Symonds & Tattersall data set 2: Phylogenies used in the analyses Ramphastidae: The intergeneric phylogeny was that produced by Moyle (2004, his Fig. 4a). Relationships within the genus Ramphastos were taken from Weckstein (2005, his Fig. 4). Relationships within Pteroglossus were taken from Eberhard & Bermingham (2005, their Fig. 4). Relationships within Aulacorhynchus were taken from Puebla-Olivares et al. (2008), with additional information on the position of Aulacorhynchus huallagae in a trichotomy with A. haematopygius and A.caeruleicinctus/A. prasinus taken from Schulenberg & Parker (1997). Relationships within Selenidera are from Cracraft & Prum (1988, Fig. 2). Andigena nigrirostris was placed as sister-group to the other three species in the genus based on the taxonomic account by Sibley & Monroe (1990). Newick format: ((Ramptoco,((Rampambi,Ramptuca),(Rampsulf,((Rampbrev,Rampvite),Rampdico)))),((((((Pter beau,Pterbito),Pterazar),((Pterplur,Ptercast),Pterarac)),(Ptertorq,Pterfran)),((Pterviri,Pterinsc),B ailbail)),(((((Aulapras,Aulacoer),(Aulahual,Aulahaem):0.0001),Aulasulc),Auladerb),((Andinigr ,(Andihypo,(Andicucu,Andilami):0.0001)),(Selespec,(Seleculi,((Selemacu,Selegoul),(Selerein, Selenatt):0.0001))))))); Lybiidae: The intergeneric phylogeny was that produced by Moyle (2004, his Fig. 4a). The phylogeny for the genus Pogoniulus is that of Moyle (2002). All remaining information on intrageneric resolution is taken from taxonomic estimations as provided by Short & Horne (2001). Newick format: (((Tracpurp,Tracvail):0.0001,((Traceryt,Tracmarg),Tracdarn)),((((((Gymnslad,Gymnpeli),Gym ncalv),Gymnbona),(Stacleuc,(Stacanch,Stacwhyt))),(Stacoliv,Buccduch):0.0001),((((((Pogopus i,Pogochry),Pogoscol),Pogoatro),(Pogosubs,Pogobili)),((Pogoleuc,Pogocory),Pogosimp)),(Tric hirs,(((Tricdiad,Tricfron):0.0001,Tricleuc),(((((Lybitorq,Lybiguif),(((Lybichap,Lybileuc),Lybir ubr),Lybiviei)),(Triclacr,Tricmela)),Lybiundu):0.0001,((Lybimela,Lybimino),(Lybibide,(Lybid ubi,Lybiroll))))))))); Psittaciformes: The primary intergeneric phylogeny was that of Wright et al. (2008, their Fig. 2). The position of the ground parrot, Pezoporus wallicus as sister-clade to Neophema was taken from Leeton et al (1994), the position of Lathamus and Psitteuteles from Christidis et al. (1991), and Callicephalon from Brown & Toft (1999). Relationships within the genus Cacatua were taken from Brown & Toft (1999) with additional resolution for C. pastinator, C. tenuirostris and C. sanguinea from Adams et al. (1984). Relationships within Platycercus were from Ovenden et al. (1987), and within Calyptorhynchus we used Saunders (1979). Relationships within Neophema were those used by Burton et al. (2008). Newick format: (((Calybank,(Calylath,((Calyfune,Calylati),Calybaud)):0.0001),((((((((Cacatenu,Cacasang),Cac apast),Cacagale),Cacalead),Cacarose),Callfimb),Probater),Nympholl)),(((((Polyswai,(Polyanth, Polyalex):0.0001),Aproeryt),Alisscap),(Eclerora,Geofgeof)),(((((((((Platcale,Plateleg),((Platexi m,Platadsi),Platvenu)),Platicte),Barnzona),((Psepdiss,(Psepchry,(Psepvari,Psephaem):0.0001):0 .0001),Purpspur)),Northaem),Lathdisc),(((((Neopsple,Neoppulc),Neopeleg),(Neopchrg,(Neopp etr,Neopchrs):0.0001):0.0001),Pezowall),Neopbour)),(((Psitvers,((Glospusi,(Glosconc,Glospor p):0.0001),(Trichaem,Tricchlo)):0.0001),Meloundu),Cycldiop)))); Estrildidae: The intergeneric phylogeny was taken from Sorenson et al. (2004), with resolution of all intrageneric relationships taken from Christidis (1987). Newick format: ((Eryttric,Erytgoul),(Hetepect,((Loncpunc,(Lonccast,Loncflav)),((Staggutt,(Stagbell,Stagocul)), (((Neocmode,Neoctemp),(Neocrufi,Neocphae)),((Poeppers,(Poepcinc,Poepacut)),(Emblpict,(Ta engutt,Taenbich)))))))); Galliformes: Intergeneric relationships taken from the supertree compiled by Kolm et al. (2007, their Fig 1a), with resolution of the genus Lagopus from Lucchini et al. (2001). Newick format: ((Oreopict,(Callcali,Colivirg)),(Melegall,(Bonaumbe,(((Lagomutu,Lagolago),Lagoleuc),(Falcca na,(Centurop,(Dendobsc,(Tympcupi,Tympphas)))))))); Spheniscidae: We used the phylogeny of Ksepka et al (2007, their Fig. 3). Newick format: ((Aptepata,Aptefors),((Pygoadel,(Pygoanta,Pygopapu)),((Eudymino,(Sphemage,Sphedeme)),( Megaanti,(Eudychrc,((Eudychrl,Eudyschl),(Eudyscla,(Eudypach,Eudyrobu)))))))); Laridae: We used the phylogeny of Pons et al. (2005, their Fig. 1). 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