Borrero, F.J. Colombian non-marine mollusks
LIST OF NON-MARINE MOLLUSCA FROM COLOMBIA
The taxonomic order follows…..
Within Genera and Subgenera, species have been arranged alphabetically
A total of XX species are recorded. Whenever available, the maximum number of different
colletion records for each species are included, and these are organized by “Departamentos”
(Colombian political divisions).
Colombia is a megadiverse country, the following table is given as comparison with other
countries.
Reason for megadiversity (discuss and make specific to mollusks):
- Andes split (cordilleras), lots of Andean mountains, valleys, large rivers, lowlands
- 2 oceans
- bridge between Central and South America (Darién/Chocó biogeografico)
- convergence of Andean, Amazon, llanos and Caribbean provinces.
The fauna of San Andrés y Providencia is different from that of the continent. Obviously it can
not be thought of as South American, but more like Caribbean, or Central American. Several
families present in San Andrés are not present in South America (Cerionidae, Polygyridae, etc.).
A
Why this list is valuable.
1-Biodiversity assessment and changes thereof. Conservation.
2-Further guidelines on ecological units within country (snails are less mobile….good indicators).
3- Anyone wishing to work in a particular group will have an account of what is where.
4- assessing/foreseeing effects of dams and large habitat changes
5- parasitology
6- pearl cultivation
7- Selecting alternative species for heliciculture
8- environmental management guidelines
The interest is not only to record all species that have been collected in Colombia, but to
examine/present the maximum extent of each species geographic distribution within the country.
In this manner, information on local extinction may be ascertained, and weighed against records
of habitat change.
This account is likely more complete for mainland Colombia, and less so for the Islands (San
Andres & Providencia, Gorgona, etc.).
There have been the following significant periods of collecting in Colombia: Pilsbry, Bequaert,
Medem, FLMNH/Thompson/hhSmith, B.Malkin (1968-71, mainly Bulimulidae)
Discuss diversity of families.
1- Bulimulidae most diverse. Partly true, probably partly they are very noticeable,
fashionable, and pretty. They are also exciting and people set out to find them. However,
it is noteworthy that Bulimulidae s.s. is not particularly diverse in Colombia in terms of
Borrero, F.J. Colombian non-marine mollusks
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number of subgenera, as compared to other South American countries. Specifically, Perú
and Ecuador are the most diverse (39 and 25 subgenera, respectively), and Colombia (12)
appears similar to Venezuela (12) and Brazil (16) (Breure, 1974).
2- Regarding degree of endemism of subgenera of Bulimulidae s.s., Colombia has no
endemic subgenera, whereas Perú has 25(64% of total), Ecuador has 14 (56%),
Venezuela has 2 (17%), and Brazil has 8 (50%) (see Table 1 in Breure, 1974).
There appears to be a geographic trend in degree of endemism of subgenera, where Costa
Rica (4 subgenera), Panamá (3) and Colombia (12), all have no endemic subgenera, whereas
beginning with Ecuador endemism increases, reaching a maximum in Perú, with the highest
number of subgenera, and of endemism at this level (see Breure, 1974). This is almost
certainly due to the South American origin of Bulimulidae, which migrated towards Central
America, the West Indies and North America later in the Miocene or Pliocene (Breure, 1974).
3- Clausiliidae- less diverse, perhaps partly artificial due to lesser conspicuossness.
[… “Dr. Paul Ehrmann has remarked of the genus Nenia that Ecuador and northern Peru are its
distribution center; the group here reaches its distribution acme of differentiation, and is most
numerous in species” (Pilsbry, 1919)]
4- Families for which revisionary work or monographs are available.
Amazonas
Caldas
Cordoba
Magdalena
Quindío
Tolima
Antioquia
Arauca
Atlántico
Bolívar
Caquetá
Casanare
Cauca
Cesar
Cundinamarca Guanía
Guaviare
Huila
Meta
Nariño
Norte de Santander
Risaralda
San Andrés y Providencia
Santander
Valle del CaucaVaupés
Vichada
Boyacá
Chocó
La Guajira
Putumayo
Sucre
(After Cowie & Thiengo, 3003):
New Granada = present-day Colombia and Panama from 1830 to 1903
= only Colombia from 1903 on
Location of specimens:
ANSP – Academy of Natural Sciences, Philadelphia, PA
Basel ??
BMNH - British Museum Natural History, London
CAS- California Academy of Sciences, San Francisco, CA
CM - Carnegie Museum, Pittsburgh, PA
CMC- Cincinnati Museum Center, Museum of Natural History, Cincinnati, OH
DMNH- Delaware Museum of Natural History, Wilmington, DE
Edinburgh - Royal Scottish Museum, Edinburgh (data in Solem, 1966)
FLMNH – Florida Museum of Natural History, Gainesville, FL
FMC – Field Museum Natural History, Chicago, IL
FML – Fundación Miguel Lillo, Tucumán, Argentina
ICNB- Instituto de Ciencias Naturales, Bogotá, Colombia
IRSN - Institut Royal des Sciences Naturelles, Brussels
Jackson - Ralph Jackson Collection, Cambridge, Maryland (data in Solem, 1966)
MCZ - Malacology Collection, Harvard University, Cambridge, MA
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MLP - Museo de La Plata, La Plata, Argentina
MNHN - Muséum National d’Historie Naturelle, Paris
NMWZ – National Museum of Wales-Zoology
NRS - Naturhistoriska Riksmuseet, Stockholm
OSUM- Ohio State University Museum of Diversity, Columbus, OH
RMNH - Rijksmuseum van Natuurlijke Historie, Leiden
SMF - Natur-Museum Senckenberg, Frankfurt am Main
UMNG-MT – Universidad Militar Nueva Granada, Santafé de Bogotá, Colombia
USNM – US National Museum Natural History- Smithsonian Inst., Washington D.C.
UVZ – Universidad del Valle-Cali, Colombia (Zoology)
ZMA, ZMA/K - Instituut voor Taxonomische Zoölogie (Zoölogisch Museum), Amsterdam
K (Kuiper)
ZMH - Zoologisches Museum der Universität Hamburg
ZMHB- Museum für Naturkunde der Humbold-Universität, Berlin
ZMUZ - Zoologisches Museum, Universität Zürich, Zürich (data in Solem, 1966)
CLASS GASTROPODA
SUBCLASS PROSOBRACHIA
ORDER ARCHAEOGASTROPODA
Family Ceresidae
Archecharax glaeserius Thompson, 1980
Valle del Cauca: [Holotype FLMNH 24355, Coll.:F.G.Thompson/1969, 3 km W Atoncelo, 1380
m (Thompson, 1980)]
Family Neritidae
Neritina latissima Broderip, 1833 [Nerita latissima Broderip]
[“Columbia” ANSP 37344]
Neritina spp. Caribbean
Theodoxus meleagris (Lamarck, 1822)
Magdalena?: [Don Diego River, 0.5 mi. above river mouth, DMNH 132237]
Theodoxus luteofasciatus
?: [Pacific] DATA ???
Family Helicinidae
Alcadia spectabilis (Poey, 1846)
Magdalena: [Sierra Nevada de Santa Marta, 1800m, Coll.K.-J.Götting 1976-78 (Götting, 1978)]
Helicina appuni Martens, 1873
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Magdalena: [(H. cf. appuni Martens): Sierra Nevada de Santa Marta, Ecoandes Transect Alto
Buritaca, 1100m, Coll. T.van der Hammen et al./Aug.1977, RMNH, ICNB (Breure, 1977b)]
?Helicina bicincta Gloyne, 1872 [Colombia ?]
“Central America; upper Amazon region” (Simone, 2006) [thus Colombia?]
Helicina cacaguelita Pilsbry & Clapp, 1902
Magdalena: [Lectotype ANSP 83164, Sierra Nevada de Santa Marta, Cacaguelito
(=Cacaguelito), 1500 ft., forest ground under decaying leaves, Coll.:H.H.Smith/1898]
Helicina colombiae E.A.Smith, 1878
?: [San Sebastian, New Granada (Baker, 1923)]
Helicina colombiana Philippi, 1847 [RELATIONSHIP WITH Helicina colombiae E.A.Smith,
1878???]
Magdalena: [(H. cf. colombiana Philippi): Sierra Nevada de Santa Marta, Ecoandes Transect
Alto Buritaca, 700-900m, Coll. T.van der Hammen et al./Aug.1977, RMNH, ICNB (Breure,
1977b)]
?: [“...from Colombia...” (Baker, 1923)]
Helicina (Helicina) concentrica Pfeiffer, 1848. [Breure, 1977b]
Venezuela, Brazil, Peru, Colombia (Simone, 2006)
Valle del Cauca: (Restrepo et al., 1983): [Paso de la Bolsa, UVM 82-153/1] [Hacienda El
Castillo, Jamundí, UVZ 089/2, 090/3, 091/2, 093/4, 094/18] [Vivero El Vínculo, Buga, UVZ 82239/16] [Jardín Botánico Mata de Guadua UVZ 82-307/1] [La Paila, UVZ 103/9, 104/12, 105/10]
[Hacienda Formosa, Ansermanuevo, UVZ 82-039/22, 82-218/1, 82-052/1]
Cundinamarca: [(H. c. pandiensis Wagner), Pandi (Baker, 1923]
?: [“New Grenada”, Swift Collection, ANSP 14602]
Helicina cucutensis ( Pilsbry, 1939)
Norte de Santander: [Lectotype ANSP 168432, Cúcuta, 07°55’N-072°31’W, Coll.: Brother
Nicéforo María (Pilsbry, 1939); Paralectotype ANSP 358659]
Helicina fasciata providentiae Pilsbry, 1930
San Andrés y Providencia: [Providencia Isld., Coll.: Pinchot Expedition, April 1929: Holotype
ANSP 150717, Valley North of High Peak (12°21’N-081°20’W); Paratypes 150151, 150161,
top of ridge N of High Peak; Paratype ANSP 150169, west slope below cleft of Split Hill]
[Providencia Isld., Coll.: K.Emberton/1987: W slope of N half of Split Hill, ca. 100 ft. below
base of main cliff (13°30’N-081°30’W), on cockspur acacia, ANSP A12126H; 50 ft. below short
ridge between Twin Pinnacles of High Peak, on N side, ANSP 368422, ANSP A12127G]
Helicina heighwayana Dall, 1909
Chocó: [Holotype USNM 111074: “Near the Atrato River, in the Sierra Darién, Coll.: A.E.
Heighway (Dall, 1909). Solem (1966) considers this location as part of modern Panamá] [Acandí:
FLMNH 95289; Coll.: F.Keep, FLMNH 1600151]
Borrero, F.J. Colombian non-marine mollusks
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Helicina (Helicina) microdina huberi (Breure, 1976)
Norte de Santander: [33 km S. Cúcuta, selva veranera, 750 m, Coll. Breure 2May/75, RMNH
55121 (Breure, 1976b)]
Helicina nemoralis ? ????
Magdalena: [Aracataca, Coll.Rehn & Hebard/1920, ANSP 46579]
Helicina ocanensis (A.J. Wagner, 1905)
Norte de Santander: [Ocaña (Baker, 1923)]
Helicina rawsoni Pfeiffer
San Andrés y Providencia: [San Andrés Isld., Airport, Coll.:D.G.Robinson/1982, FLMNH 49010]
[Providencia Isld., conch farm, Coll.: D.Addison/1991, FLMNH 194097]
Helicina riparia Pfeiffer, 1854
Bolívar: [“Calaman”, New Granada (=Calamar) (“...from near Calamar, Colombia” in Baker,
1923), Swift Collection, ANSP 14612]
Antioquia: [Aracataca, Col.: Rehn and Hebard, 1920; ANSP 46579 (Baker, 1923)]
Helicina rhynchostoma Shuttleworth in Pfeiffer, 1865
Chocó: [”Near the Atrato River, in the Sierra Darién, Coll.: A.E. Heighway (Dall, 1909). Solem
(1966) considers this location as part of modern Panamá]
?: [as H. r. rhynchostoma “Shuttleworth” Pfeiffer (1865), “Campanera” (Baker, 1923)]
Helicina sanctaemarthae Pilsbry & Clapp, 1902
Magdalena: [Lectotype ANSP 83165, Paralectotype ANSP 358660, Santa Marta Mountains at
El Líbano, 6000 ft. (10°15 N- 074°14 W), on trees, specially palm leaves, Coll.:H.H. Smith/1898]
[Sierra Nevada de Santa Marta, 50 km SE Sta.Marta, just W of Río Buritaca along trail to Ciudad
Perdida, 650 m elev., Coll.:D.K.Dorman/1989, FLMNH 137922; 550 m., near trail beside Río
Buritaca,1989, FLMNH 137927]
Helicina (Helicina) sanctaemartae Breure, 1977 [(WHAT’S THE RELATIONSHIP OF THIS
SPECIES AND H. sanctaemarthae Pilsbry & Clapp, 1902???)]
Magdalena: [Holotype ICNB, Sierra Nevada de Santa Marta, watershed between Río Frío and
Quebrada del Páramo, 2900m, Coll.A.M.Cleef et al./Aug.1977; Idem, Filo Buritaca, 2300m
(RMNH 55312/1) (Breure, 1977b)]
Helicina steindachneri Wagner, 1905
Valle del Cauca: [Cordillera Occidental, locations along the road Cali-Buenaventura (Restrepo &
Breure, 1987)]
Antioquia?: [Frontino (Baker, 1923)]
Helicina sylvatica Orbigny, 1837
?: [mentioned by Haas (1948) in comparison to H. weyrauchi Haas, 1948 from Peru (Orbigny,
1837 in op.cit.]
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Helicina superstructa Wagner, 1905
Antioquia?: [Frontino (Baker, 1923)]
Helicina (Helicina) unizonata Haas, 1966
Putumayo: [Holotype FMC 114098, Región of Nueva Granada, Upper Río Putumayo,
downstream from Puerto Asís, 76°27’W-0°24N, Coll.:J.M.Idrobo, July 1957]
Helicina sp.1
San Andrés y Providencia: [Providencia Isld, in hills near center, Coll.: C.R.Gilbert et al./1971,
FLMNH 217529]
Helicina sp.2
Magdalena: [Coll.: H.H.Smith, El Líbano, near Santa Marta, FLMNH 95352]
Helicina sp.3
Magdalena: [Cacaguelita near Santa Marta, 2000 ft, Coll.: H.H.Smith/1899, FLMNH 95356]
Helicina sp.4
Cesar: [Chiriguana, Coll.: Rosenberg, FLMNH 237584]
Lucidella lirata (Pfeiffer, 1847) [? Colombia. Distribution includes Panama and Venezuela
(Martens, 1890-1901 in Pérez & López, 2003)]
ORDER MESOGASTROPODA
Family Cyclophoridae
Systematic arrangement follows Bartsch & Morrison (1942).
Subfamily Amphicyclotinae
Buckleyia bifasciata Mousson
Antioquia: [Jericó, Coll. H.Damel (=Hno. Daniel), USNM 414156]
Filocyclus delphinulus (Mousson) (? Colombia)
?: [“The shells described by Mousson were collected by Gustav Wallis in northen South America.
He collected in parts of Colombia, Ecuador, and the Amazon region...” (Bartsch & Morrison,
1942)]
Calaperostoma hidalgoi (Crosse)
?: [“New Grenada (Colombia)”, Coll.Cuming, USNM 316134]
Calaperostoma leai Bartsch & Morrison, 1942
Nariño: [“Jumaco” (=Tumaco), western Colombia, Coll.Cuming, USNM 525747]
Calaperostoma cumingi (Sowerby)
Nariño: [Island of Tumaco, USNM 307420]
Calaperostoma sp.
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Cauca (??): [“Pereiza” (???), FLMNH 165437]
Calacyclotus atratensis, Bartsch & Morrison, 1942
Chocó: [Type USNM 206291, “Mountains near the mouth of the Atrato River in Colombia”,
Coll. A.E.Heighway]
Calacyclotus olssoni (Pilsbry, 1926)
Chocó: [Acandí, boundary between Panamá and Colombia, Coll.A.E.Heighway, USNM 341767]
Subfamily Aperostominae
Aperostoma (Incidostoma) incomptum (Sowerby)
?: [Type USNM 515940, no locality data, from the J.B.Henderson collection (Bartsch &
Morrison, 1942)] [(Neocyclotus incomptus Sowerby), “Colombia”, CMC C11066, C11067]
Aperostoma (Incidostoma) kobelti Bartsch & Morrison, 1942
Cundinamarca: [“Lake of Tedropalo, between Bojac (=Boyacá) and Tene(?)”, USNM 37582]
Aperostoma (Incidostoma) nirafe Bartsch & Morrison, 1942
Norte de Santander: [Type USNM 380795, “dense jungle, 13 km S of Puerta Santos,
Coll.R.H.Palmer]
Aperostoma (Incidostoma) pergrandis (Kobelt)
?: [“New Granada (Colombia)”. It is possible that this is taxon is actually based on an inmature
sspecimen of Aperostoma (Incidostoma) malleatum Bartsch & Morrison, 1942 (Bartsch &
Morrison, 1942)]
Incidostoma sp.1
Valle del Cauca: [Atuncelo, 9 km. N Dagua, 2800 ft., Coll.: F.G.Thompson/1969, FLMNH
252309; 3 km.W Atuncelo, 4500 ft., FLMNH 258060]
Incidostoma sp.2
Cundinamarca: [Bogotá, FLMNH 164711]
?: [Manipos, FLMNH 164710]
Aperostoma (Austrocyclotus) limellum Bartsch & Morrison, 1942
Magdalena: [Type USNM 515906, Rio Cesar Valley, Coll. O.Haught]
Aperostoma (Austrocyclotus) peilei (Gude, 1912)
Antioquia: [Type (in British Museum), Alejandría, 50 miles from Medellín, 5600 ft, Coll.:
T.P.Sharman (Gude, 1912; Bartsch & Morrison, 1942)]
Aperostoma (Neocyclotus) corpulentum (E.A.Smith, 1878)
Magdalena: [San Andrés, Rio Frío Basin, Sierra Nevada de Santa Marta, 1200 mt., Coll.
H.Pittier, USNM 192951] [Santa Marta Mountains at El Líbano, 6500 ft elevation, under
decaying laves in forest]
?: [“Colombia”, FLMNH 243781]
Borrero, F.J. Colombian non-marine mollusks
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Aperostoma (Neocyclotus) dysoni Fischer & Crosse, 1886 (?Colombia: Bartsch & Morrison,
1942 cite 13 subspecies of A.(N.) dysoni, none present or reported from Colombia)
?: [cited from Colombia by Martens, 1890-1901( Perez & López, 1993)]
Aperostoma (Neocyclotus) sanctamarthae Pilsbry & Clapp, 1902
Magdalena: [Type CM 62.15065, “Las Nubes Estate, Santa Marta Mountains”, 4000 ft., Coll.:
Mr.&Mrs.H.H.Smith, 1898 (Pilsbry & Clapp,1902; Clapp, 1920; Bartsch & Morrison, 1942)]
Aperostoma (Neocyclotus) smithi Pilsbry & Clapp, 1902
Magdalena: [Lectotype ANSP 83166, Santa Marta Mountains at El Líbano, 6500 ft., 04° 55’N075° 02’W, Coll.:H.H.Smith/1898. Under decaying leaves in forest (Pilsbry & Clapp,1902]
Aperostoma (Neocyclotus) wetmorei Bartsch & Morrison, 1942
Magdalena: [Type USNM 536033, Tierra Nueva, Sierra Negros, Coll.A.Wetmore &
M.R.Carriker, Jr.] [San Lorenzo, collado between the radio-telegraph stations, approx. 2600m,
Coll. Coll.K.-J.Götting 1976-78 (Götting, 1978)]
Neocyclotus sp.
Magdalena(?)/La Guajira(?): [Sierra Nevada de Santa Marta, Coll.:D.K.Dorman/1973, FLMNH
22263]
Aperostoma (Aperostoma) bogotense (Pfeiffer)
Cundinamarca: [Bogotá, USNM 316133] [Neocyclotus bogotense (Pfeiffer, 1855), Bogotá,
FLMNH 129965]
?: [(Aperostoma bogatensis Pfeiffer), “New Grenada”, ANSP 12950]
Aperostoma (Aperostoma) caucaense (Da Costa, 1901) (Bartsch & Morrison, 1942 state:
“...looks as if its peculiarities were due to an injury” [thus may not be a true species])
Cauca: [“Province of Cauca” (Da Costa, 1901, in Bartsch & Morrison, 1942)]
Aperostoma (Aperostoma) cingulatum (Sowerby, 1843)
Cauca?: [“Cauca”: USNM 307422; FLMNH 243780]
Antioquia: [Jerico: USNM 414158; USNM 424727, Coll.H.Daniel]
Cundinamarca: [Bogotá, FLMNH 129966] [Neocyclotus cf. cingulatus, El Totumo, 3.5 km NE
Tocaima, Coll.Correal & van der Hammen, 1992, fossil specimens (Correal & van der Hammen,
2003)]
Chocó(?)/Risaralda(?): [“Pereira Cauca”(????), FLMNH 129967]
Huila: [Neiva, FLMNH 129968]
?: [“Colombia”, USNM 307419] [“South America”, USNM 307457]
Aperostoma (Aperostoma) columbiense (Da Costa)
Cundinamarca: [Near Bogotá (Baker, 1923)]
?: [“New Grenada (Colombia)”, Coll.Sowerby and Fulton, USNM 307432]
Aperostoma (Aperostoma) confusum (Sykes)
Chocó: [Acandí, border between Colombia and Panamá, Coll.A.E.Heighway, USNM 341766]
Antioquia: [“...occurs elsewhere at Antioquia, Colombia (Pilsbry, 1926)]
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Aperostoma (Aperostoma) cumingi Bartsch & Morrison, 1942
?: [Type USNM 307416, “Colombia”, Coll.Cuming]
Aperostoma (Aperostoma) dunkeri (Pfeiffer, 1856)
Valle del Cauca: [Rio Paila Valley, 1800mt, Coll.H.Pittier, USNM 251170]
[“Colombia”, FLMNH 243784]
Aperostoma (Aperostoma) filioliratum (Sowerby) [spelling? filoliratum ¿?]
