Chapter 50: Biomes Chapter 52: Population Ecology Definitions : Population: Deme: Population density: Dispersion, types of: Life Tables: Study of survivorship at different ages of a population, usually by studying a cohort Cohort: Using life tables: life tables can be used to generate survivorship curves: Type I Type II Type III Fig. 52.3 Reproductive tables: used to measure fertility at different ages, Fig 52.3 Life History Reproductive designs/strategies Big-bang reproduction/semelparity iteroparity Why one design versus the other? Population Growth Measuring Population Growth Change in pop. Size= Births – Deaths ΔN = B-D Δt b = births /year = birth rate B = births in a given year B= bN B= .034 * 500 B= 17 (method to measure expected births) Use the same for deaths… d = death rate ΔN = bN -dN Δt One othe simplification… b-d represents rate of growth…if positive then …..if negative then ΔN = rN Δt This is exponential growth, results in a -shaped curve FIG 52.8, 52.9 Carrying capacity !! ΔN= rmax (K-N) Δt K K= carrying capacity, maximum population size environment can support What happens as population size (N) nears carrying capacity (K)? Fig 52.10, 52.12 K-selection: density dependent selection, maximizes population size, species that are successful at high densities. Ex fruit flies r-selection: density independent, reproductive success greater at low population sizes Density dependent factors Density Independent factors Fig. 52.13, Attaining equilibrium in a population based on birth and death rates….mathematical models Fig 52.13 Case I: density-dependent birth rate and death rates Case II: density dependent birth rate, density independent death rate Case III: density independent birth rate, density dependent death rate Assumptions? Population growth always leads to negative effect, what about too few? Allee effect Do experimental cases match mathematical models? What about response time to effects? Immigration and Emigration? Growth cycles: What causes cyclical population cycles? Boom and bust cycles Human population growth Exponential growth… Noah’s ark? Age structure diagrams: Fig 52.22 Notes Ch. 53 Community Ecology Community: populations in a given area Species richness Relative abundance How are communities built? What factors effect communities? Two theories FIG 53.1 Individualistic hypothesis: populations grow in same area because they need similar abiotic requirements…study populations independently Predicts that each population grows according to its own tolerance to abiotic factors Interactive hypothesis: communities are populations tied together, act as integrated unit Predicts that populations dependent on each other will create clusters Evidence seems to support former in plant communities unless dramatic shift in abiotic factors Other hypotheses? Rivet model: analogy…not all rivets are needed on wing but start taking them out?... Redundancy model: similar to individualistic, species are redundant, if one predator disappears, others take place. Example: wolves in Yellowstone How could this be applied to plants? What might happen if an animal disappears? How could this affect plants? Scriptural? Our job to understand how God designed communities…what is our job? Interspecific interactions and Community structure Competition Interspecific competition Competition for limited resources The Competitive Exclusion principle: SURVIVOR! Outcompete, outsurvive, outresource Gause: if two species compete for the same resource, one will inevitably outcompete What determines competitive exclusion? Use of resources... Ecoligical niche: use of abiotic and biotic factors, the role of an organism in the environment Apply to competitive exclusion principle…two species cannot coexist if their ________ are the same Resource partitioning: two species can coexist if one uses slightly different resources Character displacement: character displacement refers to differences(displacement) in structures or shapes (characters) examples, Galapagos finches that are usually allopatric with similar beaks do show differences in beaks on islands where they are both found. FIG 53.2 and 53.3, 53.4 Also Anole lizards Predation Predators, herbivores, parasites Defenses? Animal defenses: ?? let’s think Passive defenses Active defenses Unusual defenses Chemical Mechanical Adaptive coloration Cryptic coloration Aposematic coloration Mimicry Batesian mimicry Mullerian mimicry FIG 53.7, 53.8 Plant defenses: strychnine, morphine, nicotine, mescaline, tannins, chemicals responsible for cinnamon, cloves, peppermint! Parasites: endoparasites vs ectoparasites Mutualism: Two species that Commensalism: hitchhikers Tropic structure illustrates community dynamics Feeding relationships between populations Quaternary consumers Tertiary consumers Secondary consumers Primary consumers Primary producers (also autotrophs) Multiple Food chains create food web Fig 53.11 Some animals operate at multiple trophic levels: ex’s? Observations show that most food chains average five trophic levels WHAT limits the food chain ? Why only five on average? Two Hypothesis: Energetic hypothesis Only 10% of energy passed from one level to another (Why?) 100kg of producer→10kg for herbivore→1kg for carnivore What does this model predict Approach from mathematical equation, if 100kg increases… How to test for this prediction? Fig 53:13 Dynamic stability hypothesis Food chains are susceptible to disturbances in population numbers, higher levels need to be able to recover from “wobbles”, if the food chain is longer, larger “wobbles”, harder to recover Prediction? How to test? Compare communities that are Dominant species and Keystone species Dominant species have the highest biomass…why dominant? Think back to competition Keystone species are species that have dramatic effect if removed Why? Most are predators that influence other populations How to test? Fig 53: 14 and 15 Studies in Community disturbance Natural weather events Examples: Fig 53: 16,17, 18 Positive or negative? Succesion: Development in communities Primary succession Secondary succession Pioneer species: mainly r-selected species Climax species: mainly K-selected Soil changes as succession occurs