are also many introduced conifer species growing in the city. © 2002 Seattle Urban Nature Project. All rights reserved. 6 Cover class: One of eight categories which represents the proportion of ground surface included in a polygon covered by a particular species (out of 100%). The categories are: “1” = a trace or single individual of a species, “2” = 0-5% coverage, “3” = 5-10%, “4” = 10-25%, “5” = 25-50%, “6” = 50-75%, “7” = 7595%, and “8” = 95-100%. For example, a field with 100% grass would be classified as an “8”, while an area with 1/3 grass, 1/3 blackberry, and 1/3 pavement would receive a cover class of “5” for each species. An important consideration is the layering of coverages. A grassy area completely shaded by trees will have high cover classes (“7” or “8”) for both the grass and the trees. Data Sheet: The form used in the field to describe polygon characteristics. It contains: park name and code; staff member’s name who described the polygon; date of survey; codes to identify the aerial photograph print used to designate the polygon; polygon number; habitat type; habitat description; special features; the vegetation table; and the wildlife table. Deciduous: Trees or shrubs that lose most of their leaves in the fall and grow a new set of leaves in the spring. Native species include: Big-leaf Maple (Acer macrophyllum), Bitter Cherry (Prunus emarginata), Cottonwood (Populus trichocarpa), Oregon Ash (Fraxinus latifolia), Red Alder (Alnus rubra), and Willows (Salix spp.). There are also many introduced varieties growing in the city. Digitize: An electronic process by which lines that have been physically drawn onto an aerial photograph are converted into digital data for use in a computer database. This is the critical link in the GIS mapping process. By attaching a specific area on the ground to a data field in the computer, other data on the characteristics of the area can be coupled to the specific area. Use of the GIS allows the information to be accessed, displayed, analyzed, and compared to other areas. Digitizing also creates a spatial database that displays the polygons, as they exist on the ground, proportional to other features like roads, bodies of water, etc. and allows polygons to be measured (area, distance between polygons, etc.). This spatial database is the computerized GIS map. Dominant: Describes a tree or other plant species that comprises most of the canopy in a polygon. Habitat Key: A series of questions about the polygon that have two answer choices for each question. The chain of decisions leads to determination of habitat type. These decisions include determining the amount, type and size of forest, grass and shrub areas, the amount of pavement, buildings and other development; and the impact of mowing and other maintenance. This type of key is called a dichotomous key. Habitat Description: A brief summary of features of the polygon that are not captured by the special features or the vegetation table. Possible wetlands, horticultural © 2002 Seattle Urban Nature Project. All rights reserved. 7 varieties, weeding and replanting activity are a few of the many subjects discussed in this part of the data sheet. Habitat types: Fifty-eight categories that reflect the kinds of environments encountered in Seattle. They are listed below and described in the habitat description: Beach and Dune Broadleaf Evergreen (3 size classes) Cliff Conifer Broadleaf Evergreen Mixed (3 size classes) Conifer Deciduous Mixed (5 size classes) Conifer Forest (5 size classes) Deciduous Broadleaf Evergreen Mixed (4 size classes) Deciduous Forest (5 size classes) Developed Pasture Grassland Heavy Development Herbaceous Row Crop Lacustrine Landscaped Forest Landscaped Grassland Landscaped Shrubland Landscaped Tree Savannah Light Development Medium Development Orchard or Vineyard Palustrine Aquatic Bed Palustrine Emergent Wetland Palustrine Forested Wetland Palustrine Open Water Palustrine Scrub/Shrub Wetland Riparian Forest (4 size classes) Riverine Consolidated Substrate Riverine Tidal Riverine Unconsolidated Substrate Rock and Talus Shrub Savannah Shrubland Sparsely Vegetated Habitat Tree Savannah Invasive species: Numerous plant species that are not native to the Seattle area have been intentionally or unintentionally introduced and thrive in this region. These species can be a detriment to the native habitats in the city. Scot’s Broom (Cytisus scoparius ), Himalayan Blackberry (Rubus discolor), English Ivy © 2002 Seattle Urban Nature Project. All rights reserved. 8 (Hedera helix), English Holly (Ilex aquifolium), Cherry Laurel (Prunus laurocerasus), Clematis (Clematis vitalba ), Japanese Knotweed (Polygonum cuspidatum), Morning Glory (Convolvulus sepia), Reed Canary Grass (Phalaris arundinacea), Purple Loosestrife (Lythrum salicaria), and Yellow Flag Iris (Iris pseudacorus) are aggressive plants that are expanding their range in the city. Control in some areas has been very effective. Many areas are not controlled. Impervious surface: Ground cover that does not allow water to penetrate to the soil. Examples include pavement, buildings, plastic sheeting, and very compacted ground. Turf is partially pervious, because it allows a fraction of the water to percolate into the soil. An old growth forest is highly pervious, absorbing rain at multiple canopy layers. Jurisdictional delineations: The Washington State Department of Ecology and US Army Corps of Engineers have established criteria for delineating the edges of wetlands based on the presence of certain soils, vegetation, and water in the soils that indicate saturation for a portion of the growing season. These edges determine buffer areas where building and grading is limited. The wetland edges that appear on the Seattle Urban Nature Project maps and data are not jurisdictional delineations. They are core areas with distinct wetland properties (lots of standing water, many plants that require wet conditions, etc.). Delineating the edges with established criteria would expand the areas of many of these described wetlands. Landscaped: An area that has been regularly mowed, manicured, pruned or recently planted. Mulch: The buildup of leaves, needles, cones, branches, and fallen trees that creates the rich forest soils where woodland plants thrive. Insects, fungi, worms and moles help convert mulch to soil. Each type of tree or animal species gives a different “flavor” or chemistry to the mulch. Palustrine: A fresh water wetland ranging from mid-slope seeps and willow thickets to large open ponds. Polygon: An area on a map or aerial photograph that shows a single patch of a habitat type. Polygons are usually irregular in shape. Habitat polygons were drawn onto aerial photographs in the field and show the boundary of an area with a particular habitat type. Once digitized, polygons contain all of the information that exists about that area (plant and animal species, special features, habitat type, habitat description, etc.). Riparian: Streamside vegetation that is regularly flooded or influenced by the stream’s water table. © 2002 Seattle Urban Nature Project. All rights reserved. 9 Roads and Trails Criteria: There are many types of roads and trails in Seattle. Seattle Urban Nature Project maps show only the regularly driven roads and paved trails over five feet wide. Savannah: Scattered tall woody plants in an area of generally shorter plants (usually grass). Tall plants are trees or shrubs (creating tree or shrub savannah) and short plants are shrubs in a tree savannah or grasses and forbs (in either savannah type). The habitat key and descriptions provide specifics about the canopy cover of trees and shrubs in savannahs. In Seattle, savannahs usually receive some level of management to maintain the dominance of grass species. Size class: One of five tree diameter categories determined by the diameters of the dominant trees in the polygon (< 5”, 5-15”, 15-20”, 20-30”, and > 30”). Spatial database: A collection of polygons, points or features that are arranged according to their position on the ground and in relation to other polygons, roads, water features. Special Features: Features recorded in the field on the data sheet about a polygon including: “snag” (dead tree > 5” in diameter), “down log” (also >5” in diameter), “large tree” (> 35” in diameter), “seep” (a damp or boggy area), “slide” (mud or land slide), “gully erosion” (where water is cutting a ravine), and “refuse” (e.g. garbage, abandoned vehicles, yard waste dumps). © 2002 Seattle Urban Nature Project. All rights reserved. 10