Cundinamarca: [Bogotá, Museum Da Costa (Bartsch & Morrison, 1942)]
Aperostoma (Aperostoma) fischeri (Hidalgo, 1867)
?: [“Colombia”, FLMNH 243790] [Neocyclotus fisheri, Hidalgo, “Columbia” (Webb, 1948)]
[“New Grenada” USNM 307424]
Aperostoma (Aperostoma) giganteum (Reeve, 1842)
Chocó: [Neocyclotus giganteum (Reeve, 1842), Acandí, FLMNH 129971] [Aperostoma gigantea
Gray, On the Atrato River slope of Sierra Darién at about 1000ft, Coll.: A.E. Heighway, ANSP
98287 (Dall, 1909; Pilsbry, 1926). Solem (1966) considers this location as part of modern
Panamá]
Caldas: [Marmato (Pilsbry, 1926)]
Cauca: [(Neocyclotus giganteum Gray):Isla de Tumaco, W.Colombia, CMC C????]
Valle del Cauca: [Incidostoma cf. giganteum (Reeve, 1842), Cordillera Occidental, locations
along the road Cali-Buenaventura (Restrepo & Breure, 1987)]
Aperostoma (Aperostoma) inca (Orbigny, 1835)
From Simone (2006): Colombia, Bolivia, Brazil, Paraguay, Peru, Argentina. A. colombiensis
(Ferussac in Orbigny, 1835)(non DaCosta, 1901) is a synonym.
[Syntype BMNH 1854.12.4.287/7; 34 mm (Simone, 2006)]
Aperostoma (Aperostoma) laxatum (Sowerby, 1849)
Cundinamarca: [Neocyclotus laxatus Sowerby, Fusagasugá, Coll.M.L.Jaume, March 15, 1937,
CMC C6746??]
?: [“Columbia. Mus.Cuming” (Bartsch & Morrison, 1942)]
Aperostoma (Aperostoma) paezense Bartsch & Morrison, 1942
Huila: [Type USNM 251169, Valle of the río Paez, in the Central Cordilleras, Coll.H.Pittier]
Aperostoma (Aperostoma) paezicolum Bartsch & Morrison, 1942
Huila: [Type USNM 524047, Rio Paez Valley, 2500mt., Coll.H.Pittier]
Aperostoma (Aperostoma) pailaense Bartsch & Morrison, 1942
Valle del Cauca: [Rio Paila Valley, 1300mt, Coll.H.Pittier, USNM 251171]
Aperostoma (Aperostoma) perezi (Hidalgo, 1866)
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Valle del Cauca: [Neocyclotus cf.perezi (Hidalgo, 1866), Loboguerrero, offered for internet sale:
Firama, 2004]
Aperostoma (Aperostoma) popayanum (Lea, 1839)
Cauca: [Syntype USNM 104453X, near Popayán, Coll.:Gibbon]
Caldas?: [Marmato, Coll.Bland, USNM 307406]
Valle del Cauca: (Restrepo et al., 1983): Paso de la Bolsa UVZ 82-155/9, 82-156/5; Vivero “El
Vínculo”, Buga, UVZ 82-264/8; Jardín Botánico Mata de Guadua, Tuluá, UVZ 133/5, 81-229/2;
La Paila, UVZ 125/3, 126/3; Hacienda “Formosa”, Ansermanuevo, UVZ 82-053/5, 82-038/15]
Santander: [Socorro, FLMNH 164706]
?: [(as Cyclotus papayanus (Lea, 1836), “Colombia”, DMNH 177862]
Aperostoma (Aperostoma) simile Bartsch & Morrison, 1942
Cundinamarca?: [Type USNM 535989, “Bogotá, Coll.Sowerby & Fulton, USNM 307398, same
locality]
?: [New Grenada, USNM 307400]
Aperostoma (Aperostoma) umbilicatum Bartsch & Morrison, 1942
Cundinamarca: [Type USNM 307428, Bogotá]
Aperostoma (Aperostoma) utriaense Bartsch & Morrison, 1942
Chocó: [Type USNM 524068, Puerto Utría]
Aperostoma (Cyclohidalgoa) belli belli (Beddome, 1908)
Only known from one record, with uncertain locality.
Antioquia?: [(Beddome, 1908; Bartsch & Morrison, 1942): Near Zaragoza at 800 ft in dense
forest, Coll.: E.Bell] [It could also be from the towns called by the same name in Cauca or Valle
del Cauca]
Aperostoma (Cyclohidalgoa) belli haughti Bartsch & Morrison, 1942
Santander: [Type USNM 524136, and Paratypes USNM 472798, south part of Santander,
Coll.O.Haught]
Norte de Santander: [Poteria translucida translucida (Sowerby, 1843): “Cucutá” (=Cúcuta),
ANSP 12963 (Baker, 1923)]
Aperostoma (Cyclohidalgoa) translucidum translucidum (Sowerby, 1843)
Casanare: [Neocyclotus translucidum translucidum (Sowerby, 1843)(N. translucidus (Sowerby,
1843), Abbott, 1989) Orocué, FLMNH 164708]
?: [“New Grenada”, USNM 307424, 307405, 307408]
Incerticyclus, a pseudogeneric term (Since knowledge of the operculum is required to ascertain
Cycloporids’ generic and subgeneric name, Incerticyclus has been erected as a “superspecific
designation for species that has been christened but that in [their] monographic endeavors [they]
are unable to locate properly for lack of the characters mentioned. ... It is not to be considered a
genus, nor is it ever to have a type, but it is to remain an open catch-all” (Bartsch & de la Torre,
1942)).
Borrero, F.J. Colombian non-marine mollusks
11
Incerticyclus distinctus (Sowerby)
?: [Bay of Montija, West Colombia, Coll.H.Cuming]
Poteria niceforana (Pilsbry, 1939) (The genus Poteria is .... “a nude name......he lumps here
about everything known from Jamaica at that time [1840-1850] except what had been segregated
to form....Cyclotus lineatus” (Bartsh & de la Torre, 1942)). These authors list several species of
Poteria in 2 subgenera, all restricted to Jamaica. Further, no P. nicerorana is listed in their
monograph. The actual generic affiliation of “Poteria niceforana” is thus uncertain, as is the
actual species it may represent).
Norte de Santander: [Holotype ANSP 168431, Cúcuta, 07°55’N-072°31W, Coll.: Brother
Nicéforo María]
Family Ampullariidae
Systematic arrangement follows Reeve (1856), Alderson (1925), and Cowie & Thiengo (2003).
Asolene crassa (Swainson, 1823)
Distribution: Guyana, French Guyana, Surinam, Venezuela, Colombia, Ecuador, Brazil (Simone,
2006)
Magdalena: [Ampullarius monticolus (Vernhout, 1914), a synonym of crassa Swainson, 1823
(Refs. in Cowie & Thiengo, 2003)] [(as A. monticolus Vernhout): beach of Isla de Salamanca,
and in fast flowing streams ending in Sevilla, Toribío and Córdoba Rivers in NE area of Ciénaga
(Cosel, 1986]
Atlántico: [(as A. monticolus Vernhout): Laguna de Luruaco (carretera Cartagena-Barranquilla
(Cosel, 1986)]
Asolene platae (Maton, 1809)
Distribution: Bolivia, Colombia, Brazil, Argentina, Paraguay, Uruguay (Simone, 2006)
Asolene sp.
?: [Colombia, 34mm up (Femorale Ltda, 2001/2003)]
Marisa cornuarietis (Linnaeus, 1758).
Distribution: Cuba, Costa Rica, Panama, Trinidad, Tobago, Venezuela, Colombia, Guyana,
French Guyana, Surinam, Bolivia, Brazil Paraguay, Argentina, Uruguay (Simone, 2006)
La Guajira: [Rio Hacha: 11 15N 074 13W, USNM 153452; 11°34N 072°58W, ANSP 322236]
Magdalena: [Ponds and flooded areas in Isla de Salamanca and Isla de los Gómez, as well as
rivers ending in Cienaga Grande de Santa Marta (Cosel, 1986)]
Atlántico: [“Barranquilla, Magdalena River marshes”, Coll.: H.W.Kale/1977, FLMNH 243515]
[Coll.:T.Link, USNM 364420] [Río Magdalena, 1 km.SE Sabana Grande, Coll.: Thomerson et
al./1973, FLMNH 24787]
Bolívar: [Ciénaga of Rio Magdalena, 10 km. E Cartagena, Coll.: C.R.Gilbert/1969, FLMNH
39828] [Cartagena, CMC C11094-11097] [Coll.: S.Smith, USNM, 314986; Coll.:T.Link, USNM
364416] [ponds and lakes between Cartagena and Barranquilla (Bandel, 1976 in Cosel, 1986)]
Córdoba/Sucre: [Between Tolú and Coveñas; Los Volcanes, Sinú; Laguna de Momil, E of Lorica;
Caño Viejo, Pivijay (Pain, 1956)]
Borrero, F.J. Colombian non-marine mollusks
12
Marisa rotula (Mousson, 1869) [May be a synonym of M. cornuarietis (Linnaeus)]
Atlántico: [Marsh, Sabanagrande, Coll.: F.G.Thompson/1990, FLMNH 174358] [Laguna
Totumo at Volcán Totumo, Coll.: F.G.Thompson/1990, FLMNH 174360]
?: [“Lower Magdalena River” (Cowie & Thiengo, 2003)]
Marisa sp.
Atlántico: [Cartegena (=Cartagena), MCZ 316340]
?Pomacea (subgenus?) amazonica Reeve, 1856 [? Colombia]
“Amazon River” (Simone 2006).
?Pomacea (subgenus?) armeniacum Hupé, 1857 [? Colombia]
“Amazon River” (Simone 2006).
?Pomacea (subgenus?) castelnaudii Hupé, 1857 [? Colombia]
“Amazon River” (Simone 2006).
?Pomacea (subgenus?) lineata (Spix, 1827)
Distribution: Jamaica, Colombia, Venezuela, Ecuador, Guayana, Brazil, Argentina,
Uruguay (Simone 2006).
Pomacea (Effusa) aurostoma (Lea, 1856)
Distribution: Colombia, Venezuela (Simone, 2006)
?: [Syntypes USNM 106273, “Colombia”,Coll.: J.Totten, Ident.by I. Lea,1856; Syntype USNM
106299X] [“New Grenada”, Ampullaria aurostomus Lea, Coll.R.Swift, ANSP 50693]
Atlántico: [Luruaco (Laguna Luruaco) (Pain, 1956)]
Norte de Santander: [Cucutá (=Cúcuta), Wheatley Collection, ANSP 124619 (Baker, 1930; Pain,
1956)]
Pomacea (Efffusa) baeri (Dautzenberg, 1902)
?: [Distribution including Colombia (Refs. in Cowie & Thiengo, 2003)]
Pomacea (Effusa) castelloi (Sowerby, 1894)
Colombia, Surinam (Simone, 2006)
Meta?: [Lectotype BMNH 1893.5.29.3, Rio Meta, S.E. Bogotá]
?: [Colombia, MZSP 1841 (Simone, 2006)]
Pomacea (Effusa) chemnitzii (Philippi, 1852)
Magdalena: [La Cueva, Santa Marta (Pain, 1956)] [synomym P. porphyrostoma (Reeve, 1856)
(Cowie & Thiengo, 2003): Rivers ending in Ciénaga Grande de Santa Marta, and lakes and
ponds in Isla de Salamanca and between Barranquilla (Atlántico) and Santa Marta; Isla de Los
Gómez (Cosel, 1986)]
Bolívar: [(as P. porphyrostoma Reeve) Cartagena, DMNH 187256]
Córdoba/Sucre: [(as Ampullarius porphyrostomus Reeve) Rivers and lakes and ponds along
Magdalena River between Tolú and Coveñas, and Laguna de Momil, E of Lorica (Pain, 1956)]
Borrero, F.J. Colombian non-marine mollusks
13
?: [“Colombia”, ANSP 120419 (Baker, 1930)] [Distribution including Colombia (Refs. in Cowie
& Thiengo, 2003)]
Pomacea (Effusa) columbiensis (Philippi, 1851)]
??: [Syntypes ZMHB 1343, “Westkolumbien” (probably “Rio Pastása in Andibus orientalibus,
Colombia”) (Refs. in Cowie & Thiengo, 2003)] [“Columbia”, ANSP 120419]
Cundinamarca: [P. columbiensis (Reeve), Bogotá, FLMNH 87517]
Pomacea (Effusa) columellaris (Gould, 1848) [(?Colombia). This species’ distribution is
restricted to Perú (Cowie & Thiengo, 2003; Ghesquiere, 2003)]
Chocó: [Limnopoma columellaris (Gould, 1848), Bahía de Solano, floor of dense forest above
beach [(however, this is a freshwater snail!], Coll.:D.R.Paulson/1961, FLMNH 20469]
Pomacea (Effusa) cornucopia (Reeve, 1856)
?: [Syntype BMNH 20020651, “Columbia” (Refs. in Cowie & Thiengo, 2003)]
Pomacea (Effusa) cumingii (King & Broderip, 1831) [Subspecific taxon Ampullaria cumingi
urabaensis Pain, 1856 is present in Colombia, although, the stated distribution of cumingii King
& Broderip does not include Colombia (Cowie & Thiengo, 2003)]
Antioquia: [Holotype and 3 Paratypes NMWZ 1981.118.00114, Golfo de Uraba; additional
paratypes at MCZ]
Chocó: [Camp La Nueva, Rio Truandó, Coll. F.J. Barcroft, ANSP 195769 (Pilsbry, 1956)] [Rio
Atrato, Coll. J.Brouwer (Pain, 1956)]
Pomacea (Effusa) fasciata (Roissy, 1805) [?Colombia: The distribution of fasciata Roissy, 1805
is Jamaica only, but a synonym, buxea Reeve, 1856 is reported from Colombia]
??: [Pomacea (Effusa) buxea (Reeve, 1856), distribution including Colombia (Refs. in Cowie &
Thiengo, 2003)]
Pomacea (Effusa) flagellata (Say, 1829)
Bolívar: [Ampullaria flagellata Say, “Carthagena”, CMC C11516, C11517, C11530]
[“Colombia” DMNH 187244; “Northern Colombia (Magdalena drainage area)” (Cowie &
Thiengo, 2003; Ghesquiere, 2003)]
Pomacea (Limnopomus) forbespoma [Not listed in Cowie & Thiengo (2003)]
?: [Colombia 06°01 N-077°22 W, USNM 515453]
Pomacea (Effusa) glauca (Linnaeus, 1758) [Subspecies minuscula Baker, 1930 and planorbula
Philippi, 1851 reported from Colombia (Cowie & Thiengo, 2003)]
Distribution: Trinidad, Barbados, Guadeloupe, Dominica, Martinique, Saint Lucia, Bolivia,
Colombia, Venezuela, Guyana, Surinam, French Guyana, Brazil (Simone, 2006)
Meta: [Coll.: F.Medem/1974, USM 853420; Coll.: S.Tello/1974, USM 853421] [Rio Guatiquia,
Caños Negros, 18 km. on road from Villavicencio, Coll.: C.Gilbert et al./1973, FLMNH 39827]
Atlántico: [Pomacea (Efffusa) oligista (Pilsbry & Olsson, 1953) is considered a synonym of
planorbula Philippi, 1852, which is considered a subspecies of glauca Linnaeus , 1758 (Cowie &
Thiengo, 2003)]. Holotype ANSP 189546, 10 spec.dry, Ciénaga de Luruaco, Road Cartagena-
Borrero, F.J. Colombian non-marine mollusks
14
Barranquilla, 11° 00’N-074° 48’W, Coll.:A.A.Olsson/ March 1952; Paratypes ANSP 18547,
365367, USNM 605847] [Lago Luruaco, Coll.: F.G.Thompson/1990, FLMNH 174356]
?: [“Cuenta, New Granada”, Wheatley Collection, ANSP 120420 (Baker, 1930)]
[“Colombia”(?)(Baker, 1930): Coll.Schaufuss, ANSP 50610; ANSP 50601] [“Northern South
America (Atlantic drainage)” (Morrison, 1946)] [Ampullaria glanca Linn, “Colombia”, ANSP
50610] [(Ampullaria prunulum Reeve 1856 (=P.glauca planorbula Philippi, 1852), “Colombia
or Panama”, Syntypes BMNH 20020679 (Cowie & Thiengo, 2003)]
Pomacea (Effusa) guaduasensis (Anderson, 1928)
Cundinamarca: [Holotype CAS 2721, San Juán de Río Seco, on the eat border of the upper
valley of the Magdalena River]
Pomacea (Effusa) haustrum (Reeve, 1856) (Not mentioned as present in Colombia by Cowie &
Thiengo, 2003)
Valle del Cauca: [From small pond 1 km.S of Jamundí (15 km.S of Cali), Coll.: G.Kattan/1990,
FLMNH 190627]
Pomacea (Effusa) hollingsworthi (Pain, 1946)
Colombia (Simone, 2006)
Cundinamarca: [Holotype BMNH 1946.6.24.24, 64mm; Paratype NMWZ.1981.118.00198, in a
swiftly flowing stream with a rocky bed near Bogotá]
Pomacea (Effusa) novaegranadae (Busch, 1859)
??: [Syntypes BMNH 20020671, “New Granada (in 1859=present-day Colombia and Panama)”
(Refs. in Cowie & Thiengo, 2003)]
Pomacea (Effusa) ocanensis (Kobelt, 1914)
Norte de Santander: [Ocaña in Neu-Granada (=Colombia), ZMUH (Cowie & Thiengo, 2003)]
Pomacea (Effusa) palmeri (Marshall, 1930)
Distribution: Colombia (Simone, 2006)
Santander del Norte: [Type USNM 380696, small stream in dense jungle, 13 km S of Puerto
Santos, Coll.:R.H. Palmer; Paratypes: USNM 380697 (55mm); 380698 (embryonic shells),
same location, Coll.:R.Palmer (Marshall, 1930)]
Santander: [Quebrada Gómez railway, Puerto Wilches, Coll. J. Brouwer (Pain, 1956)]
Pomacea (Effusa) patula (Reeve, 1856)
?: [Syntypes BMNH 20020673, distribution includes New Granada (=Colombia and
Panama)(Cowie & Thiengo, 2003)]
Pomacea (Effusa) pealiana Lea, 1838
Distribution: Colombia.
Bolívar: [Lectotype ANSP 192933 (35mm), Turbaco, 10°20’N-075°25 W, Coll.: T.R.Peale,
1944; Paralectotypes MCZ 161600]
Antioquia: [Turbo region, Golfo de Urabá, Coll. J.Brouwer (Pain, 1956)]
Borrero, F.J. Colombian non-marine mollusks
15
Pomacea (Effusa) producta (Reeve, 1856)
Distribution: Amazon River (Simone, 2006)
?: [Syntypes BMNH 20020678, “F.Magdalen [=Colombia], Neu Granada (Colombia or
Panama)(Refs. in Cowie & Thiengo, 2003)]
Meta: [Region of Villavicencio, Coll.V.Guevara, pre-1935 (Pilsbry, 1935)]
Pomacea (Effusa) semitecta (Mousson, 1873)
Distribution: Colombia, Venezuela (Simone, 2006)
?: [Holotype ZMUZ 283 (50mm)] [(Ampullaria semitecta Mousson), cited as a Colombian
species by Haas (1951) when describing records from Lambadero, Zulia, Venezuela]
[Distribution includes Colombia (Refs.in Cowie & Thiengo, 3003)]
Santander/Boyaca: [Anca de Caballo, Rio Ermitaño (tributary of Magdalena River), Magdalena
Valley (Pain, 1956)]
Pomacea (Effusa) superba (Marshall, 1926)
Distribution: Colombia
Atlántico: [(A. superbus Marshall), Holotype USNM 362863 (45mm), Cienaga Totuma, Coll.
T.A.Link (Marshall, 1926)] [Coll. J. Brouwer (Pain, 1956): Laguna de Luruaco; Puerto Caimán]
Bolívar: [Coll. J. Brouwer (Pain, 1956): Los Volcanes, Sinú; Laguna de Momil, E. of Lorica;
Road between Sincerín and San Cayetano; Arroyo Mancomoján, headwaters between El Carmen
& Ovejas]
Casanare: [Coll.: S.Tello/1974, USNM 853422]
Pomacea (Effusa) urceus (Müller, 1774)] Ampullaria urceus Müller, 1774
Distribution: USA, Mexico, Trinidad, Venezuela, Guyana, French Guyana, Colombia, Peru,
Brazil (Simone, 2006).
Antioquia: [Puerto Berrío, Río Magdalena. Coll. R.W.Barker (Pain, 1956)]
Santander: [as Pomacea urceus guyanensis (Lamarck, 1819): Carare, region, south of
Barrancabermeja, Magdalena Valley, Coll. J. Brouwer (Pain, 1956)]
Arauca: [Rio Arauca at City Arauguita, Coll.: C.R.Gilbert et al./1970, FLMNH 22110, 22111]
?: [“Columbia”, ANSP 50665] [Distribution includes Colombia (Cowie & Thiengo, 2003;
Ghesquiere, 2003)]
Pomacea (Effusa) zeteki Morrison, 1946 (The distribution given in Cowie & Thiengo (2003)
restricts it to Panama)
Bolívar: [09°17N-075°08W, Coll.: Daniel/1933, USNM 444122]
[“Colombia”: Coll.:T.Link, USNM 364416; Coll.:A. Heighway, USNM 206286, 206287]
Pomacea (Effusa) zonata (Spix in Wagner, 1827)
Distribution: Colombia, Brazil (Simone, 2006)
?: [Syntypes, ZSM (Simone, 2006)] [Distribution includes Colombia (Refs. in Cowie & Thiengo,
2003)]
Pomacea sp.1
Amazonas: [Leticia-ditch N Tarpon Zoo, Coll.: R.M.Shelley/1977, FLMNH 28000]
Pomacea sp.2
Borrero, F.J. Colombian non-marine mollusks
16
Magdalena: [2 km.W of Caracoti, 100m., Coll.:F.G.Thompson/1990, FLMNH 174357]
Pomacea sp.3.
Atlántico: [Marsh, Sabanagrande, Coll.: F.G.Thompson/1990, FLMNH 174359,174361] [Laguna
Totumo at Volcán Totumo,Coll.: F.G.Thompson/1990, FLMNH 174362,174363]
Family Annulariidae
Choanocoma (Choanopomops) barkeri (Haas, 1951)
La Guajira: [Coll.: O.Haught/1943, USNM 599653]
Magdalena: [Coll.:O. Haught/1944, USNM 5434469]
Choanopoma (Choanopomops) barkeri perijensis Solem, 1960
Magdalena: [Paratype USNM 473939, Ident. G.Solem/1960; Paratype USNM 473929,
Coll.:O.Haught]
Torrella simpsoni simpsoni Henderson & Bartsch
Atlántico: [Soledad Cave in central portion, Coll.:1933, FLMNH 119058]
Chondropoma hjalmarsoni Pfeiffer
San Andrés y Providencia: [Providencia Isld, Airport, Coll.:D.G.Robinson/1982, FLMNH 49009]
Parachondria pyrostoma Pilsbry, 1930
San Andrés y Providencia: [Holotype ANSP 150931, and Paratype ANSP 150932, San Andrés
Island, North Cliff, 12°30’N-081°40’W, Coll.: Pilsbry & Pinchot/1929] [San Andrés Isld.: Coll.:
D.R.Paulson et al./1966, FLMNH 218203] [(Annularia pyrostoma (Pilsbry, 1930b) San Andrés
Islnd., In moist eroded pocket on soil and dead leaves, at base of limestone cliff on north end of
island, NE face, 12°30’N 081°60’W, Coll.: K.Emberton/1987, ANSP 368417 A12126D]
Family Hydrobiidae
Andesipyrgus sketi Hershler & Velkovrh, 1993
Santander: [Holotype USNM 860574, Coll.:B.Sket/1984, La Paz River, 06°11’N-073°35’00 W;
Paratype USNM 860575]
Aroapyrgus colombiensis Malek & Little, 1971
Valle del Cauca: [Type? ANSP 321818. (Mikkelsen & Bradford, 1997: Holotype DMNH
041542; 91 Paratypes DMNH 041543), Tributaries of Río Pichindé, near Pichindé, Municipio
de Cali, Coll.E.A. Malek & M.D.Little, 1968. Host of Paragonimus caliensis and P. emberai,
n.sp. ]
Aroapyrgus sp.”a”
Magdalena: [Sierra Nevada de Santa Marta: Donama, Coll.: K.J.Götting/1976, FLMNH 223516;
San Pedro de la Sierra, Coll.: K.J.Götting/1976, FLMNH 223518]
Aroapyrgus sp.”b”
Magdalena: [Rio Guachaca, E of Santa Marta, Coll.: K.J.Götting/1976, FLMNH 223517]
Borrero, F.J. Colombian non-marine mollusks
17
Pyrgophorus coronatus (Pfeiffer, 1840)
?: [Distribution includes Colombia (Hershlers & Thompson, 1992 in Pérez & López, 2003)]
Pyrgophorus parvulus (Guilding, 1828)
Magdalena: [Ciénaga Grande de Santa Marta, Station near 166, Coll.:R.von Cosel/1970,
FLMNH 224038; All over Ciénaga Grande de Santa Marta, including Isla de Salamanca and Isla
de los Gómez, in ponds and lakes (Cosel, 1986)] [Buritaca River (Götting, K., in Cosel (1986)]
Pyrgophorus sp.
Magdalena: [Ciénaga Grande de Santa Marta: Río de Grapo, 1 m elev., Coll.:R.von Cosel/1971,
FLMNH 93950, 225085; Coll.:R.von Cosel/1971, FLMNH 93952, 93970; Near center, 10°50’N74°23’W, Coll.:D.K.Dorman/1989, FLMNH 137911, 137912]
Lithococcus venustus Pilsbry, 1950.
Nariño: [Type: ANSP 186145, Paratypes ANSP 182273, Between Boca de Napi and Suárez, on
high rock cliffs, and good current, Coll. A.A.Olsson, Dec.19-20/1935 (Pilsbry, 1950)]
Cicinatia sp. (Roldán, 2001)
Potamopyrgus sp. (Roldán, 2001)
Lyrodus sp. [“…widespread from southern United States (Texas) to Central America , the West
Indies, and south to the Argentine in Latin America.” (Morrison, 1946).
Family Pleuroceridae
? Doryssa asperata Lamarck 1822 [? Colombia]
South America (Simone, 2006)
Doryssa sp.
?: [in river swamp, “2 mi. N Lebeic” (??), Coll.: E.R.Allen/1957, FLMNH 18564]
Family Thiaridae
?Aylacostoma strigillata (Dunker, 1843) [?Colombia]
Orinoco River (Simone, 2006)
Melanoides (Melanoides) tuberculata (Müller, 1774) [Introduced European species]
Valle del Cauca ??: [Club Campestre Farallones de Cali, Municipio de Jamundi (?), in small
creeks, Rio Pance Drainage, Coll. F.J.Borrero/June 2004, CMC C
]
?: [(Introduced species, Roldán, 2001)] [Velásquez et al., in press (2004?)]
Thiara granifera (Lamarck, 1822)
Valle del Cauca ??: [Club Campestre Farallones de Cali, Municipio de Jamundi (?), in small
creeks, Rio Pance Drainage, Coll. F.J.Borrero/June 2004, CMC C
]
Borrero, F.J. Colombian non-marine mollusks
18
Pachychilus laevissimus (Sowerby, 1824)
[“Colombia”, FLMNH 87181]
Pachychilus sp. (Roldán, 2001)
Hemisinus binneyi Tryon
?: [New Granada, 2 specs, dry, ANSP 26786]
SUBCLASS PULMONATA
ORDER BASOMMATOPHORA
Family Ellobiidae
See Gibson-Smith & Gibson-Smith, 1982 for Ellobium, Melampus, Detracia.
Detracia globulus (Orbigny, ??)
Nariño: [Tumacao Island (Tumaco) (Morrison, 1943)]
Detracia sp.? Atlantic Coast ?.
Ellobium stagnalis Orbigny, 1853
Nariño: [Tumaco Island, Coll.Cuming, USNM 119545 (Morrison, 1943)]
Melampus coffeus (Linne, 1758)
Magdalena: [Various locations within Ciénaga Grande de Santa Marta, including Isla de
Salamanca and Isla de los Gómez, and Ciénaga de Cuatro Bocas, associated to muddy bottoms
and mangrove roots (Cosel, 1986)]
Bolívar: [Cartagena area (Cosel, 1986): Channel going into El Laguito of El Rodadero;
Cartagena; Isla Grande in Islas del Rosario]
Melampus piriformis (Petit)
Nariño: [Tumaco Island (Morrison, 1943)]
Tralia sp.? Atlantic Coast?
Family Lymnaeidae
Lymnaea (Pseudosuccinea) columella (Say, 1817)
Distribution: Central and South America, west of Andes (Simone, 2006)
[Malek & Cogswell/1980 : Depts. Valle del Cauca, Cauca, Meta, Nariño. L.bogotensis and L.
peregrina are synonymns]
Cundinamarca: [(L. bogotensis Pilsbry, 1935) Type ANSP 164566, Bogotá, Coll.Hno.Apolinar
María, pre-1935 (Pilsbry, 1935)]
Lymnaea cousinni Jousseaume, 1887 [Programa de Biodiversidad Chile]
Borrero, F.J. Colombian non-marine mollusks
19
Lymnaea selli (Preston)
Cundinamarca: [Usaqueu (=Usaquén), Coll.R.P.E.Rochereau, pre-1935 (Pilsbry, 1935:
“described from Bogotá”)]
Lymnaea truncatula ?? (María Imelda, Univ.de Antioquia Project, 2003)
Family Physidae
Aplexa (Stenophysa) rivalis (Maton & Rackett, 1807) [Introduced Euopean species]
San Andrés y Providencia: [Old Providence (=Providencia Isld.), in debris on the beach south of
Isabel, Coll.Pinchot Expedition, April 1929 (Pilsbry, 1930b)]
Aplexa (Stenophysa) marmorata (Guilding, 1828)
Distribution: Central and South Americas (Simone, 2006)
Magdalena: [(as Physa marmorata Guilding, 1828, from Cosel, 1986): in aquatic plants at mouth
of Sevilla and Fundación Rivers, Ciénaga Grande Santa Marta; more common upstream. Rivers
Buritaca, Don Diego, Palomino and other rivers East of Santa Marta, as well as other rivers and
streams in Sierra Nevada de Santa Marta]
[“Colombia”, USNM 103236]
Physella venustula (Gould)
Valle del Cauca: [3 km.W of Loboguerrero, Rio Dagua, 457 m, Coll.: F.G.Thompson/1969,
FLMNH 219026]
Physella sp.
Cundinamarca: [31.1 km.NW Meta Dept.Line, Quebrada Chirajara, Coll.:C.R.Gilbert/1973,
FLMNH 92820]
Family Planorbidae
B. canonica (Uribe, 1950) [Velazquez et al., 2002]
Biomphalaria amazonica (Paraense, 1966)
Distribution: Amazon Basin (Simone, 2006)
Amazonas: [Leticia (Velázques et al., 2002)]
Biomphalaria glabrata (Say, 1818) [? Colombia]
Distribution: Central and South Americas (Simone, 2006)
Biomphalaria kuhniana (Clessin, 1883) (Dejong et al., 2001)
Distribution: Colombia, Surinam, Brazil (Simone, 2006)
Antioquia: [Segovia; Porce, Llanogrande (Velázques et al., 2002)]
Meta: [Acacías (Velázques et al., 2002)]
[These authors find that all the above Colombian species (straminea, peregrina, canonica &
oligoza) are actually B. amazonica and B. kuhniana]
Borrero, F.J. Colombian non-marine mollusks
20
B. oligoza (var. B. philippiana) (Uribe, 1950)
Distribution: Only cited from Brazilian locatlities (Simone, 2006)
?: Reported by Velazquez et al. (2002)
B. peregrina (Malek, 1985)
Distribution: South America (Simone, 2006)
?: Reported by Velazquez et al. (2002)
Biomphalaria prona (Martens)
Magdalena: [E of Santa Marta, Río Guachaca, Coll.: K.J.Götting/1976, FLMNH 93948, 93949]
Biomphalaria straminea (Dunker, 1848) (Barbosa, 1968)
Distribution: Central America, Guyana, Venezuela, Brazil (Simone, 2006)
[Velazquez et al., 2002]
?Biomphalaria teganophila teganophila (Orbigny, 1835) [? Colombia]
Distribution: South America (Simone, 2006)
Biomphalaria sp.
Magdalena: [Ciénaga Grande, near Center, 10°50’N-74°23’W, Coll.:D.K.Dorman,1989,
FLMNH 137913]
Biomphalaria sp.
Magdalena: [(From Cosel, 1986): Río Sevilla and other rivers that empty into the Ciénaga
Grande de Santa Marta; Ciénaga Grande proper; rivers and streams East of Santa Marta]
Bolívar: [streams and canals near Cartagena (Cosel, 1986)]
Biomphalaria sp.
Valle del Cauca: [3 km W of Cisneros,1500 ft., Coll.:F.G.Thompson/1969, FLMNH 270870]
Gyraulus sp.
Magdalena: [Río Fundación, 1 km upstream from mouth at Ciénaga Grande,
Coll.:D.K.Dorman/1989, FLMNH 137931]
Drepanotrema cultratum (Orbigny, 1845) [Given as synomym of D. depressissimum (Moricand,
1839) by Simone (2006)]
Distribution: Cuba, Venezuela, Bolivia, Brazil, Peru, Paraguay, Argentina, Uruguay (Somone,
2006)
Magdalena: [“...from a pond in the desert at Bolivar, Santa Marta...” (Baker, 1930)]
?Drepanotrema cimex (Moricand, 1839) [? Colombia]
Distribution: Central and South Americas (Simone, 2006)
?Drepanotrema lucidum (Pfeiffer, 1839) [? Colombia]
Borrero, F.J. Colombian non-marine mollusks
21
Distribution: Central and South Americas (Simone, 2006)
Drepanotrema sp.
Valle del Cauca: [Carmelo (El Carmelo?), 1000 ft., Coll.: F.G.Thompson/1969, FLMNH 270852]
Helisoma canonicum (Cousin) (?Colombia)
Cundinamarca: [Bogotá, Coll.pre-1935 (Pilsbry, 1935: “The specimens agree well with the
account of this Ecuadorian species”)]
Helisoma sp. (Roldán, 2001)
Planorbula (Tropicorbis) pallida (Adams, 1846)
Magdalena: [“...in a pond in the desert at Bolívar, Santa Marta, Colombia”(?),
Coll.Univ.Michigan Expedition, Aug/1920 (Baker, 1930: “The jaw and radula...are very similar
to...P.kuhniana” (DeJong)]
ORDER STYLOMATOPHORA
Family Veronicellidae
Sarasinula dubia (Semper, 1885) [?Colombia]
Distribution: Mexico, Central America, Venezuela, Brazil (Thomé, 1989; Simone, 2006) [Thus,
very likely in Colombia]
Sarasinula plebeia (Fischer, 1868)
Distribution: Pantropical; USA, Mexico Central America, Antilles, Colombia, Venezuela, Brazil,
Chile (Simone, 1006)
Atlántico: [Barranquilla, in gardens, Coll.K.-J.Götting, 1976-78 (Götting, 1978)]
Veronicella laevis Blainville
San Andrés y Providencia: [(V. laevis laevis Blainville), Old Providence (=Providencia Isld.),
Coll.Pinchot Expedition, April 1929 (Pilsbry, 1930b] [Providencia Isld, hill nr.center of isld,
from trees, Coll.: C.R.Gilbert et al./1971, FLMNH 255567]
Veronicella sloanei (Cuvier, 1817)
San Andrés y Providencia: [Isla de Providencia, Coll. 1829, ANSP A1078 (Voucher for Thome’s
Neotropical Veronicellidae study)]
Veronicella sp.1
San Andrés y Providencia: [Providencia Isld., Coll.: K.Ebel/1998, FLMNH 271154]
Veronicella sp.2
[“Colombia”, Site/Stat.No.54-5171, Coll.:1954, USNM 57940]
Vaginula minuta (Srth., 1913) (Shuttleworth?)
Antioquia: [“...from near Angelopolis, Colombia...” (Baker, 1927)]
Borrero, F.J. Colombian non-marine mollusks
22
Vaginulus occidentalis (Guilding, 1823) [Host of Angiostrongylys costaricensis Morera &
Cespedes, 1971 in Colombia (Malek, 1981)]
Magdalena: [Veronicella occidentalis (Guilding, 1825), Aracataca, Coll.:1920, ANSP A1086]
[“Colombia”, Station 53-12849, Coll.:1943, USNM 574919]
Forcatulus coerulescens (Semper, 1885)
Distribution: Brazil, Venezuela, Colombia (Simone, 2006)
Family Philomycidae
Philomycus sp.
[“Colombia”, Site/Stat.No.67-2264, Coll.:1967, USNM 769140]
Family Pupillidae
Pupoides albilabris nitidulus (Pfeiffer, 1839)
La Guajira: [1-2 km S of RioHacha, Coll.: P.W.Hummelinck/1937; FMC 65311, 65313; NE of
RioHacha, FMC 65312]
Family Valloniidae
Circinaria ponsonbyi Pilsbry & Clapp, 1902
Magdalena: [“Santa Marta Mountains” at Cacaguelito,1500 ft., under dead leaves,
Coll.:Mr.&Mrs.H.H.Smith, 1898 (Pilsbry & Clapp, 1902; Clapp, 1920)]
Circinaria ponsonbyi var clara Pilsbry & Clapp, 1902
Magdalena: [“Santa Marta Mountains” at El Líbano, 6000 ft.,Coll.: Mr.&Mrs.H.H.Smith, 1898
(Pilsbry & Clapp; 1902, Clapp, 1920)]
Pupisoma dioscoricola (Adams, 1845) [Introduced Central American Species ?]
La Guajira: [2 km S of RioHacha, Coll.: P.W.Hummelinck/1937, FMC 296624]
??: [Considering its extensive distribution in North, Central, and South America, this species is
likely present in many locations in Colombia]
Pupisoma macneili? (Clapp, 1928)
La Guajira: [2 km S of RioHacha, Coll.: P.W.Hummelinck/1937, FMC 296623]
Pupisoma sp.
San Andrés y Providencia: [Providencia Islnd, 50-100ft below short ridge between Two
Pinnacles of High Peak, 13°30’N-081°30W, In wet plametto leaves in high ferms, Coll.
K.Emberton/ 20 Oct.1987, ANSP A12127I]
Family Vertiginidae
Gastrocopta barbadensis barbadensis (Pfeiffer, 1853)
La Guajira: [NE of RioHacha, Coll.: P.E.Hummelinck/1937, FMC 65277; 1-2 km S of RioHacha,
FMC 65278-280]
Borrero, F.J. Colombian non-marine mollusks
23
Gastrocopta contracta (Say)
San Andrés y Providencia: [Old Providence (=Providence Isld.), wood on the ridge north of High
Peak, Coll.Pinchot Expedition, April 1929 (Pilsbry, 1930b] [Providencia Isld.: 50-100 ft., below
short ridge between Twin Pinnacles of High Peak, North Side, in wet palmetto litter in high ferns,
Coll.:K.Emberton/1987, ANSP 368430; Next to small dry stream bed above path, below Spanish
Saddle on Trail from bottom house to High Peak, under leaf litter, ANSP 368431] [St.Andrew
Isld. (=San Andrés Isld., Coll.: Pinchot Expedition, April, 1929 (Pilsbry, 1930b]
Gastrocopta colombiana (Pilsbry, 1921)
Atlántico: [Caribbean coastal region: Puerto Colombia (Pilsbry, 1921)
La Guajira: [RioHacha area: Near RioHacha, Coll.:Thompson & Al López/1966, FLMNH 65396,
65273, 65394; 1-2 km S of RioHacha, Coll.: P.W.Hummelinck/1937, FMC 65273, 65394; NE of
RioHacha, FMC 65393]
Gastrocopta geminidens (Pilsbry, 1917) [?Colombia. Distribution includes Costa Rica and
Venezuela (Zilch, 1959; Refs. in Pérez & López, 2003)]
Gastrocopta pellucida (Pfeiffer, 1841)
San Andrés y Providencia: [Providencia Isld.: Conch farm, Coll.: D.Addiso/1991, FLMNH
194096; Hillside at W end of Manzanille (= Manchioneal) Bay, S End Of Island, from near Sea
Level to the top Of The Ridge, 13°30’N-081°30’W, Virginn forest, leaf litter (primarily fronds
from smooth-stemmed palmetto) and rocks, very wet from recent rains, Coll.K.Emberton,
24Oct1987, ANSP 368439] [St.Andrew Isld. (=San Andrés Isld., Coll.: Pinchot Expedition,
April, 1929 (Pilsbry, 1930b]
?: [Considering its extensive distribution in North, Central (Costa Rica, Panamá), and South
America (Ecuador) (Martens, 1890-1901 in Pérez & López, 1993), this species is likely present
in the Colombian mainland]
Gastrocopta servilis (Gould, 1843)
San Andrés y Providencia: [Old Providence (=Providence Isld.), wood on the ridge north of High
Peak, Coll.Pinchot Expedition, April 1929 (Pilsbry, 1930b] [Providencia Isld.,
Coll.:K.Emberton/1987: 50 ft., below short ridge between Twin Pinnacles of High Peak, North
Side, in wet palmetto litter in high ferns, ANSP A12126M; Coconut Grove between beach and
wooded hillside, E end of Manzanille (=Manchioneal) Bay, S end of island, under driftwood logs
after rains, ANSP 368435, A12128L] [Providencia Isld.: conch farm, Coll.: D.Addison/1991,
FLMNH 194095] [St.Andrew Isld. (=San Andrés Isld., Coll.: Pinchot Expedition, April, 1929
(Pilsbry, 1930b]
??: [Distribution includes Panamá and Venezuela (Refs. in Pérez & López, 2003), thus it may be
in Colombia’s mainland]
Vertigo sp.
San Andrés y Providencia: [Providencia Isld., Coll.: K.Emberton/1987: 50-100 ft. below short
ridge between Twin Pinnacles of High Peak, on N side (13°30’N 081°30’W), in wet palmetto
litter in high ferns, ANSP 368429 A12127E; next to small dry stream bed above path, several
Borrero, F.J. Colombian non-marine mollusks
24
hundred ft. below Spanish Saddle on trail from bottom house to High Peak, under leaf litter
beside rocks, 45° angle, ANSPA 12128A]
Family Strobilopsidae
Strobilops piratica Pilsbry, 1930
San Andrés y Providencia: [Type ANSP 150860, Paratypes ANSP 150862, Old Providence
(=Providence Isld.), on the summit of the ridge north of high peak, in debris of forest, Coll.
Pinchot Expedition, April 1929 (Pilsbry, 1930b]
Strobilops sp.
San Andrés y Providencia: [Providencia Isld., Coll.: K.Emberton/1987: ca. 50 ft. below short
ridge between Twin Pinnacles of High Peak, on N side (13°30’N-081°30’W), in palmetto litter,
high ferns, ANSP 368424; on coconut grove between beach and wooded hillside, E end of
Manzanille (=Manchioneal) Bay, S end of island, under drift logs, ANSP A12128F]
Family Clausiliidae
Alopia cyclostoma cyclostoma (Bielz, 1858) (=Clausilia cyclostoma Pfeiffer, 1849)
Cauca: [Popayán, NMWZ PU4027]
Nenia (Andinia) barcrofti Pilsbry, 1957
Cundinamarca/Meta?: [Type and 2 Paratypes ANSP 203475, Monteredondo, km 73 from
Bogotá to Villavicencio, Coll.: F.Medem/1956 (Pilsbry, 1957)]
Gonionenia dohrni Pfeiffer, 1860
Norte de Santander?: [As Nenia Dohrn Pfeiffer, 1860: Syntype SMF 62 027, Ocana (=Ocaña ?)
(Zilch, 1960, as Gonionenia dohrni (L.Pfeiffer)]
[“Colombia”, ANSP 105072] [=Clausilia dohrni Pfeiffer, 1860, “Colombia”, NMWZ PU4106,
PU6328]
Nenia epistomium (Kuster in Martini & Chemnitz, 1847)
Cundinamarca?: [“Bogota”, DMNH 152656]
Nenia karsteniana magistra Sowerby
Cundinamarca: [Santa Fé de Bogotá, FLMNH 268819] [=N.karstesiana (Dohrn, 1859),
Monteredondo, Coll.:1956; FMC 114119; Selva húmeda fría, Coll.: P.I.Scheible/1960, FMC
114120] [Nenia magistra Sabana de Bogotá, on coffee trees, NMWZ PU 4879; Bogotá, Coll.:
McAndrew, NMWZ PU6306; N. (Neniops) magistra, Santa Fé de Bogota, SMF 62 055 (Zilch,
1960)] [Pilsbry (1957) reported Nenia magistra and N. karsteniana as distinct species, both from
near Bogotá]
?: [“Colombia”, FLMNH 268821]
Nenia kusteniana (Dohrn, xxxx)
Cundinamarca: [Sasaima, DMNH 161399]
Boyacá: [Muzo, DMNH 152655]
Borrero, F.J. Colombian non-marine mollusks
25
Nenia perarata (von Martens, 1873)
Antioquia?: [Frontina (=Frontino?), Nueva Granada, FMC 521; Frontino (SMF 62 037) (Zilch,
1960)]
?: [“Colombia”, FLMNH 210219] [“Colombia” altitude 8000 ft, NMWZ PU6307]
Nenia smithiae Pilsbry, 1901
Magdalena: [Sierra Nevada de Santa Marta at El Líbana (=El Líbano), 6000-7000 ft. in moss on
tree trunks, Coll.: H.H.Smith/1898. Pilsbry (1901) mentioned “Types in ANSP and of George H.
Clapp”] [Paratypes FLMNH 49822, Near Santa Marta, El Líbano, 6000 ft., Coll.:H.H.Smith]
[Socorpa Mission, Sierra de Perijá, Coll.: B.Malkin/1968, FMC 168529-533] [San Lorenzo,
approx. 2600m, Coll., K.-J.Götting, 1976-78 (Götting, 1978)]
Clausilia stomium (Küster)
Cundinamarca: [Villeta, Bogotá, NMWZ PU4202, PU4203] [Alto del Frigo near Sabana de
Bogotá, NMWZ PU6384]
[“Colombia”, NMWZ PU6330]
Clausilia pseudoepistomium (Bourguignat) (same as C. epistomium (Küster) ?)
[“United States of Colombia”, NMWZ PU5401]
Columbinia columbiana (Polinski, 1924)
Valle del Cauca ?: [Rio Aguaca-Tal, Columbia (=Rio Aguacatal?), SMF 30 012 (Zilch, 1960)]
[Cordillera Occidental, locations along the road Cali-Buenaventura (Restrepo & Breure, 1987)]
[(=?)Nenia columbiana (Polinski, 1924), Syntype NMWZ PU3908, Aguacatal, 2000 ft., West
Cordillera]
Nariño: [Santiago, Coll.: E.I.Schlinger & E.S.Ross/1955, FMC 106233]
Family Orthalicidae
Subfamily Orthalicinae
Orthalicus adamsoni maculatus Pilsbry, 1899
Cundinamarca: [Lectotype ANSP 78569, Bogotá (Breure & Schouten, 1985)]
Orthalicus bensoni (Reeve, 1849)
Distribution: Guyana; Amazon River; Peru? (Simone, 2006). As Oxystyla bensoni: “Colombia”,
1 ½ inches (Webb, 1948).
Putumayo: [Santa Rosa de Sucumbios, Río San Miguel, Coll.: B.Malkin, FMC 174915]
[“Colombia” FLMNH 202995]
Orthalicus bifulguratus (Reeve, 1849)
?: [Lectotype BMNH 1975583, “Andes of Columbia” (Breure & Scouten, 1985)]
Orthalicus fulvescens Pfeiffer, 1856
La Guajira: [Type Locality: Río de la Hacha, Colombia (Parodiz, 1962)]
Borrero, F.J. Colombian non-marine mollusks
26
Orthalicus jamaicensis Pilsbry
Bolívar: [Puerto Colombia, Coll.:Lowe, FLMNH 158866]
Orthalicus maracaibensis Pfeiffer, 1856
Atlántico: [Edge of Laguna de Guájaro, Coll.: G.R.Dolmatoff/1956, FMC 114088] [Barranquilla,
DMNH 152701]
Bolívar: [Lower Río Magdalena, Río Viejo, Coll.: F.Medem/1956, FMC 57025, 114069;
Hacienda “Jesús del Río, Loma Santa, Munic.Zambrano, Coll.:G.R.Dolmatoff, FMC 114109]
[Sincelejo, Coll.: F.Medem/1956, FMC 57026; Coll.: G.Dahl/1956, FMC 114068, 114071]]
[Magangué Dique, Río Cauca S of Magdalena junction, Coll.: L.H.Roa, FMC 109734]
[Tolúviejo, Río Pechelín, Coll.: F.Medem et al./1964, FMC 187553, 187563-187564, 187579]
[Gambote, Canal del Dique, Road Cartagena-Sincelejo, Coll.: L.A.Cepeda/1965, FMC 187565]
[Tierra Bomba Isld. In front of Cartagena, Coll.: L.Cepeda/1963, FMC 187578]
Cesar: [foothills of Sierra de Perijá, 10-12 km E of Becerril, Coll.: B.Malkin/1969, FMC 172999,
173006]
La Guajira: [Between Cabo de la Vela and Village El Cabo, Coll.: F.Medem/1965, FMC 187561]
[Piojo, Coll.: H.LeNesour/1964, FMC 187580] [Urumita and Rio Hacha (Pilsbry, 1889]
Magdalena: [Bonda region: 8 mi SE of Santa Marta, FMC 61063, 100746; Coll.: G.Clapp/1902,
FMC 84175, 84176; Shore of Quebrada Matogiro, Coll.:F.Medem & C.C.Velásquez/1964, FMC
187562] [Santa Marta, Río Manzanares, Coll.: H.N.Howland/1902, FMC 13589] [Near Alto Las
Minas, Road Fundación-Valledupar, Coll.:R.W.Barker, FMC 109719] [Pozos Colorados, ~11
km W of Santa Marta, Coll.: B.Malkin/1969-70, FMC 173066, 187567] [area of Parque Nacional
Natural Tayrona: road to Villa Concha, Coll.: C.A.Velásquez & L.A.Cepeda/1964, FMC
187612; Bahia de Cinto, Coll.K.-J.Götting, 1976-78 (Götting, 1978); Bahia de Neguange,
Coll.K.-J.Götting, 1976-78 (Götting, 1978)]
Santander: [Malagavita, Coll.: R.E.Acosta/1960, FMC 114070]
Valle del Cauca: [Palmaseca, Lower Río Bolo, between Cali & Palmira, Coll.: J.C.Cubillos/1965,
FMC 187566]
Orthalicus maracaiboensis maracaiboensis (Pfeiffer)
Atlántico: [Barranquilla, Coll.: H.Dybas, 1938, FMC 17512, 17513]
Magdalena: [Orthalicus maracaibensis maracaibensis (Pfeiffer), Santa Marta, Coll.:H.H.Smith,
FLMNH 109542]
Orthalicus maracaibensis imitator (Pilsbry)
Atlántico: [Puerto Colombia, FLMNH 109544] [Barranquilla (Breure & Schouten, 1985)]
Bolívar: [Puerto Colombia, Coll.:Lowe, FLMNH 158867]
Magdalena: [Santa Marta, Coll.: H.H.Smith, FLMNH 109545]
Orthalicus obductus (Shuttleworth, 1856)
Magdalena: [Aracataca, Coll.: Rehn & Heberd/1920, FMC 78726]
?: [“Colombia”, 53mm and up (Femorale, 20040]
Orthalicus ponderosus (Strebel, 1882)
?: [“Finea Bonilla, Candeloria” (=Finca Bonilla, Candelaria), Coll. 14April/1965, DMNH
067277 (could be Antioquia, Valle del Cauca, Huila, Sucre, Atlántico, Magdalena, Cesar,
Boyacá, or Casanare)]
Orthalicus princeps fischeri (Martens, 1893)
Borrero, F.J. Colombian non-marine mollusks
27
Distribution: (As Orthalicus fischeri Martens, 1983) Guatemala, Peru? (Simone, 2006).
Sucre: [Sincelejo, near Colegio Bolívar, Coll.: F.Medem/1956, FMC 115806]
Orthalicus pulchellus (Spix, 1827)
Distribution: Venezuela, Colombia, Bolivia, Brazil Paraguay (Simone, 2006).
[Type, ZSM, 45mm (Simone, 2006)].
Orthalicus tricinctus Martens, 1893
[“Colombia” (Breure & Schouten, 1985)]
Sultana (Melaniorthalicus) adamsoni (Gray, 1834) [=?? Porphyrobaphe adamsoni Gray]
Caldas: [Quebrada Gallinazo, trib.of Río Chinchiná, 6 km from Manizales, Coll.: J.M.Martínez,
FMC 114067]
Cundinamarca: [“Bogotá”: Coll.Fulton, CMC C10818; DMNH 183731] [(as P. adamsoni):
Villeta, near Bogotá, DMNH 137596] [“Guaduas, Colombia” (Abbott, 1989)]
Tolima: [Basis of Páramo, near Tolyma (=Tolima) volcano (Breure & Schouten, 1985)]
Sultana (Melaniorthalicus) atramentaria (Pfeiffer, 1855)
Boyacá: [Below Soatá, and road to Río Chicamocha, arid zone, in cactus-like plants, Coll.:
R.Jaramillo/1958, FMC 114056, 114131] [San Mateo, Marvoques Villarreal, Coll.
J.Matau/Jan.1942, DMNH 129037]
Cundinamarca: [Bogotá, Coll.Fulton, CMC C11118]
Putumayo: [Puerto Asís, Upper Río Putumayo, Coll.: F.Medem/1958, FMC 114057] [Santa Rosa
(Kofan Indian Village), Headwaters of Río San Miguel, Coll.: B.Malkin & P.Burchard/1970,
FMC 168190]
Norte de Santander: [Pamplona (Abbott, 1989)]
Santander: [Near El Galán, N of Bogotá, FMC 109717] [Near San Gil, N of Bogotá, FMC
109718] [San Gil: Coll.Hno.G.Victor, pre-1935 (Pilsbry, 1935); Juvenile, Coll.1931, DMNH
148706] [Capinanejo, Coll.P.E.Acosta, 1960, FMC 114055] [Miranda, Coll.P.E.Acosta/1960:
FMC 114058; Trail to El Salado, FMC 114130] [San Andrés, Quebrada de Pangote, Coll.:
P.E.Acosta/1960, FMC 115801]
?: [Columbia, SMF 155 498 (Zilch, 1960)]
Sultana kelletti Reeve, 1850
Cundinamarca: [(as Porphyrobaphe fungairinoi), Bogotá, DMNH 137597]
Sultana (Melaniorthalicus) powisianus (Petit, 1843)
Quindio: [Syntype MNHN, “Montagne de Quindiu” (Breure & Scouten, 1985)]
Caldas: [Marmato & forests on Cauca River (Pilsbry, 1899)]
Santander: [San Gil, juvenile, DMNH 213707]
Valle del Cauca: [Cali, in W Colombia, Coll.Hno.Apolinar María, pre-1935 (Pilsbry, 1935)]
[Atuncelo area: 3 km.W, 4500 ft., Collec.: F.G. Thompson,1969, FLMNH 21276; 9 km. N of
Dagua, 2800 ft., Collec.: F.G. Thompson/1969, FLMNH 21277]
Cauca(?)or Risaralda(?): [Metorthalicus powisianus (Petit, 1843), “Cauca Dept. Pereira”,
Coll.:1932, FLMNH 166256-257]
Borrero, F.J. Colombian non-marine mollusks
28
Sultana sultana (Dillwyn, 1817)
Distribution: Northern South America, Peru, Bolivia (Simone, 2006)
Córdoba: [Upper Río Uré, trib.of R.San Jorge, Coll.:CVM.Expedition/1963, FMC 187607]
Caquetá ??: [S. sultana (Dillwyn, 1817), Río Mecaya, trib.to Upper Caquetá, Coll.: H.Granados
& H.Arévalo/1957, FMC 114060]
Chocó: [Pangala, about 40 km N of Palestina, Río San Juán, Coll.: B.Malkin/1972, FMC 170627]
Putumayo: [Santa Rosa (Kofan Indian Village), Headwaters of Río San Miguel, Coll.: B.Malkin
& P.Burchard/1970, FMC 168192]
Vaupés: [S. sultana (Dillwyn, 1817), Caño Tucunemita, 4 hrs on foot W of Lago El Dorado,
Coll.: F.Medem/1958, FMC 114059]
Corona regalis (Hupé, 1857) [relationship between C. regalis, C. regalis loroisiana & C.
loroisiana ?]
Distribution: Brazil, Peru, Ecuador (Simone, 2006)
Corona regalis loroisiana Hupe 1857
Distribution: (As Corona loroisiana Hupé, 1857) Peru, Brazil (Simone, 2006)
[“North Colombia”, FMC 421]
Corona regulis Hupe
[“New Grenada”, Coll.:J.G.Anthony, ANSP 1217]
Corona regina (Férussac, 1821) [Simone has it as Fèrusaac in Bowdich, 1822)]
Ditribution: Northern South America (Simone, 2006).
[Lectotype MNHN (45 mm); Paralectotype MNHN (35mm) (Simone, 2006)]
Putumayo: [Santa Rosa de Sucumbios, Río San Miguel, Coll.: B.Malkin& P.Burchard/1970-71,
FMC 170613-614, 170616]
Valle del Cauca: [Restrepo et al.,1983: Paso de la Bolsa, UVZ 82-154/1; Hacienda El Castillo,
Jamundí, UVZ 8186/1; Hacienda Formosa, Ansermanuevo UVZ 82-037/1]
[“Colombia”, FLMNH 206236]
Hemibulimus (Myiorthalicus) dennisoni (Reeve, 1848)
Caldas: [Marmato (Pilsbry, 1899)]
Valle del Cauca: [Type [Unconfirmed?] MCZ 216993, Bitaco, Coll. E.N.Kozloff, July 1954, 1
spec. (MCZ online catalog June 2007)]
[Rio Aguacatal, SMF 155 499 (Zilch, 1960)] [Bosque de Yotoco, Colombia; F.J Borrero’s
collections, 1979-82, deposited at UVZ] [Cordillera Occidental, locations along the road CaliBuenaventura (Restrepo & Breure, 1987)]
Cundinamarca: [Near Bogotá (Abbott, 1989)] [Bogotá, CMC C11117]
Cundinamarca?/Valle del Cauca?: [“Canca, Bogotá”, Coll.Fulton, CMC C10727 (There is a
town called Canca in Valle del Cauca)]
Hemibulimus dennisoni carus Pilsbry, 1909
Cauca: [Quilichao (=Santander de Quilichao): Lectotype ANSP 97716, 5500ft (Breure &
Scouten, 1985); Coll.: H.C. Fulton, FMC 146210; CMC C11116]
?: [“Colombia”: as Orthalicus carus Pilsbry, DMNH 015395; as Porphyrobaphe (Hemibulimus)
carus, Coll.Fulton, CMC C10728]
Borrero, F.J. Colombian non-marine mollusks
29
Hemibulimus excisus Martens, 1885
Cauca: [as H. magnificus (Pfeiffer): “near Popayán” (Pilsbry, 1899; Breure & Schouten, 1985)]
?: [“S Columbia”, SMF 90 698 (Zilch 1960)] [“Colombia” (p. 140, Parkinson, 1987)]
Porphyrobaphe approximata Fulton, 1896
Cundinamarca: [Bogotá (Breure & Schouten, 1985)] [Near Bogotá (Abbott, 1989]
Porphyrobaphe dennisoni (Reeve, 1848) [=Hemibulimus dennisoni (Reeve, 1849) ?]
Cauca: [“Bogotá”, FLMNH 22164]
?: [“Cauca River”, FLMNH 166228] [Bogotá-Cauca, FLMNH 166229]
Porphyrobaphe denisoni var marmatensis Pilsbry, 1899 [=Hemibulimus dennisoni (Reeve,
1849) ?]
Caldas: [Holotype ANSP 26124, Marmato (Breure and Schouten, 1985), Coll.Bland]
Porphyrobaphe denisoni obscurata Mousson, 1873
Cundinamarca: [Bogotá (Breure & Schouten, 1985)]
Porphyrobaphe (Porphyrobaphe) iris (Pfeiffer, 1852) (=P.wallisianus Mousson, 1873, which is
a variety of P. iris)
Antioquia: [La Ceja (Type locality), DMNH 137598; “Páramo de Frontino”, FLMNH 166233;
Betania, DMNH 142265; Yarumal, DMNH 137600]
Cundinamarca: [Lectotype ZMUZ 511864, “near Bogotá”, Coll. Wallis/1872 (Breure, 1976)]
[Bogotá, FLMNH 166232]
Caldas: [Pensilvania, DMNH 137599] [Laguneta, NRS 2748 (Breure & Schouten, 1985)]
?: [“Mountains of Colombia” (Abbott, 1989)] [“North Colombia”, FMC 410, 31522]
Porphyrobaphe iostoma (Sowerby)
See Univalle database
Porphyrobaphe irroratus (Reeve, 1849)
Antioquia?: [“La Ceya” (La Ceja ?), DMNH 137598]
[“Colombia”: FLMNH 166235; DMNH 015318] [P. irrorata: “Colombia”, FMC 31164;
“Columbia” CMC C10829]
Porphyrobaphe yatesi (Pfeiffer, 1855) [=Porphyrobaphe sarcostoma Ancey, 1903]
?: [“Colombia (?)” (Ancey, 1903, 1904)]
Porphyrobaphe sp.
Nariño: [El Morro, near Tumaco, Camp Pesmaco Co., Coll.: F.Medem & C.A.Velásquez/1965,
FMC 187285]
Family Orthalicidae
Subfamily Bulimulinae
Bulimulus bilineatus Sowerby 1833b [It may be a Bostryx (Breure, 1879), see Bostryx]
[“Columbia”, ANSP 25504,]
Borrero, F.J. Colombian non-marine mollusks
30
Bulimulus buenavistensis (Pilsbry, 1897)
Córdoba: [=Bulimulus corneus (Lea, 1838), Holotype ANSP 192931, Paratype USNM 105078,
“Buenavista”, Coll.T.R.Peale (Abott, 1955; Breure, 1979)]
Magdalena: [Codazzi, Coll.: O.Haught/1943, FMC 72759]
Bulimulus cacticolus (Reeve, 1849)
La Guajira: [“Santa Marta”, Coll.:H.H.Smith, FLMNH 109611]
Bulimulus corneus (Sowerby, 1833). [“Terrestrial species, lives on grass at low heights from the
floor” “distributed from Guatemala to Colombia (Martens, 1890-1901). B. unicolor (Sowerby) is
apparently a synonymn (Pérez y López, 1993)
Bulimulus dysoni (Pfeiffer, 1846)
Magdalena: [4 mi N of Bonda, Coll.: H.H.Smith/1900, FMC 78785]
Bulimulus effeminatus (Reeve, 1848)
[Lectotype BMNH 1975508, “Valley of the Madeleine” (=Río Magdalena, Colombia) ( Breure,
1978)]
Bulimulus erectus (Reeve, 1849) [Cited as Drymaeus (check subgenus) erectus (Reeve, 1849) by
Simone (2006)]
Distribution: Venezuela, Brazil (Simone, 2006), Colombia (this paper).
Atlántico: [Barranquilla, Coll.: H.Dybas/1938, FMC 17511]
Bulimulus eyriesii (Drouët, 1959)
La Guajira: [Coll.: L.Zanella/1949, FMC 109695]
Bulimulus glandiniformis Sowerby, 1892
Cundinamarca: [U.S. of Colombia, Bogotá (Breure, 1979)]
Bulimulus hyaloideus (Pfeiffer)
Cundinamarca: [Utica, Bogotá-La Dorada Railroad, 105 km NW of Bogotá,
Coll.:L.Zanella/1949, FMC 109693] [Valley of Río Bogotá, Coll.: H.C.Raasvelt/1948, FMC,
109696]
Bulimulus incolor Sowerby
[“New Granada”, Coll.: Luders, ANSP 5255]
Bulimulus juvenilis Pfeiffer, 1855
Cundinamarca: [Lectotype BMNH 1975161, Santa Fé de Bogotá (Breure, 1979)]
Norte de Santander: [(B.cf. juvenilis Pfeiffer)10km S of Cúcuta, 375m (Breure’s Sta.182)(Breure,
1977a)]
Bulimulus krebsianus (Pilsbry, 1897)
Borrero, F.J. Colombian non-marine mollusks
31
Bolívar: [Type ? ANSP 71434, , “Carthagena” (Cartagena)] [Convento de La Papa, near
Cartagena, Coll.H.Sturm (Breure, 1977a)]
Magdalena: [Río Cesar Valley, Coll.: O. Haught/1938, FMC 72760] [Near Valledupar, FMC
109686] [“Specimens collected between Santa Marta and El Rodadero are tentativelu referred to
this species (SMF 245412/4 and 245413/2 respectively”, Coll. H.Sturm (Breure, 1977a)]
La Guajira: [Coll.: L.Zanella/1949, FMC 109681]
Bulimulus marmatensis Pfeiffer, 1855
Caldas: [Marmato, U.S. of Colombia, (Breure, 1979)]
Bulimulus pallidior Sowerby
[“New Grenada”, ANSP 25703 ]
Bulimulus aff. pilosus (Guppy)
Córdoba: [Río Sinú, La Floresta, Coll.: L.Zanella/1950, FMC 109691]
Bulimulus tenuissima Férussac, 1832 (Colombia??)
Bolívar: [Cartagena, CMC C?? (material is in a problem box)]
Bulimulus vestalis Albers (?Colombia)
Amazonas?: [“north bank of River Marañon” (Haas, 1951) (Thus Colombia?)]
Drymaeus (Drymaeus) alabastrinus Da Costa, 1906
Tolima: [Holotype BMNH 1907.11.21.16, Honda, (Breure, 1979)]
Drymaeus (Drymaeus) angusta Da Costa, 1906
Cundinamarca: [Holotype BMNH 1907.11.21.14 (Breure, 1979)]
Drymaeus (Drymaeus) antioquensis (Pfeiffer, 1855)
Antioquia: [Lectotype BMNH 1975450, “Prov.de Antioquia, New Grenada”, (Breure & Eskens,
1981)]
Drymaeus atteneatus (Pfeiffer) [Drymaeus (Drymaeus) attenuatus (Pffeifer, 1853a)]
Arauca: [Coll.: M.Medem-Cortez/1975, USNM 853418]
Drymaeus (Drymaeus) baranguillanus (Pfeiffer, 1853)
Atlántico?: [Lectotype BMNH 1975452, “Baranguilla in Andibus Columbianus”
(Barranquilla ??), (Breure, 1979)]
Drymaeus (Drymaeus) bellus Da Costa, 1906
Meta?: [Holotype BMNH 1907.11.21.32, San Martin, (Breure, 1979)]
Drymaeus (Drymaeus) blandi (Pilsbry, 1898)
Boyacá: [Muzo, FLMNH 161214]
Cundinamarca: [Páramo de Usaquén, Cordillera Oriental, N of Bogotá, Coll.: J.I.Idrobo, 1961,
FMC 114078] [San Miguel (Fusagasugá), Coll.: Richter/1943, FMC 114105]
Borrero, F.J. Colombian non-marine mollusks
32
Tolima: [near Fresno: Lectotype ANSP 25780 (Breure, 1979); FMC 78761]
Vaupés: [Barranco Picrue, Coll.: 1957, FMC 114121]
Drymaeus (Drymaeus) bogotensis (Pfeiffer, 1855)
Cundinamarca: [Santa Fé de Bogotá: Lectotype BMNH 1975191 (Breure & Eskens, 1981);
Mesitas del Colegio, RMNH (op.cit.); FMC 31476] [“Bogotá Depto.”, FLMNH 206150]
Caldas/Risaralda?: [“between Salamina and Cabuyal, west of (Bland)” (Pilsbry, 1897-8)]
? Drymaeus (Drymaeus) bolivianus Pfeiffer, 1846. [Questionable entry: Breure (1979) lists
“Drymaeus (D.) bolivianus Pfeiffer, 1846a: 34 (Merida, Andes of Bolivia [sic]) [LT BMNH
1975444]
Valle del Cauca: [Type [unconfirmed?] MCZ 216995, 2 km S. of Vijes, Coll. E.N.Kozloff, 25
June 1954, 6 specs. (MCZ online catalog, June 2007)]
Drymaeus (Drymaeus) cantatus medinamus Pilsbry, 1935
Cundinamarca: [Holotype ANSP 164752, Region of Medina, eastern slope of the Cordillera, E
of Bogotá, Coll.M.Gonzalez, pre-1935 (Pilsbry, 1935; Breure, 1979)]
Drymaeus (Drymaeus) caucaensis Da Costa 1898
Valle del Cauca?: [Holotype BMNH 1907.11.21.43, “Valley of the Cauca River” (Breure, 1979;
Pilsbry 1898)]
Drymaeus (Drymaeus) chimborasensis (Reeve, 1848)
Cundinamarca: [Sitio Tequendama I, Soacha (Correal & van der Hammen, 1977)]
Drymaeus (Drymaeus) cognatus Pilsbry, 1901
Cundinamarca: [Lectotype ANSP 78543, Bogotá (Breure,1979)]
Drymaeus (Drymaeus) comis Preston, 1907
Cundinamarca: [Bogotá (Breure,1979)]
Drymaeus (Drymaeus) confluens (Pfeiffer, 1855)
Caldas?: [Lectotype BMNH 19755196, Marmato (Breure, 1979)]
Drymaeus (Drymaeus) dacostae Sowerby, 1892
Cundinamarca: [Holotype BMNH 1907.11.21.51, Bogotá, (Breure, 1979)]
Drymaeus (Drymaeus) decoratus (Lea, 1838)
Bolívar: [near Carthagena (=Cartagena) (Breure, 1979).
Drymaeus decoratus goniobasis Pilsbry, 1898
Magdalena?: [Holotype ANSP 25803, near Santa Ana (Breure,1979)]
Drymaeus (Drymaeus) elegantissimus (Mousson, 1873)
Cundinamarca: [Bogotá, FMC 31419]
?: [ZMUZ 512110, 512111 (Breure, 1976)]
Borrero, F.J. Colombian non-marine mollusks
33
Drymaeus (Drymaeus) exoticus Da Costa, 1901
[Holotype BMNH 1907.11.21.38, Upper Magdalena River (Breure, 1979)]
Drymaeus (Drymaeus) expansus (Pfeiffer, 1848)
Distribution: Brazil (Simone, 2006), Colombia.
Putumayo: [Puerto Asís, Upper Río Putumayo, F.Medem, 1958, FMC 114079]
Drymaeus (Drymaeus) eversus Mousson, 1873 (?Colombia)
[“Nördlichem Süd-Amerika” (Breure, 1979)]
?Drymaeus (Drymaeus) fabrefactus Reeve, 1848 (?Colombia)
?: [Lectotype, BMNH 1975531, “New Granada, Province of Merida” (Venezuela) (Breure,
1979)]
Cundinamarca?: [Santa Fé de Bogotá (Pilsbry, 1898); “Bogotá” (Baker, 1926]
Drymaeus (Drymaeus) felix (Pfeiffer, 1862)
Cundinamarca: [Bogotá, DMNH 194935, 183910] [Bogotá, Distrito Especial, FLMNH 109230,
161241, FMC 146882]
Meta: [“road from Llanos of San Martín, and low land, and in forest at Lebonday, 2100m”
(Pilsbry, 1897-8)] [Villavicencio Region, Coll.: Guevara, pre-1935, FMC 78749]
Drymaeus (Drymaeus) flexuosus (Pfeiffer, 1852)
Cauca: [“Quilachao” (Santander de Quilichao), FLMNH 161244, 109238]
Meta: [(Drymaeus flexosus) Villavicencio, Coll.: H.Dybas/1838, FMC 3377]
Caldas: [Marmato: FLMNH 109239; “Marmato” (Breure & Eskens, 1981); DMNH 025595]
Valle del Cauca: [3 km.W Atuncelo, 4500 ft., Coll.: F.G.Thompson/1969, FLMNH 176885]
?: [“Yerico” (=Jerico?), DMNH 164056 (could be Antioquia, Huila, Tolima, Magdalena,
Santander, Boyacá, or Valle del Cauca)]
Drymaeus flexuosus megas Pilsbry 1944
Huila: [Holotype ANSP 179981, Upper Magdalena Valley near Suesa 03° 15N-075° 15W,
Coll.:A.A.Olsson/1943]
Drymaeus (Drymaeus) fresnoensis Pilsbry, 1898
Tolima: [Holotype ANSP 25836, Fresno (Breure, 1979)]
?Drymaeus (Drymaeus) fucatus Reeve, 1849 (?Colombia: Could be Sibundoy Ecuador, not
Colombia)
Putumayo?: [New Granada, Sebundoi (=Sibundoy?), (Breure, 1979)]
Drymaeus (Drymaeus) geometricus (Pfeiffer, 1846)
Cundinamarca: [Bogotá, FMC 77575]
?: [Lectotype BMNH 1975564, “Novae Granadae, vallis Magdalenae” (Breure, 1979)]
Risaralda/Caldas?: [Forests in the mountains below Ervé, on the road to Santa Ana (Pilsbry,
1898)]
[“New Granada”: Coll.: T.Bland, ANSP 25770; FMC 78750] [“Colombia”, FLMNH 206172]
Borrero, F.J. Colombian non-marine mollusks
34
Drymaeus (Drymaeus) inclinatus Pfeiffer, 1862 (?Colombia)
[Lectotype, BMNH 1975532, “New Granada”, Breure (1979); U.S. of Colombia (Pilsbry, 1898)]
Drymaeus (Drymaeus) incognita Da Costa, 1907
Cundinamarca: [Holotype BMNH 1907.11.21.24, Bogotá (Breure, 1979)]
Drymaeus (Drymaeus) leai Pilsbry, 1898 (new name for Bulimus gracilis Lea, 1838 (=Drymaeus
(Drymaeus) gracilis Lea) (Breure, 1979).
Bolívar: [Type USNM 105,154; near Carthagena (Pilsbry, 1898); Breure, 1979)]
Magdalena: [Mountains near Santa Marta, 3600 ft, Coll.:H.H.Smith, FLMNH 109285]
[“Colombia”, FLMNH 109286]
Drymaeus (Drymaeus) membielinus (Crosse, 1867)
Cundinamarca: [Bogotá, Distrito Especial, FLMNH 161263] [“Bogota” (=Bogotá): DMNH
183909; D. mebielinus (Crosse, 1867), FMC 31408]
Drymaeus (Drymaeus) murrinus (Reeve, 1848)
Cundinamarca: [Lectotype BMNH 1975213, Santa Fé de Bogotá (Breure, 1979)] [Localities
near Bogotá, Coll. H.Sturm, SMF 245399-404 (Breure, 1977a): Tibatitá; 6 km N Usaquén;
valley E of Calle 82; 18 km N of Calle 79] [Bogotá, Distrito Especial, FLMNH 109324, 161268]
[Bogotá: FMC 77538, 77577; DMNH 184294]
Nariño??: [“Drymaeus sp. aff. murrinus”, 5 mi E of Santiago, Coll.:E.S.Ross &
E.I.Schlinger/1955, FMC 106317]
Drymaeus (Drymaeus) narcissus (Albers, 1854)
?: [“U.S. of Colombia”, FMC 94876]
Drymaeus (Drymaeus) notabilis Da Costa, 1906
Antioquia: [Holotype BMNH 1907.11.21.5, “Antioquia” (Breure, 1979)]
Drymaeus (Drymaeus) notatus Da Costa, 1906
Antioquia: [Holotype BMNH 1907.11.21.6, “Antioquia” (Breure, 1979)]
Drymaeus (Drymaeus) pamplonensis Pilsbry, 1939
Norte de Santander: [Holotype ANSP 170699, Pamplona, Coll.: Brother Nicéforo María (Pilsbry,
1939)]
Drymaeus (Drymaeus) pealianus Lea, 1838
?: [Holotype ANSP 192929, “near the rapids of Angostura, Colombia”, Coll.T.R.Peale (Abbott,
1955; Breure, 1979) (Vichada?-banks of Orinoco River; Antioquia?-Munipality; Tolima?Mount)]
Drymaeus (Drymaeus) poecilus (d’Orbigny, 1835) [“Columbia (locality data degraded)”, ANSP
25824]
Borrero, F.J. Colombian non-marine mollusks
35
Drymaeus (Drymaeus) plicatoliratus Da Costa, 1898
Cundinamarca: [Holotype BMNH 1907.11.21.120, Bogotá (Breure, 1979)]
Drymaeus (Drymaeus) pseudofusoides Da Costa, 1906
Cundinamarca: [Holotype BMNH 1907.11.21.11, Bogotá (Breure, 1979)]
Drymaeus (Drymaeus) rehderi Parodiz, 1962
Antioquia: [Holotype, USNM 590653, Valdivia, 4500ft on Cordillera Oriental,
Coll.M.A.Carriker, June/1948]
Drymaeus (Drymaeus) rugistriatus Haas, 1952. [Haas, 1952]
Valle del Cauca: [Restrepo et al., 1983: Paso de la Bolsa UVM 82-152/1, Puerto Tejada UVM
81-239/1, Hacienda “E; Castillo” Jamundí UVM 8162/3, 8164/3,82054/2]
Drymaeus (Drymaeus) sanctaemarthae Pilsbry, 1901
Magdalena: [Jiracasaca, NW slope of Santa Marta Mountains (Breure, 1979)]
Drymaeus (Drymaeus) smithii Da Costa, 1898
Cundinamarca: [Lectotype BMNH 1907.11.21.52, Bogotá (Breure, 1979)]
Drymaeus (Drymaeus) solidus Preston, 1907
Cundinamarca: [Syntype BMNH, Bogotá (Breure, 1979)]
Drymaeus (Drymaeus) spadiceus Da Costa, 1906
Cundinamarca: [Holotype BMNH 1907.11.21.15, Bogotá (Breure, 1979)]
Boyacá: [Muzo, DMNH 164093]
Drymaeus (Drymaeus) spectatus Reeve, 1849
?: [Lectotype BMNH 1874.12.21.15, New Granada (Breure, 1979)]
Cauca: [“road between Popayan to the Hacienda Sotara… at about 2400m (Stübel)(Pilsbry 18978)]
Drymaeus (Drymaeus) subsemiclausus (Petit, 1843)
Quindío?: [Syntype in the Paris Museum, “Montagne de Quindiu” (Breure & Eskens, 1981)]
?: [Province of Bogota, Colombia (Pilsbry, 1898)]
Drymaeus (Drymaeus) subventricosus Da Costa, 1901
Cundinamarca: [Holotype BMNH 1907.11.21.37, Bogotá (Breure, 1979)]
Drymaeus (Drymaeus) trivittatus Mousson, 1873
Cundinamarca: [Bogotá (Pilsbry, 1898)]
Drymaeus (Drymaeus) tusagasuganus Pilsbry, 1935
Cundinamarca: [Holotype ANSP 164570, Tusagasugá (=Fusagasugá), Coll.V.Guevara, pre-1935
(Pilsbry, 1935; Breure, 1979)]
Borrero, F.J. Colombian non-marine mollusks
36
? Drymaeus (check subgenus) vannattai Pilsbry, 1898 [? Colombia]
Distribution: Amazon River (Simone, 2006).
Drymaeus (Drymaeus) ventricosus Preston, 1907
Cundinamarca: [Bogotá (Breure, 1979)]
Drymaeus (Drymaeus) vicinus Preston, 1907
Cundinamarca: [Bogotá (Breure, 1979)]
Drymaeus (Drymaeus) villavicencioensis Breure, 1977
Meta: [Holotype SMF 245417, Villavicencio, Coll.F.Schremmer/1970 (Breure, 1977a;1979)]
Drymaeus (Drymaeus) violaceus (Mousson, 1873)
Cundinamarca: [Lectotype ZMUZ 512087, “near Bogotá”, Coll. Wallis, 1872 (Breure, 1976)]
Caldas: [Marmato, Coll. Bland (Pilsbry, 1897-8)]
Cauca ??: [“Cauca”, Coll.: F.Bonis, FMC 78780]
[“Colombia”, FLMNH 206222]
Drymaeus (Drymaeus) ziczac Da Costa, 1878
[Holotype BMNH 1907.11.21.46, “Valle of the Rio Cauca”, (Breure, 1979)]
Drymaeus (Drymaeus) zingarensis Restrepo & Breure, 1987
Valle del Cauca: Restrepo & Breure (1987): [Holotype UVM 83-010, Finca Zingara, km. 6 on
the trail km.19 road Cali-Buenaventura-La Paz, 2000 m. (3°31’N-76°36’W), Coll.:
M.Giraldo/1983; Paratypes UVM 81-252/1, 82-032/1, 82-194/2, 83-007/1] [Locations on road
Cali-Buenaventura: km. 14, Bosque de San Antonio, W of Cerro de la Horqueta, 2100 m.
(3°28’N-76°40’W), UVM 82-131/2, 82-133/3, 82-164/1, 83-036/1, 83-115/1, RMNH 55838/1
(Paratypes); km.16, Bosque de San Pablo, 1900 m. (3°29’N-76°38’W), UVM 81-060/1, 82210/1, RMNH 55839/1 (Paratypes)]
Drymaeus (Mesembrinus) cactivorus Broderip in Broderip & Sowerby, 1832
Bolívar?: [“ad Montem Chris in Columbia” (=Montecristo ?) (Breure, 1979)]
Drymaeus (Mesembrinus) colmeiroi (Hidalgo, 1872)
Valle del Cauca: [Road to Campo Alegre, Lago Calima, 1600m, Coll.: J.Cantera (Breure &
Eskens, 1981)]
Drymaeus (Mesembrinus) columbianus (Lea, 1838)
?: [Lectotype ANSP 192932, “about 100 miles up the Magdalena River, Colombia”,
Coll.T.R.Peale (Abbott, 1955; Breure, 1979)]
Drymaeus (Mesembrinus) depictus (Reeve, 1849)
[“New Granada”: Lectotype BMNH 1975529 (Breure, 1979); ANSP 25831, 25832]
Drymaeus depictus pervariabilis (Pfeiffer) [“Colombia”: FMC 77559; Pilsbry (1898)]
?Drymaeus (Mesembrinus) deshayesi (Pfeiffer, 1845) (?Colombia)
Borrero, F.J. Colombian non-marine mollusks
37
[“New Granada”, Lectotype BMNH 1975526 (Breure & Eskens, 1981)]
Drymaeus (Mesembrinus) dominicus Reeve, 1850 [“New Granada”, ANSP 25843]
?Drymaeus (Mesembrinus) dubius Pfeiffer, 1853 (?Colombia)
[“Andes of New Granada”, Lectotype BMNH 1975519 (Breure, 1979, 1981)]
Drymaeus dubins Pfeiffer ? [“New Granada”, Coll.: Luders, ANSP 5259]
Drymaeus dybuys Pfeiffer, 1853 [“Colombia”, Breure (1979)]
Drymaeus (check subgenus) edmulleri (Albers, 1854)
Distribution: Marañón River, Colombia (Simone, 2006)
Amazonas?: [Marañón River, Colombia, ZMB 111929, 30mm (Simone, 2006)]
Drymaeus (Mesembrinus) fasciarum Pilsbry, 1939
Norte de Santander: [Holotype ANSP 168429, Cúcuta, Coll.: Brother Nicéforo María (Pilsbry,
1939)]
Santander: [~ 15km S of Málaga, Río Servitá valley, ca.1800m (A.S.H.Breure Sta.184), at
RMNH (Breure, 1977a)]
?Drymaeus (Mesembrinus) fidustus (Reeve, 1849) [?Colombia: Could be Sibundoy Ecuador, not
Colombia]
Putumayo?: [Lectotype BMNH 1975517, New Granada, Sebundoi (Sibundoy?) (Breure, 1979)]
Drymaeus (Mesembrinus) flavidus (Menke, 1829) (?Colombia)
??: [“from Venezuela to Panamá” (Haas, 1966) (Thus Clombia?)]
Drymaeus (Mesembrinus) gorgonensis Haas, 1966
Cauca: [Holotype FMC 114164, Island of Gorgona, 78°11’W-3°N, Coll.F.Medem, Feb.1961]
Drymaeus (Mesembrinus) gratus (Pfeiffer, 1856)
[(=Drymaeus columbiensis Pfeiffer, 1855), Lectotype BMNH 1975521, “Columbia” (Breure,
1979)]
Cundinamarca: [Sitio Tequendama I, Municipio de Soacha (Correal & van der Hammen, 1977).
?: [Internet paper, Andes Orientales (Rodríguez, 1999)]
Drymaeus (Mesembrinus) koppeli (Sowerby, 1892)
Cundinamarca: [Bogotá: Lectotype BMNH 1907.11.21.133 (Breure, 1979); FMC 31393]
[Bogotá, Distrito Especial, FLMNH 109279- 280]
Drymaeus (Mesembrinus) lacteus (Lea, 1838)
??: [Lectotype ANSP 192930, “about 100 miles up the Magdalena River”, Coll.T.R.Peale
(Abbott, 1955; Breure, 1979)]
Magdalena: [Near Codazzi, Coll.: O.L. Haught/1943, FMC 117070]
Drymaeus (Mesembrinus) laetus Reeve, 1849
Cundinamarca: [Sitio Tequendama I, Soacha (Correal & van der Hammen, 1977)]
Borrero, F.J. Colombian non-marine mollusks
38
Putumayo: [Lectotype BMNH 1975534, New Granada, Sebundoi, northern Ecuador
(=Sibundoy?) (Pilsbry, 1898; Breure, 1979)]
Nariño: [Pasto (Pilsbry, 1898)]
Drymaeus (Mesembrinus) loxanus (Higgins, 1872) [not listed in Breure (1979) for Colombia]
Cauca: [Between Popayán and Hacienda Sotará, at about 2400m (Pilsbry, 1898)]
?Drymaeus (Mesembrinus) manupictus Reeve, 1848 (?Colombia)
?: [Lectotype BMNH 1975522, “Venezuela” and “Merida, Columbia” (Breure, 1979; Breure &
Eskens, 1981; “Andes of Colombia (Pilsbry, 1898)]
Drymaeus (Mesembrinus) menkei (Gruner, 1841)
?: [“Colombia” (Femorale.com)]
Drymaeus (Mesembrinus) meridanus Pfeiffer, 1846
[“Colombia”, FLMNH 176954]
Drymaeus (Mesembrinus) muliebris Reeve, 1849
?: [Andes of Colombia (Pilsbry 1898)]
[“New Granada”, ANSP 25757]
Drymaeus (Mesembrinus) multilineatus (Say, 1825)
Magdalena: [Near Santa Marta, Coll.: H.H.Smith, FLMNH 109323] [Bahia de Neguange,
Coll.K.-J.Götting, 1976-78 (Götting, 1978)] [Between Santa Marta and El Rodadero, SMF
245414/2; Between Santa Marta and Ciénaga, SMF 245415/1 (Breure, 1977a)]
[Near Valledupar, Coll.: R.W.Barker, FMC 109677]
Atlántico: [Barranquilla & Savanilla (Pilsbry, 1899]
Bolívar: [Cartagena (Pilsbry, 1899]
Drymaeus (Mesembrinus) niceforonis Pilsbry, 1939
Norte de Santander?/Cundinamarca?: [Type ANSP 170697, “Colombia, Santa Librada”, Coll.:
Brother Nicéforo María (Pilsbry, 1939)] [Santa Librada is a town in N.de Santander, and a
Quebrada (creek) in Cundinamarca]
Drymaeus (Mesembrinus) nigrofasciatus (Pfeiffer, 1846) [should it have parenthesis? Not by
Simone (2006)]
Distribution: Venezuela, Brazil (Simone, 2006), Colombia (this paper).
Amazonas: [Puerto Nariño, Coll.:D.Avella/1958, FMC 114082]
Cundinamarca: [Bogotá, Distrito Especial, FLMNH 109325, 109326, 161271] [Bogotá, FMC
77560, 102483] [“Bogotá Depto.”, FLMNH 206192] [La Calera, just NE of Bogotá, Coll.: O
Haught, FMC 72764] [Laguna de Hernández, Sabana de Bogotá, Coll.: F.Medem/1958, FMC
114080] [Rocks W of Laguna de La Herrera, Munic.Mosquera, Sabana de Bogotá, Coll.:
R.Jaramillo/1960, FMC 114083] [Ubaté, Coll.: J.Carvajal/1958, FMC 115897]
Valle del Cauca: [D.(Mesembrinus) nigrofasciatus (Restrepo et al., 1983): Meléndez, Cali UVM
8181/1, Guadalupe, Cali UVM 82-162/1, La Paila, UVM 8174, 8180/3, Hacienda “Formosa”,
Ansermanuevo, UVM 82-041/8, 81-273/1] [Hacienda “El Castillo” Municipio de Jamundí;
Borrero, F.J. Colombian non-marine mollusks
39
common in bambú forests, and in low bushes along sugar cane plantations; (F.J.Borrero 19801982, unpublished observations)]
Santander: [Capitanejo, Coll.: P.E.Acosta, 1960, FMC 114081]
Drymaeus nigrofasciata elongatulus Pilsbry, 1898
Valle del Cauca: [Carmelo, 1000 ft., Coll.: F.G.Thompson/1969]
[“New Grenada”(locality data degraded), Coll.: Bt.fr. Seymour, ANSP 4346] D.nigrofasciatus
elongatulus Pilsbry, 1898
[“Colombia”, FLMNH 176962] [Syntype BMNH, “Novae Granadae, vallis Magdalenae”
(Breure, 1979)]
Drymaeus (Mesembrinus) parvus Lea, 1838
Bolívar: [near Carthagena (=Cartagena) (Breure, 1979)]
Drymaeus (Mesembrinus) pertristis Pilsbry, 1898
Norte de Santander: [Ocaña, Coll.: Wessel, FMC 31388]
?Drymaeus (Mesembrinus) pervariabilis Pfeiffer, 1853 (?Colombia)
[“New Granada”: Lectotype BMNH 1975547 “Novae Granadae” (Breure, 1979); Coll.:Luders,
ANSP 5240]
Drymaeus (Mesembrinus) rufescens pinchoti (Pilsbry, 1930)
San Andrés y Providencia: [Holotype ANSP 150852, Old Providence Isld., ½ mi. inland from
Southwest Bay] [Old Providence, Southwest Harbor, DMNH 164079] [Providencia Isld.:
13°30N-081°30W, on Cockspur acacias, one on bromeliad, Coll.:K.Emberton/1987, ANSP
A12126I; Hills near center of isld., from trunks of smooth-barked, greenwood trees, primarily
Cockspur, Coll.: C.R.Gilbert et al.,1971, FLMNH 176977; Manchioneel Hill, Coll.:
J.Geister/1970, FLMNH 176977] [Old Providencia Isld: Mount back of Isabel, Coll.:
Pilsbry/1929, FMC 78770; Saddle between Split Hill and next peak, Coll. H.A.Pilsbry
23Apr.1929, DMNH 135067, 026872, 030138, 007344; Coll. Barbour et al./March 1931,
DMNH 164080] [Saddle nr. Split Hill, FMC 146662]
Drymaeus rufescens providentialis Pilsbry, 1930
San Andrés y Providencia: [Providencia Isld., Coll.: K.Ebel/1998, FLMNH 271152-153]
Drymaeus (Mesembrinus) semifasciatus (Mousson, 1869)
?: [Lectotype ZMUZ 512301, “Columbia”, Coll.Wallis, 1869 (Breure, 1976)]
Meta: [Villavicencio, Coll.: H.Dybas/1938, FMC 3378]
Drymaeus (Mesembrinus) semimaculatus Pilsbry, 1898
Bolívar: [Type USNM 105, 153 Carthagena (Pilsbry, 1898)] [(=Drymaeus maculatus Lea, 1838),
near Carthagena(=Cartagena) (Breure, 1979)]
Drymaeus (Mesembrinus) sykesi Da Costa, 1906
Cundinamarca: [Holotype BMNH 1907.11.21.4, Bogotá (Breure, 1979)]
Drymaeus (Mesembrinus) studeri (Pfeiffer, 1847)
[Lectotype BMNH1975480, “Merida, New Granada”(Venezuela) (Breure & Eskens, 1981)]
Borrero, F.J. Colombian non-marine mollusks
40
Bolívar: [Rioviejo, Lower Río Magdalena, Coll.: F.Medem/1956, FMC 115803]
?Drymaeus (Mesembrinus) studeri primula Reeve (Uncertain taxon) [Breure & Eskens (1981)
list Drymaeus primulus (Reeve, 1848) as corresponding to material from Venezuela. They also
mention that Pilsbry (1898) synonimized this taxon with D. studeri (Pfeiffer, 1847), but
dissagree on this action on the basis of shell differences. Thus, the status of D.studeri primula is
uncertain]
[“Colombia”, FLMNH 177005]
Drymaeus (Mesembrinus) tenuilabris (Pfeiffer, 1866)
?: [Lower Magdalena (Pilsbry, 1898); “...probably lower valley of River Magdalena, Colombia”
(Baker, 1926)] [“from Venezuela and Colombia” (Haas, 1966)]
Drymaeus (Mesembrinus) translucens (Broderip in Broderip & Sowerby, 1832)
Cauca: [27 mi S of Popayán, Coll.: E.I.Schlinger & E.S.Ross/1955, FMC 106234]
Drymaeus (Mesembrinus) tribalteatus Reeve, 1848
Cundinamarca: [Santa Fé de Bogotá (Breure, 1979)]
?Drymaeus (Mesembrinus) venosus Reeve, 1848 [?Colombia: Could be on the Venezuelan side,
not Colombia]
Vichada?: [“Angostura, banks of the Orinoco” (Breure, 1979)]
Drymaeus (Mesembrinus) virgo (Lea, 1838)
Bolívar: [Type USNM 105,076, near Carthagena (=Cartagena) (Pilsbry, 1898)]
[“Colombia Mendez” (Tolima?), FMC 78783]
Drymaeus cleefi Breure & Eskens, 1981
Santander: [Holotype RMNH 55375, Paratypes RMNH 55376/9, ICNB/9, Páramo del
Almorzadero (06°59N’-72°44”W), Piedra de Molino, 4225m, Coll.:mA.M.Clef, J.Aguirre &
H.Hooghiemstra, 1978 (Breure & Eskens, 1981)]
Drymaeus equatorianus Smith
Nariño: directly across river from Chical, Ecuador, 1220m., Coll.:E.W.Schupp/1979,
FLMNH 26613]
Drymaeus fallax chicoensis Breure, 1977
Cundinamarca: [Holotype SMF 245405, Bosque de Chicó (NE Bogotá), 2700-2800 m, Coll.
H.Sturm (Breure, 1977a; 1979)] [Drymaeus chicoensis, Internet paper (Villegas, 2000)]
Drymaeus obliquus (Reeve)
Cundinamarca: [NB 18, Munic.Bojacá, Las Mercedes, road to Fute, S.of Laguna de
Herrera at limit of Sabana de Bogotá, Coll.: J.Hernández/1962, FMC 187576] [Suba,
Sabana de Bogotá, Coll.:Mejía/1962, FMC 187577]
Drymaeus roseatus montanus Pilsbry, 1901
Borrero, F.J. Colombian non-marine mollusks
41
Distribution: For Drymaeus roseatus (Reeve, 1848), Colombia, Venezuela, Brazil (Simone,
2006)
[Lectotype of D. roseatus (Reeve, 1848), BMNH 1975309, 37mm (Simone, 2006)]
Magdalena?/La Guajira?/Cesar?: [Western part of Santa Marta Mountains, Las Pantidas
(Breure, 1979)]
________________________________________________________
Drymaeus sp.
Chocó: [Acandí, Coll.: F.Keep, FLMNH 109122]
Drymaeus sp.
Valle del Cauca: [Atuncelo, 9 km.N Dagua, 2800 ft., Coll.:F.G.Thompson/1969, FLMNH 24412]
[MAY NOT BE THE SAME SPECIES, 3 km. W Cisneros, 1500 ft., Coll.: F.G.Thompson/1969,
FLMNH 176762]
Drymaeus sp.
Putumayo: [Santa Rosa de Sucumbios, Río San Miguel, Coll.: B.Malkin & P.Burchard, 1970,
FMC 187568] [Between Alpichaque & San Antonio, Santa Rosa de los Cofanes, below San
Antonio del Guamués, trib.of Upper R.Putumayo, Coll.: H.LeNestour/1965, FMC 187569]
Drymaeus sp.
Magdalena ?: [Sierra Nevada de Santa Marta: San Sebastián de Rábago, B.Malkin, 1968, FMC
157540, 161099; El Pueblito, Parque Nacional Tairona, Coll.: B.Malkin & P.Burchard/1970,
FMC 187599]
Bolívar: [tree close to bridge over Río Pechilín, road from Toluviejo to Sincelejo, Coll.:
F.Medem & C.A.Velásquez/1964, FMC 187575]
Cesar: [MAY NOT BE THE SAME SPECIES Sierra de Perijá, trail El Roncón, Finca San José,
10-25 km E Becerril, Coll.: B.Malkin/1969, FMC 163702, 173004, 173005, 173007-173011,
173857-173860; 15 km S of Becerril, La Jagua, FMC 187600]
[Socorpa Mission area, Sierra de Perijá, Coll.: B.Malkin/1968, FMC 187555, 187581, 187602604] [Valledupar, Coll.: B.Malkin/1968, FMC 187601]
Córdoba: [MAY NOT BE THE SAME SPECIES, Upper Río Uré, Trib.of Río San Jorge, Coll.:
L.Cepeda/1963, FMC 187572]
Drymaeus sp.
Cundinamarca: [Bogotá, Distrito Especial, FLMNH 109117, 109131] [Las Mercedes, Rd. to Fute,
SW slope of Tena, Munic.Bojacá, Coll.: Camacho/1962, FMC 187571]
Dryptus adoptus Reeve, 1849
Vichada?: [Banks of the Orinoco Breure (1979), (thus Colombia?)]
Dryptus funckii Nyst, 1843 [“Colombia”, Abbott, 1989] [May not be in Colombia: Breure (1979)
notes that its type locality is “province of Cumaná (Venezuela)]
Dryptus guerini (Pfeiffer, 1846)[form of D. funcki (Nyst, 1843) ??]
[Lectotype BMNH 1975272, Neu Granada (Breure, 1979)] [U.S. Colombia (Pilsbry, 1899);
“Colombia” (Abbott, 1989)]
Dryptus leptochilus Pfeiffer, 1848
Norte de Santander: [“prope La Baja in prov.Pamplona, Novae Granada“ (Breure, 1979)] [Baker
(1926) list this report as a synonym of Plekocheilus (Dryptus) pardalis (Ferussac)]
Borrero, F.J. Colombian non-marine mollusks
42
Dryptus lindeni Reeve, 1848
Norte de Santander: [La Baja in prov.Pamplona, New Granada (Breure, 1979)]
[(Strophocheilus lindeni): “New Grenada”, Coll. W.F.Webb, CMC C11024]
Dryptus moritzianus (Pfeiffer, 1847)
Santander del Norte: [Pamplona (Pilsbry, 1899)]
?: [(as Plekocheilus moritzianus Pfr.): “Alto Arauca”, DMNH 128665; “Sta. Librada” (=Santa
Librada), DMNH 128666 (could be Cauca, Tolima, Cundinamarca, or Vichada)]
[“Colombia”, FMC 31040; FLMNH 22129]
[“New Granada”, ANSP 3577-3579, 50486]
Dryptus moritzianus wilsoni Pilsbry, 1895
[(Bulimulus moritzianus wilsoni Pilsbry): Lectotype ANSP 7775, no type locality given (“New
Grenada”) (Breure, 1979)]
Dryptus pardalis (Férussac, 1821)
Norte de Santander: [La Baja, Province Pamplona, eastern Colombia (Baker, 1926)] [see
possible confussion with D. leptochilus (Pfeiffer)]
Cesar: [foothills of Sierra de Perijá, 10-30 km E of Becerril, El Roncón, Coll.: B.Malkin, 196869, FMC 163692-163694]
Magdalena: [Socorpa Mission trail, Becerril, Sierra de Perijá, Coll.:B.Malkin/1968, FMC 159207]
?: [“Sta. Librada”, DMNH 141725 (could be Cauca, Tolima, Cundinamarca, or Vichada)]
[ “Colombia”, Coll.Webb, CMC C11035]
Dryptus stuebeli Martens, 1885
Cundinamarca: [Fusagasugá, 1700 m (Pilsbry, 1899; Breure, 1979)]
Naesiotus cleeforum Breure, 1978
Valle del Cauca: [Holotype FLMNH 22769, 3 Paratypes FLMNH 22770/3, RMNH 55173/2, 9
km.N Dagua, Atunceto, 3500 ft., Coll.: F.G.Thompson/1969, ] [Paratypes FLMNH 22771/44,
RMNH 55172/7, Balneario Santa Rita-Cali, 1000 ft., Coll.:F.G.Thompson/1969 (Breure, 1978)]
Naesiotus gerenorum Breure, 1977
Boyacá: [Holotype RMNH 55115, Paratypes RMNH 55116, SMF 245410, FMNH 178108:
Upper Río Chicamocha Valley, few km S of Soatá, 2100 m (Breure, 1977a;1979)]
Naesiotus jullensorum Breure, 1977
Boyacá: [Holotype RMNH 55113, Paratypes RMNH 55114, SMF 245411, FMNH 178104:
Upper Río Chicamocha Valley, few km S of Soatá, 2100 m (Breure, 1979)]
Naesiotus quitensis catlowiae (Pfeiffer, 1854) CHECK WITH BREURE !!!! [is it Bulimulus
quitensis (Pfeiffer 1847)? Is it Bulimulus clatowae Pfeiffer 1847?]
Nariño: [Sandona, MCZ 173381, 5 specs, dry] [Pasto, MCZ 173382, 5 specs, dry]
Plekocheilus (Plekocheilus) appuni (Dunker, 1875)
Magdalena?/La Guajira?: [Type?, Sierra Nevada de Santa Marta (Breure, 1979)]
Borrero, F.J. Colombian non-marine mollusks
43
Magdalena: [Sierra Nevada de Santa Marta, watershed between Río Frío and Quebrada del
Páramo, 2875m, in humid forest (selva andina), Coll.A.M.Cleef et al./Aug.1977, RMNH/1
(Breure, 1977b)]
Plekocheilus (Plekocheilus) blainvilleanus (Pfeiffer, 1847)
[“Colombia”, FLMNH 109839-840] [Abbott, 1989; Parkinson, 1987]
Plekocheilus (Plekocheilus) cardinalis Pfeiffer, 1853
[“Colombia”, Coll.:1898-1899, ANSP 78579]
Plekocheilus (Plekocheilus) fulminans (Nyst, 1843)
Cundinamarca: [Monteredondo, Coll.: F.Medem/1956, FMC 115805]
Vichada?: [Angostura, (Vichada?-banks of Orinoco River) CMC C10997]
[“Columbia”, Coll.:Thorey, ANSP 328096]
?: [P. fulminans fulminans Nyst “has a wide distribution throughout Venezuela and Northeartern
Colombia” (Haas, 1955)]
?Plekocheilus (Plekocheilus) loveni Pfeiffer, 1848 (?Colombia)
[(Breure, 1979): Type locality “New Granada, Colonia Tovar” (Venezuela)]
Plekocheilus (Plekocheilus) lugubris Dunker, 1882
Nariño: [Pasto, “Columbiae australis” (Breure 1979)]
Plekocheilus (Plekocheilus) plectostylus (Pfeiffer, 1848)
[(Breure, 1978): Type locality “Chachopo province of Merida New Granada” (Venezuela)]
?: [“Colombia”: DMNH 183698; (Webb, 1948)] [“Columbia”, Coll.Fulton, CMC C11070]
Plekocheilus (Eurytus) ameguinoi Parodiz, 1962
?: [Type Locality: “Nueva Granada” (Colombia) (Parodiz, 1962)]
Plekocheilus (Eurytus) ampullaroides (Mousson, 1873)
Cundinamarca: [Lectotype ZMUZ 511584, “near Bogotá”, Coll. Wallis/1872, (Breure 1976;
1979)]
Nariño: [El Palusar, 1300 m, NRS 2754]
?: [“Villavicencio”, DMNH 141730 (could be Meta [most liquely], or Córdoba)]
Plekocheilus (Eurytus) apolinari (Pilsbry, 1935)
Meta: [Holotype ANSP 164566, region of Villaviencio, Coll. V.Guevara, pre-1935 (Pilsbry,
1935)]
Cundinamarca: [Monteredondo, Coll.: I. Scheible/1960, FMC 114075]
Plekocheilus (Eurytus) auriformis Da Costa, 1904
Cundinamarca: [Holotype BMNH 1907.II.2.88, (Breure, 1979)]
Plekocheilus (Eurytus) castaneus (Pfeiffer, 1845)
Cauca: [“Cauca Depto.”, FLMNH 206151; “Cauco” (=Cauca), DMNH 076768]
Borrero, F.J. Colombian non-marine mollusks
44
Quindío?: [Lectotype BMNH 1975279, Paralectotype BMNH 1975280, “Vegas of the River
Quenden (=Quindío?), Colombia (Breure, 1979)]
Tolima: [Espinal, DMNH 137440]
Cundinamarca: [Guayabetal, on road between Bogotá and Villavicencio, RMNH/1 (Breure,
1978)]
Tolima/Caldas: [“Near Fresno and Santa Ana”, and “Upper Magdalena Valley”, Colombia
(Breure, 1978); Pensilvania, DMNH 137442 (could be Tolima, Caldas, Huila); La Ceja, DMNH
128687 (could be Antioquia or Huila]
Plekocheilus (Eurytus) coloratus (Nyst, 1845)
?: [Lectotype BMNH 1975295, “Andes of New Granada” (Breure, 1978)]
Cauca: [Isla de Gorgona, North portion, humid forest near encampment, Coll.: F.Medem/1961,
FMC 114095]
Cesar: [Sierra de Perijá, 25-30 km E of Becerril, Finca América, Coll.: B.Malkin/1969, FMC,
163690] [El Roncón, foothills of Sierra de Perijá, 10-12 km E of Becerril, Col.: B.Malkin/1969,
FMC 163691]
Cundinamarca: [Bogotá: FMC 31113; DMNH 076770] [Sitio Tequendama I, Soacha (Correal &
van der Hammen, 1977)]
La Guajira: [Mountains near Santa Marta: 4000 ft, Coll.:H.H.Smith/1900, FLMNH 109842;
FLMNH 109843]
Magdalena: [Sierra Nevada de Santa Marta: 50 km SE Sta.Marta, just W of Río Buritaca along
trail to Ciudad Perdida, 11°2’N-73°55’W, Coll.:D.K.Dorman/1989, FLMNH 137920, 137933;
550 m., near trail beside Río Buritaca, FLMNH 137926; 11°1’B-73°55’W FLMNH 137932; San
Sebastián de Rabago, Coll.: B.Malkin/1968, FMC 157551, 173001; Near Minca, northern slope
of Sierra, approx.700m, Coll.K.J-Götting, 1976-78 (Götting, 1978)] [Socorpa Mission, forest
only, Becerril, Sierra de Perijá, Coll.:Malkin, 1968, FMC 159206, 159208, 159209-211, 172861,
173013]
Nariño: [Río Mataje, Pacific Coast on Colombo-Ecuatorian border, Coll.: ReichelDolmatoff/1962, FMC 187609] [El Palusar, 1300 m, NRS 2754/1-2755/2 (Breure, 1978)]
Putumayo: [Santa Rosa de Sucumbios, Río San Miguel, Coll.:B. Malkin/1971, FMC 175441]
Santander: [Trail to El Salado, Coll.: P.E.Acosta/1960, FMC 114053] [Malagavita, Coll.:
P.E.Acosta, 1960, FMC 114054] [(Götting, 1978)]
Tolima: [Toche, ca. 55 km NW Ibagué, 2500 m, NRS 2752/1 (Breure, 1978)]
Antioquia: [Frontino: Coll.Fulton, CMC C11047; DMNH 141721] [(Götting, 1978): Valle del
Río Magdalena (2500m); San Juán (250m); Potrero de Venancio (1100m); Pueblo Viejo]
Plekocheilus (Eurytus) coloratus coloratus (Nyst)
Cauca: [Popayán, FMC 31060]
Pleckocheilus (Eurytus) coloratus lamarkianus (Pfeiffer, 1848)
[“North Colombia”, FMC 411] [“Colombia” ANSP 66411] [“New Granada”, Coll.: H.Cuming,
ANSP 66427] [“Andes of New Granada, 8000 ft” (Baker, 1926)]
Plekocheilus (Eurytus) lamarckianus (Pfeiffer, 1848) [Same species ??]
Magdalena: [Sierra Nueva, Sierra Negra, Coll.: Carriker, FMC 71056]
Plekocheilus coloratus subplicatus (Pfeiffer, ?1855)
Magdalena: [Cerro San Lorenzo, Sierra Nevada de Santa Marta, Coll.: Carriker/1945, FMC
71055]
Norte de Santander: [“Provincia de Ocaña” (Breure, 1979)]
Borrero, F.J. Colombian non-marine mollusks
45
Plekocheilus (Eurytus) corticosus Sowerby, 1895
Cundinamarca: [Lectotype, BMNH 1907.11.21.110, Bogotá (Breure, 1979)]
[P. coritcosus (Sowerby, 1895), Monteredondo: km 75 road BogotáVillavicencio, Coll.: P.I.Scheible & H.Frohmund/1961, FMC 114165; Coll.:
P.I.Scheible,1959, FMC 114167]
?: [(as P. apolinari Pilsbry) “Piperal”, DMNH 141724]
Plekocheilus (Eurytus) dalmasi Dautzenberg, 1900
Magdalena?/Guajira?: [2 Syntypes (IRSN), “Sierra Nevada de Santa Marta”
(Breure,1978)]
Plekocheilus (Eurytus) doliarius Da Costa, 1898
?: [San Pablo, US of Colombia, CMC C494??]
Plekocheilus (Eurytus) elaeodes (Pfeiffer, 1853)
Boyacá: [Coll.A.S.H.Breure (Breure, 1977a): 10km NE of Villa de Leiva, Sta.121; Páramo de
Toquillo near Vado Hondo, 2900-3100m, SMF]
Cundinamarca: [Localities inmediately E of Bogotá, Coll. H.Sturm, SMF 245388-398 (Breure,
1977a): Tibatitá; Above Calle 82; Slope above Calle 85 and 86; Bosque de Chicó]
Plekocheilus (Eurytus) episcopalis (Pfeiffer, 1855)
Cundinamarca: [Lectotype, BMNH 1953. 11.30.1, Bogotá (Breure, 1979)] [“Bogata”(Bogotá),
FMC 31122; “Cordillera de Bogotá”, Coll.V.Guevara, pre-1935 (Pilsbry, 1935)]
XXXX: [“Angostura” CMC C???? (Antioquia?-Munipality; Tolima?-Mount; Vichada?-banks
of Orinoco River)]
Plekocheilus (Eurytus) gibbonius (Lea, 1838) [Simone says 1836 check!]
Distribution: Colombia, Brazil (Simone, 2006)
Boyacá: [Sogamoso: FLMNH 166220; DMNH 141726] [Humbo, DMNH 128663]
Chocó: [Pangala, ca. 40 km N of Palestina, Río San Juán, Coll.: B.Malkin/1972, FMC 174879]
Cundinamarca: [Bogotá, Distrito Especial, FLMNH 109851, 166221, FMC 31074] [Bogotá,
Coll.Fulton, CMC C10787, C11080; DMNH 137441] [Las Mercedes, Municipio Sasaima, Coll.:
E.Mutis/1958, FMC 114049] [La Mesa (San Javier), Coll.: Hernández/1955, FMC 114142]
Meta: [“Bogotá”, FLMNH 22134]
Quindío: [“Quenden Mts., Colombia”(=Quindío Mountains)(Abbott, 1989)]
Huila: [Between La Plata and Tocaima (Breure, 1979)]
Magdalena: [No.12 Finca “La Granja” Cerro San Lorenzo, Sierra Nevada de Santa Marta, Coll.:
F.Medem & C.A.Velásquez et al./1964, FMC 187286-287, 187683]
Plekocheilus (Eurytus) glandiformis (Lea, 1838)
Cauca: [Volcán de Puracé, Paletará, Coll.: 1951, FMC 109715] [“La Cumbre or Popayán(??),
Coll.: Pennell & Killip/1922, FMC 78741]
Huila: [Holotype, USNM 105045, Between La Plata and Tocaima (Breure, 1979)]
Plekocheilus (Eurytus) guentheri Sowerby, 1892 [“U.S. of Colombia” (Breure 1979)]
Borrero, F.J. Colombian non-marine mollusks
46
Plekocheilus (Eurytus) jucundus (Pfeiffer, 1855)
Antioquia: [“Antioquia. Neu Granada” (Breure, 1979)]
[“New Granada” (North Colombia), FMC 31067]
Plekocheilus (Eurytus) lamarckianus Pfeiffer, 1848
[Lectotype BMNH 1975295, “Andes of New Granada, 8000 ft.” (Breure, 1979)]
Plekocheilus (Eurytus) mabillei (Crosse, 1867)
[Holotype MNHN, “Andes de Colombie”, (Breure, 1975, 1979)]
Plekocheilus (Eurytus) nachiyacu Pilsbry, 1939 [described as n.sp. by Pilsbry, 1939] [Holotype
ANSP 170693a, “Colombia”, (Breure, 1979)]
Plekocheilus (Eurytus) oligostylus Pilsbry, 1939
[Type ANSP 170696, “Colombia”, Coll.:Brother Nicéforo María (Pilsbry, 1939)]
Plekocheilus (Eurytus) piperatoides (Pilsbry, 1901)
Magdalena: [Cerro Pintado, Sierra de Perijá, Coll.: Carriker, FMC 62362]
?: [Holotype, ANSP 263536 (Breure, 1979)]
Plekocheilus (Eurytus) pulicarius (Reeve, 1848)
[Lectotype BMNH 1975281, “New Granada” (Breure, 1979)
Boyacá: [Near Chiquinquirá, road Bogotá-Leiva, FMC 109720] [Sora, W of Tunja, Coll.:
M.Breistroffer, FMC 109721] [Salchica, S of Leiva, Coll.: 1949, FMC 109722] [N.W of
Sagamoso (=Sogamoso), DMNH 076788]
Cundinamarca: [Bogotá, Distrito Especial, FLMNH 109858] [Bogotá, FMC 413, 31061, 31075,
114077] [Bogotá, Coll. ?Fulton, CMC C10792-10794] [“Near Bogotá, Colombia Extinct ?”
(Abbott, 1989)]
Nariño: [Pasto, FLMNH 166223]
Plekocheilus (Eurytus) semperi Dohrn, 1882
Antioquia: [Sonson, (Breure, 1979)]
Plekocheilus (Eurytus) speciosus (Pfeiffer, 1855)
Magdalena?/Guajira?: [Lectotype, BMNH 1975300, Sierra Nevada de Santa Marta (Breure
1978)]
Magdalena: [Sierra Nevada de Santa Marta, Ecoandes Transect Alto Buritaca, 1100-1300m, Coll.
T.van der Hammen et al./Aug.1977, RMNH, ICNB (Breure, 1977b)]
Antioquia: [Támesis, DMNH 141728]
?: [“La Ceja”, DMNH 141723 (could be Antioquia, Huila, Bolívar, Magdalena or Sucre)]
Plekocheilus (Eurytus) subglandiformis (Mousson, 1873)
Distribution: Bogotá, Colombia (Simone, 2006)
Cundinamarca: [Lectotype ZMUZ 511633, “near Bogotá”, Coll.Wallis/1872 (Breure, 1976)]
Borrero, F.J. Colombian non-marine mollusks
47
[Simone has different information, so check!: Lectotype ZMUZ 233 (34 mm), ZMUZ 233 from
Frontina, Colombia (36 mm) (Simone, 2006)]
Plekocheilus (Eurytus) taquinensis Pfeiffer, 1855
Magdalena?/La Guajira?: [Lectotype BMNH 1957.6.3.1, Sierra Nevada de Santa Marta (Breure,
1979)]
Plekocheilus (Eurytus) tetensii (Dunker, 1875)
Magdalena: [Type?, Sierra Nevada de Santa Marta (Breure, 1979)] [Sierra Nevada de Santa
Marta, Coll.A.M.Cleef et al./Aug.1977, in Páramo vegetation (Calamagrostis sp. dominant)
(Breure, 1977b): Cuchilla Guïnue, 3000m (RMNH/1); Idem, Filo Alto La Cumbre, 3500m
(RMNH/1)] [Cerro Pintado, Sierra de Perijá, Coll.: Carriker, FMC 62363] [“Indian food”
(Abbott, 1989)]
Plekocheilus (Eurytus) virgatus (Pilsbry, 1935)
Boyacá: [Holotype ANSP 164573a, Valle de Tensa, Coll. M.Gonzalez, pre-1935 (Pilsbry, 1835;
Breure, 1979)] [(as Plectostylys virgatus (Pilsbry, 1935)) Macunal, DMNH 148705]
Cundinamarca: [P.aff. virgatus (Pilsbry, 1935), Salina de Sesquilé, E of Zipaquirá, NE of Bogotá,
FMC 109742] [La Calera, Bogotá (Abbott, 1989)]
Plekocheilus (Aeropictus) calliostoma (Dohrn, 1882)
Antioquia: [“provincia Antioquia, Novae Granadae (Breure, 1979)]
Boyacá: [10 km NE Villa de Leyva, (A.S.H.Breure Sta.121, RMNH (Breure, 1977a;1978)]
Cundinamarca: [Páramo de Palacio, 3 km S of Lagunas de Siecha, Coll.: J.Hernández/1958,
FMC 114111]
? Plekocheilus (Aeropictus) cathcartiae (Reeve, 1848) (?Colombia)
?: [Lectotype BMNH 1975288, “New Granada” (Breure, 1978)]
Plekocheilus (Aeropictus) delicatus (Pilsbry, 1935)
Cundinamarca: [(Found on Opuntia sp. cacti (Breure, 1977a), Soacha, 10 km SW Bogotá:
Holotype ANSP 164577a, Coll.V.Guevara, pre-1935 (Pilsbry, 1935; Breure, 1977a); Soacha
near Bogotá: Topotypes DMNH 148704; DMNH 076765] [near Mosquera, RMNH (Breure,
1978); FMC 86665] [Sabana de Bogata (=Bogotá), DMNH 076771] [Bogotá, Distrito Especial,
FLMNH 109987, FMC 125490, FMC 126937, 202079; DMNH 160230, 160232, 162165,
083709]
Boyacá: [Alto de Verea Suscon, 7 km NW of Sogamoso, FMC 78740]
Plekocheilus (Aeropictus) latilabris (Pfeiffer, 1855)
Cundinamarca: [Lectotype BMNH 1975127, Santa Fé de Bogotá (Breure, 1979)] [Plains of
Bogotá, Distrito Especial, FLMNH 109853]
?Plekocheilus (Aeropictus) quadricolor (Pfeiffer, 1847) (?Colombia)
[“Merida Prov., Colombia” (Abott & Dance, 1989) (may just be Venezuela)]
[Lectotype BMNH 1975283, “New Granada” (Breure, 1978)]
Borrero, F.J. Colombian non-marine mollusks
48
Plekocheilus (Aeropictus) succinoides (Petit, 1840)
Cundinamarca: [P. succineoides Petit, 1840, Syntype MNHN, “les environs de Bogotá” (Breure,
1979)] [Bogotá: FMC 414, 158889] [Bogotá, Coll.?Fulton, CMC C1108??] [(Breure, 1977a),
Coll.A.M. Cleef: Páramo Laguna Verde, 3600m (ZMA); Páramo de Palacio, Laguna de Stechas,
3750m (RMNH), Lagunas de Buitrago, 3600m (RMNH); Coll. H.Löscher: Finca Chueca near
Facatativá, ca.40km W Bogotá, 2700m (SMF 242140); Coll.H.Sturm: Páramo de Palacio, 31003200m (SMF 245384), Páramo de Monserrate (SMF 245385), Páramo between Monserrate and
La Calera, 3200m (SMF245386); Coll.T.van Der Hammen: Mountain side W of Laguna Muña,
SW of Bogotá, 260-2700m (ZMA); Bogotá: SMF 21611, 242139] [“Near Bogotá, Colombia.
Extinct?” (Abbott & Dance, 1989)] [Internet paper, Andes Orientales (Rodríguez, 1999)]
[Observed by A.M.Cleef at Páramo between Cogua and San Cayitano (San Cayetano?), around
Laguna Seca (Breure, 1977a)] [Sitio Tequendama I, Soacha (Correal & van der Hammen, 1977)]
Boyacá: [(Breure, 1977a), Coll.A.M. Cleef, at ZMA: Páramo de la Rusia, 3400-3600m; Páramo
Vado Hondo, 3600m] [Observed by A.M.Cleef at Páramo de Pizba, road from Socha to La Punta,
km.61 E of Los Pinos; km 78, 1km NE opf Laguna Colorada; Páramo de La Sarna, 5 km NE
Laguna de Tota (Breure, 1977a)]
Cauca: [Río Cauca valley, SMF 45635 (Breure, 1977a)
Plekocheilus (Aeropictus) succinoides cleeforum Breure, 1977
Cundinamarca: [Holotype ZMA, Páramo de Sumapaz, Cabeceras Rio Bogotá, Alto de Torquita,
ca.3900 m. (Breure, 1979)] [(Breure, 1977a), Coll.A.M. Cleef: Páramo de Sumapáz, Cuchilla
Rabona near Andabobos, 4000m (RMNH)]
Meta: [(Breure, 1977a), Coll.A.M. Cleef: Nevado de Sumapáz, 3800-3900m (ZMA)]
Plekocheilus (Aeropictus) veranyi (Pfeiffer, 1847)
Cundinamarca: [Sabana de Bogotá: Cerro de La Herrera, near Laguna de La Herrera on Opuntia,
2600m, Coll.:H.B.Bake/1955, FLMNH 184701; Laguna de Hernández, Coll.: F.Medem/1958,
FMC 114074; Mondeñedo, above Mosquera, Coll.: J.Carvajal/1957, FMC 114112]
?: [“Bulimus scytodes Pfeiffer 1853a, type locality Andes of Colombia is now considered a
junior subjective synonym of P.veranyi” (Breure, 1978)]
Plekocheilus scytodes Pfeiffer, 1853
?: [“Andes de Colombia” (Breure, 1979)]
Plekocheilus (Aeropictus) veranyi (Pfeiffer, 1847) (?Colombia)
?: [“Chachopo, Merida Prov., Colombia (Abbott,1989)(probably just Venezuela]
Plekocheilus (Aeropictus) zilchi Breure, 1977
Boyacá: [Holotype SMF 245387, SW Labranza Grande, 5°33’N, 72°35W), 1140m, Quebrada
Comojoque, Coll. H.Sturm (Breure, 1977a)]
??: [“Colombia”, BMNH 1975496]
Plekocheilus (Eudolichotis) bisuturalis (Pilsbry, 1895)
Norte de Santander: [as Auris distortus, Lectotype ANSP 25421a (Simone has it as Holotype
ANSP 25421 (42mm)) San José de Cúcuta (Breure, 1979)]
Cundinamarca: [as Auris bisuturalis bisuturalis (Pilsbry, 1896): Monteredondo, km 75 on road
Bogotá-Villavicencio, Coll.: P.I.Scheible & H.Frohmund, FMC 114169]
Borrero, F.J. Colombian non-marine mollusks
49
Eudolichotis distorta (Bruguiére, 1789)
Distribution: Trinidad, Venezuela, Colombia, Brazil (Simone, 2006)
Plekocheilus distorta gracilis (Pilsbry, 1896)
Norte de Santander: [Lectotype ANSP 25422, Cúcuta, Pamplona (Breure, 1979)]
?: [(as Auris bisuturalis Pilsbry), DMNH 075368] [“Andes Mountains”, DMNH 183930]
?Plekocheilus (Eudolichotis) dillwynianus (Pfeiffer, 1853) (?Colombia)
?: [“Lectotype BMNH 1975145, “Andes of New Grenada” (Breure, 1978)]
[Plekocheilus (Eudolichotis) dillwyanus Pfeiffer, 1853] Auris (E.) dillwyana spectrum (Albers)
?: [Type Locality: ‘“New Granada” (Colombia’ (Parodiz, 1962)
Plekocheilus (Eudolichotis) glabra Gmelin, 1791 (Breure, 1979)
Distribution: Trinidad, Brazil(?) (Simone, 2006).
[“Colombia”, FMC 31471]
?Plekocheilus (Eudolichotis) perdix Pfeiffer, 1848 (?Colombia)
[(Breure, 1979): Type locality “New Granada, Agua de Obispo” (Venezuela)]
?Plekocheilus (Eudolichotis) spectrum Albers, 1854 (?Colombia)
[(Breure, 1979): Type locality “Neu Granada”]
[Breure (1979) places Eudolichotis as a subgenus of Plekocheilus. However, in that
publication doesn’t list E. bisuturalis, unless is the same as Plekocheilus (Eudolichotis)
distorta bisuturalis (Pilsbry, 1896)]
Eudolichotis bisuturalis (Pilsbry) (Bruguière, 1789, Abbott, 1989)
Chocó: [“Arcandi” (Acandí ?), Coll.: F.Keep, FLMNH 109938]
Cundinamarca: [Monteredondo area: km 73 highway Bogotá-Villavicencio, FMC 84758; Coll.:
F.Medem,1956, FMC 114076; Coll.: P.I.Scheible/1959, FMC 114170]
Eudolichotis bisuturalis gracilis (Pilsbry)
Meta: [La Macarena, Camp.No.2, legos(?) de la orilla del Guayabero, Selva Platanillos after Las
Quemas, Coll.:1959, FMC 114168]
?: [“Colombia to Venezuela, on bananas” (Abbott, 1989)
Eudolichotis sp.
Cundinamarca: [Above Bogotá, Coll.: W.Weyrauch/1968, FMC 216976]
Plekocheilus victor (Pfeiffer) [Drymaeus ??]
Putumayo: [Puerto Asís, Upper Río Putumayo, Coll.: F.Medem/1958, FMC 114073]
Plekocheilus sp. [“Colombia”, Coll.:1898-1899, ANSP 78578]
Plekocheilus sp.
Magdalena/La Guajira (?): [Sierra Nevada de Santa Marta, Coll.:D.K.Dorman/1973, FLMNH
22264, 22265]
Borrero, F.J. Colombian non-marine mollusks
50
La Guajira: May not be the same species.[“Above Manuare” (Manaure ??), Coll.: A.A.Olsson,
FLMNH 184699]
Thaumastus (Thaumastus) tatutor (Jousseaume, 1887) (?Colombia)
?: [“New Grenada (=Colombia, Ecuador and Venezuela) (Breure, 1975)]
Family Orthalicidae
Subfamily ?
Sultana (Melaniorthalicus) adamsoni (Gray, 1833) Melaniorthalicus/Metorthalicus ????
Tolima: [Paramo near volcano Tolima (Pilsbry, 1899)]
Cundinamarca: [Bogotá & Valley of Guaduas (Pilsbry, 1899); Bogotá, Distrito Especial,
FLMNH 109378] [Metorthalicus adamsoni (Gray, 1833) Bogotá, FLMNH 166247]
[“Colombia”, FLMNH 109379]
[Are they the same genus/species ???]
Sultana (Melaniorthalicus) atramentarius (Pfeiffer, 1855) [Melaniorthalicus/Metorthalicus ????]
Cundinamarca: [Metorthalicus atramentarius (Pfeiffer, 1855), Bogotá, FLMNH 166249]
Santander del Norte: [Cúcuta & Chopa-Pamplona (Pilsbry, 1899)]
Santander: [FLMNH 166250] [~ 15km S of Málaga, Río Servitá valley, ca.1800m (A.S.H.Breure
Sta.184), at RMNH (Breure, 1977a)]
[“Colombia”, FLMNH 109380]
Metorthalicus approximatus (Fulton, 1896)
Cundinamarca: [Bogotá, FLMNH 166248]
Orthalicus (Melaniorthalicus) deliciosus, Strebel, 1909
Atlántico: [Barranquilla (Breure & Schouten, 1985)]
Orthalicus (Clathorthalicus) wallisi (Strebel, 1909)
Antioquia?: [West of Frontino, Cordillera de Las Juntas, 10000ft (Zilch, 1960; Breure &
Schouten, 1985)]
Orthalicus ferussaci Martens, 1863 [Distributed from Mexico to Ecuador, including Colombia
(refs. in Pérez & López, 2003)]
Trachyorthalicus fraseri (Pfeiffer) [“Colombia”, FLMNH 109359]
Stenostylus meleagris (Pfeiffer, 1853)
?: [Type locality: “Andes of Colombia” (Breure, 1978)]
Stenostylus nigrolimbatus (Pfeiffer, 1853) (?Colombia)
?: [Lectotype BMNH 1975549, “Andes, N.Granada” (Breure, 1978)]
[Breure (1979) does not list S. goudoti, nor any Stenostylus for which the original locality is
in Colombia. He places a question mark as to whether Stenostylus exists in Colombia. He
Borrero, F.J. Colombian non-marine mollusks
51
includes S. goudoti Petit within Scutalus (Suniellus), with different protoconch than
Drymaeus.
Scutalus (Suniellus) goudoti (Petit, 1843)
Tolima: [Mount Tolima, two Syntypes, one designated Lectotype, MNHN (Breure, 1975)]
Cundinamarca: [Bogotá, Distrito Especial, FLMNH 109250] [Scutalus goudoti (Petit,
1843),Villarica, FLMNH 161325]
?: [“Colombia”, FLMNH 109251]
Rhinus constrictus (Pfeiffer, 1841)
Distribution: NE Brazil, Venezuela (Simone 2006), Colombia?.
? : [“Mendez, Andes de Granada [Colombia]”(Breure, 1978)]
Simpulopsis (Eudioptus) citrinovitrea (Moricand, 1836)
Distribution: (As E. critrinovitreus (Moricand, 1836) Brazil (Simone, 2006). Colombia.
Valle del Cauca: [3 km W Atuncelo, 4500 ft., Coll.: F.G.Thompson/1979; FLMNH 178674]
[Pichindé, 5100 ft., Coll.: F.G.Thompson/1969, FLMNH 178675] [Restrepo et al./1983:
Hacienda “El Castillo”, Jamundí, UVM 021/1; La Paila UVM 014/1; Hacienda “Formosa”,
Ansermanuevo UVM 082-040/1]
Cauca: [Quilichao, IRSN/1 (Breure, 1978)]
Bulimus sowerbyi Pfeiffer, 1847 [Breure (1979) notes that this species cannot unambiguously be
assigned to one of the genera of Bulimulinae. Thus, while its family status seems clear, its
generic affiliation remains uncertain]
[“Andes of Colombia” (Breure, 1979)]
Bulimus wallisianus Mousson, 1873
Cundinamarca: [Bogotá (Breure & Schouten, 1985)
Bostryx bilineatus (Soweby, 1833b) [CHECK FAMILY/SUBFAMILY AFFILIATION]
?: [“known from Ecuador and Western Colombia” (Breure, 1879), CMC C11742 (no known
location)]
Bostryx scutulatus (Broderip, 1832)
?: [“Colombia”, DMNH 162164]
Family Odontostomidae
?Spixia spixi paraguayana (Ancey, 1892) [very possibly in error!]
Distribution: [As Spixia paraguayana (Ancey, 1892)], known only from Corumbá, Mato Grosso
do Sul, Brazil (Simone, 2006)
Cundinamarca: [Bogotá, Distrito Especial, Corumbá, FLMNH 109792. This is most likely a
Brazilian specimen labelled as Colombian in error]
Macrodontes tenuisculptus (Parodiz, 1962)
Cundinamarca: [(as Odontostomus fasciatus tenuisculptus (Parodiz, 1962): Holotype USNM
316130, ‘“Bogotá, Colombia”, possibly in error as the genus is only known from southern Brazil,
Borrero, F.J. Colombian non-marine mollusks
52
Paraguay and northern Argentina (Parodiz, 1962); Paratypes- USNM 635300 (34mm) (Simone,
2006)]
Ringincella carinata (Pfeifer, 1853)
?: [Syntype – BMNH (15mm), “Magdalena R.; Colombia” (Simone, 2006)]
Family Urocoptidae
Microceramus bonairensis (E.A.Smith, 1898)
Magdalena: [Ranchería, NE Magdalena, Coll.: A.A.Olsson, FMC 78840] [near Valledupar, FMC
117065] [Mogdalena (=Magdalena), DMNH 077317]
Microceramus concisus (Morelet, 1849)
San Andrés y Providencia: [San Andrés Isld. N.Cliff, Coll.: Pilsbry/1927, FMC 78843]
Microceramus sp.
San Andrés y Providencia: [Providencia Isld., Manchioneel Hill, basalt & limestone plantain
field, 100 m., Coll.: J.Geister/1970, FLMNH 180318]
Family Cerionidae
Cerion flamea (Maynard & Clapp, 1912) (introduced species? Only found once (Götting, 1978))
Magdalena: [Villa Concha, near Santa Marta, Coll.K.-J.Götting, 1976-78 (Götting, 1978)]
Cerion lewisi Clench 1961
San Andrés y Providencia: [Providencia Isld.: Back of Third Turtle Marina, Coll.:
H.Parkington/1981, FLMNH 48987; Airport, Coll.:D.G.Robinson/1982, FLMNH 49006]
Family Subulinidae
Rhodea barcrofti Pilsbry, 1958.
Cundinamarca/Meta: [Type, ANSP 211335, Monteredondo, km.73 on road from Bogotá to
Villavicencio, 1700m, Coll.F.Medem] [Monteredondo, FMC 216063]
Rhodea californica (Pfeiffer, 1846)
?: [Type, SMF 157250, Cauca-Tal, Columbien. Slg. Ehrmann ex Schlüter (Wenz, 1959; Zilch,
1973)]
Valle del Cauca: [Cordillera Occidental, locations along the road Cali-Buenaventura (Restrepo &
Breure, 1987)]
[“New Grenada” (Locality data degraded), ANSP 4345]
Rhodea pfeifferi Crosse [“Crosse renamed [R. californica] on account of the erroneous location
implied by the original name” (Pilsbry, 1906). Thus, R. pfeifferi Crosse is a synonym of R.
californica (Pfeiffer)]
?: [“Colombia”, Coll.:1898-1899, ANSP 78438; “Colombia, FLMNH 164920]
Rhodea gigantea Mousson, 1873
Borrero, F.J. Colombian non-marine mollusks
53
Antioquia: [Type, Sonson, Prov.Antioquia, Neugranada. Slg. Kobelt 1875 ex J.O.Semper ex
Wallis, SMF 227708 (Zilch, 1973)] [Frontino, DMNH 152856]
Valle del Cauca: [3 km.W Atuncelo, 1500 m, Coll.: F.G.Thompson/1969, FLMNH 36126]
[“Colombia”, Coll.: Brooklyn Museum, 1952, ANSP 189393; Coll.:Dohrn, FMC 36151;
FLMNH 110905] [New Grenada, FMC 416]
Rhodea wallisiana Dohrn
?: [“From Upper Magdalena Region” (Pilsbry,1958)]
Obeliscus haplostyla (Pfeiffer)
Cundinamarca: [Bogotá, FMC 36267]
Obeliscus major (Miller)
Cundinamarca: [Bogotá, FMC 36140]
Obeliscus (Ischnocion) triptyx Pilsbry, 1907
?: [cited as “known only from Colombia” when reporting it from Chanchamayo Valley, Peru
(Haas, 1951)] [Most likely same as Leptinaria triptyx Pilsbry from“New Grenada”, ANSP 24113]
Obeliscus sp.
Valle del Cauca: [Cordillera Occidental, locations along the road Cali-Buenaventura (Restrepo &
Breure, 1987)]
Beckianum beckianum (Pfeiffer, 1846)
Distribution: Mexico to Venezuela, West Indies, Guyana, Surinam, Peru, Brazil (Correa, 199;
refs. in Pérez & López, 2003; Simone, 2006).
?: [As Opeas (Synopeas) beckianum (Pfeiffer, 1846): “New Grenada”, ANSP 24212]
Lamellaxis gracilis (Hutton, 1834)
Distribution: (West Indies, Central America, tropical and subtropical South America (Simone,
2006).
As Opeas (Synopeas) gracile (Hutton, 1834):
San Andrés y Providencia: [Old Providence (=Providence Isld.): ridge north of High Peak; beach
south of Isabel, Coll. Pinchot Expedition, April 1929 (Pilsbry, 1930b] [St.Andrew Isld. (=San
Andrés Isld.), Coll. Pinchot Expedition, April 1929 (Pilsbry, 1930b]
Bolívar: [Cartagena, Coll.:F.Medem/1955-56, FMC 54749, 114108]
[Opeas gracilis Hutton, “New Grenada”, ANSP 24148]
Lamellaxis goodalli (Miller, 1822)
Distribution: Central and South Americas, Caribbean (Simone, 2006).
Opeas micrum (Orbigny, 1835)
Distribution: [As Allopeas micra (Orbigny, 1835)], Pan-American (Simone, 2006).
Bolívar: [Cartagena, Coll.:F.Medem/1955, FMC 54748]
Syntype – BMNH 1854.12.4.87 (6mm), Rio de Janeiro, Brazil (Simone, 2006).
Borrero, F.J. Colombian non-marine mollusks
54
San Andrés y Providencia: [Old Providence (=Providence Isld.): slope below the cleft of Split
Hill, ridge north of High Peak; near SouthWest Bay; Coll. Pinchot Expedition, April 1929
(Pilsbry, 1930b] [St.Andrew Isld. (=San Andrés Isld.), Coll. Pinchot Expedition, April 1929
(Pilsbry, 1930b; Coll., 1931, J.Zetek, FMC 86111]
Opeas pumilum (Pfeiffer, 1840) [Introduced European species]
San Andrés y Providencia: [St.Andrew Isld. (=San Andrés Isld.), Coll. Pinchot Expedition, April
1929 (Pilsbry, 1930b]
[Due to its extensive distribution in Central and South America (refs. in Pérez & López, 2003), it
probably occurs in the mainland as well]
Opeas pyrgula (Schmacker & Boettger)
San Andrés y Providencia: [San Andrés Isld., at base of limestone cliff on N end of island, NE
face (12°30’N-081°60W), Coll.:K.Emberton/1987, ANSP 368414] [Providencia Isld., halfway
up to cleft in Split Hill (13°30’N-081°30W), on andesite cobble in rock pile,
Coll.:K.Emberton/1987, ANSP 368420]
Leptinaria lamellata (Potiez & Michaud, 1938)
Distribution: West Indies, Colombia, Venezuela, Guyana, Brazil (Simone, 2006), plus Central
America, Ecuador (Pérez & López, 1993).
Magdalena: [Near Pueblito, E of Santa Marta, Coll. K.-J.Götting,1976-78 (Götting,1978)]
Leptinaria ritchiei Pilsbry, 1906
Distribution: Simone (2006) reports it as only from São Miguel, Alagoas, Brazil. Colombia (this
paper).
Bolívar: [Between Sincerín & San Cayetano, 1947, FMC 109685]
Leptinaria triptyx Pilsbry, 1907?
?: [“New Grenada”, ANSP 24113] [Most likely same as Obeliscus (Ischnocion) triptyx Pilsbry,
1907?, cited as “known only from Colombia” when reporting it from Chanchamayo Valley, Peru
(Haas, 1951)]
Subulina octona (Bruguière, 1792) [Introduced African (?) species]
Type: [Haiti, Dondon. Slg. O.Boettger ex Maltzan 1889, SMF 145454a]
Magdalena: [Santa Marta area, Coll. K.-J.Götting,1976-78, common under stones and in flower
pots (Götting,1978): Cerca de Minca, S.of Santa Marta; Punta de Betín]
Atlántico: [Barranquilla, Coll. K.-J.Götting,1976-78, common under stones and in flower pots
(Götting,1978)]
Bolívar: [Monte Cristo, Río Sinú, Coll.: L.Zanella, FMC 109712]
Caquetá: [La Tagua, Coll.: F.Medem/1958, FMC 114117]
Cundinamarca: [Apulo, W of Bogotá, 1947, FMC 109706]
Chocó: [Quibdó; Río Atrato (Götting,1978)]
Valle del Cauca: [Cali, Balneario Santa Rita, 1000 ft., Coll.: F.G.Thompson/1969, FLMNH
270851] [Cali, F.J Borrero’s collections, gardens and flower pots, often with eggs, 1979-82,
deposited at UVZ] [Buenaventura, Coll.: W.Weyrauch, FMC 52407]
Cauca: [(Götting,1978)]
Borrero, F.J. Colombian non-marine mollusks
55
San Andrés y Providencia: [St.Andrew Isld. (=San Andrés Isld.): Coll. Pinchot Expedition, April,
1929 (Pilsbry, 1930b; Coll.: J.Zetek, FMC 74012, 86122]
Synapterpes (Zoniferella) albobalteatus (Dunker)
Nariño: [Pasto (Zilch, 1959)]
Synapterpes (Chryserpes) amabilis Pilsbry 1906
Caldas: [Marmato (Zilch, 1959)]
Synapterpes cf. amabilis Pilsbry, 1906
Valle del Cauca: [Cordillera Occidental, locations along the road Cali-Buenaventura (Restrepo &
Breure, 1987)]
Synapterpes auratus (Pfeiffer, 1846)
Valle del Cauca: [Cordillera Occidental, locations along the road Cali-Buenaventura (Restrepo &
Breure, 1987)]
?: [S. auratus auratus Pilsbry, “N.Grenada”, Catalog.Jun/1908, ANSP 59346]
Synapterpes glandiniformis DaCosta
Cundinamarca: [Bogotá, Catalog.March/1909, ANSP 98107]
Synapterpes (Promoussonius) incertus (Mousson, 1873)
Cundinamarca: [Type, Bogota. Slg. C. Boettger, 1906, SMF 96531 (Zilch, 1959; 1973)]
Synapterpes (Promoussonius) pennelli Pilsbry, 1926b
Valle del Cauca?/Cauca?: [Type ANSP 132451, “mountains near Popayán (Cauca), or in the
Cordillera Occidental near La Cumbre (Valle), Coll. F.W.Pennell/1922 (Pilsbry, 1926b)]
Synapterpes promoussonius Crowler
Cauca: [Volcán Furace (=Puracé), Porayan (=Popayán) area, Coll.:1957, ANSP 209783]
Synapterpes pilsbryi
?: [“Columbia” (=Colombia), Coll.Fulton, CMC C11044, 11045]
Synapterpes sp.
Valle del Cauca: [Pichindé, 5100 ft., Coll.: F.G.Thompson,1969, FLMNH 270867] [3.0 km W of
Atuncelo, 4500 ft., Coll.: F.G.Thompson/1969, FLMNH 270868]
Family Spiraxidae (Oleacinidae [Pilsbry, 1935?]
Euglandina apolinari Pilsbry, 1935
Cundinamarca: [Type ANSP 164580, Eastern slope of the Cordillera of Bogotá (Pilsbry, 1935)]
Euglandina bogotensis (Da Costa)
Magdalena: [No.14, Finca La Granja, Cerro San Lorenzo, Coll.: Medem et al./1964, FMC
187611]
Borrero, F.J. Colombian non-marine mollusks
56
Bolívar: [Tolúviejo, ca. 25 min from Tolú towards Sincelejo, Coll.: Medem et al./1964, FMC
187574]
Meta: [region of Villavicencio, ANSP 164581, Coll.pre-1935 (Pilsbry, 1935)]
Cauca: [Las Tijeras, Moscopán region, Coll.: F.Medem/1961, FMC 114063]
Chocó: [Caño Unguía, Lower Río Atrato, in a vine (bejuco), selva, Coll.: F.Medem/1958, FMC
114064]
Córdoba: [Ayapel, Río San Jorge, Coll.: F.Medem & C.A. Velásquez/1963, FMC 187573]
Nariño: [Río Mataje, Pacific coast, Coll.: G.Dolmatoff/1962, FMC 187610]
Santander: [Finca near Río Conde, Girona, Coll.: F.Medem/1960, FMC 114065]
Valle del Cauca: [Between Buenaventura and the Upper Río Calima, Coll.: I. Cabrera/1959,
FMC 114062]
Euglandina callista Pilsbry & Clapp, 1902 [Glandina callista Pilsbry & Clapp, 1902]
Magdalena: [Santa Marta at Valparaíso in forest, 4500 ft., Coll., Mr.&Mrs.H.H.Smith, 1898
(Pilsbry & Clapp, 1902; Clapp, 1920)]
Euglandina cucutensis Pilsbry, 1939
Norte de Santander: [Type ANSP 168434, Cúcuta, Coll.: Brother Nicéforo María (Pilsbry, 1939)]
Euglandina cumingii (Beck, 1837)
Atlántico: [Puerto Colombia, Coll. Hebard & Rehn, pre-1926 (Pilsbry, 1926)]
Bolívar: [hill “La Popa”, near Cartagena, Coll. A.P.Brown, pre-1926 (Pilsbry, 1926)]
Euglandina (Euglandina) cylindrus Martens
?: [Oleacina cylindrus Martens, Columbia, S.A. (Tryon, 1885)] [in describing E.(E.) cylindrus
angusta as a new subspecies, Haas (1951) mentioned that “the nominate form (i.e., Euglandina
cylindrus Martens) from the Colombian Andes has only seven whorls...]
Euglandina isabella Pilsbry
?: [“(E. apolinari Pilsbry, 1935) is related to E. isabella Pils. from northwestern Colombia...”
(Pilsbry, 1935)]
[Oleacina lucida Strebel]
?: [New Granada (Tryon, 1885)]
Euglandina ornata Pfeiffer, 1846
Quindío?: [(Achatina ornata Pfeiffer, 1846), Quendeu (Quindío) mountains (Baker, 1925)]
?: [Oleacina ornata Pfeiffer, New Granada (Tryon, 1885)]
Euglandina plicatula (Pfeiffer) [Oleacina plicatula Pfeiffer]
?: [Andes, New Granada (Tryon, 1885)]
Euglandina striata (Müller, 1774)
Distribution: Surinam, Guyana, Brazil (Simone, 2006). Colombia (this paper).
Type of E. striatula (a synonym) – NNML [what is it?] (56mm).
Caldas: [Manizales, Coll.: Richter/1942, FMC 114061]
Borrero, F.J. Colombian non-marine mollusks
57
Cauca: [Isla Gorgona, near prison camp, plantain crops, Coll.: F.Medem/1961, FMC 114159]
Chocó: [Pangala, about 40 km N of Palestina, Río San Juán, Coll.: B.Malkin &
P.Burchard/1969-72, FMC 169397, 398, 174914; Quebrada Taparral, 15 km N of Palestina,
FMC 173015]
Valle del Cauca: [(Euglandina cf. striata) Cordillera Occidental, locations along the road CaliBuenaventura (Restrepo & Breure, 1987)] [Euglandina sp. (striata?) San Pablo Forest, Finca de
Gustavo Kattan, km 16 road Cali-Buenaventura, 1900 m. (3°29’N-76°38’W), Coll.:
F.J.Borrero/1980]
Euglandina dactylus (Broderip) [may be a synonym of E. striata Müller (Tryon, 1885)]
Chocó: [Acandí, FLMNH 105936]
Bolívar: [Near Cartagena, Coll. Ll.B.Smith, pre-1926 (Pilsbry, 1926)]
Nariño: [Tumaco Island, near the Colombia-Ecuador boundary (Pilsbry, 1926)]
Valle del Cauca: [Sonson (Pilsbry, 1926)]
Euglandina sp.
Magdalena: [Socorpa Mission, Finca San José, Sierra de Perijá, Coll.: B.Malkin/1968, FMC
159202, 1509204, 205, 187598]
Euglandina sp.
[Colombia, 52mm (Femorale, 2000-2003)]
Family Streptaxidae
Rectartemon colombianus Pilsbry, 1935
Meta: [Type and 2 paratypes, ANSP 164578, Region of Villavicencio, Coll.V.Guevara, pre-1935
(Pilsbry, 1935)]
Magdalena: [(Rectartemon aff.colombianus Pilsbry), Sierra de Perijá, 1300-1400m (9°45’N73°20’W), Coll. B.Malkin/Aug.1968, FMC 168006 (Solem, 1974)]
Streptaxis anomphalus (Pilsbry, 1935)
Meta: [Type ANSP 164579, Region of Villavicencio, Coll. V.Guevara, pre-1935 (Pilsbry, 1935)]
[Selva, Platanillos, after Las Quemas, Sierra La Macarena, Camp.2, ca.300-500 m from border of
[river]Guayabero, Coll.:1959, FMC 114166]
Cundinamarca: [Monteredondo, km 76 road Bogotá-Villavicencio, Coll.: P.I.Scheible, 1956,
FMC 114003, 004, 114091]
Streptaxis candei Petit, 1842 [= S. gibbosa Pfeiffer, 1859, same as S. candeana Pfr. (Baker,
1925)]
Distribution: Venezuela, Brazil (Simone, 2006).
Syntype – ANSP 23739 (Simone, 2006).
Atlántico: [Barranquilla, ANSP 23751 (Baker, 1925)]
Bolívar: [Cartagena (Baker, 1925)]
?: [(As Streptaxis candeanus Petit, 1842), New Granada (Tryon, 1885)]
?: [(As Streptaxis candeanus Petit, 1842), New Granada (Tryon, 1885)]
Streptaxis dacostae Gude
Borrero, F.J. Colombian non-marine mollusks
58
Cauca: [“Compared with S. dacostae Gude from Cauca, “this form (S. anomphalus Pilsbry, 1935)
differs by....” (Pilsbry, 1935)]
Streptaxis dunkeri (Pfeiffer, 1845)
Distribution: Only reported from Rio de Janeiro, Brazil (Simone, 2006)
?: [New Granada (Tryon, 1885)] [(As Streptaxis funcki (Pfeiffer), listed as synomym of S.
dunkeri Pfeiffer: “...reported from New Granada...” (Pilsbry, 1935), thus possibly Colombia
and/or Panama]
Streptaxis flora Pfeiffer
?: [Andes de Colombia (Tryon, 1885)]
Streptaxis gibbosus Pfeiffer
?: [New Granada (Tryon, 1885)]
Streptaxis subregularis Pfeiffer, 1845 [Simone (2006) has it as (Pfeiffer, 1846)]
Distribution: Only reported from Brazil (Simone, 2006).
Cundinamarca: [Bogotá (Pilsbry, 1935)]
Streptaxis suturalis Martens
?: [New Granada (Tryon, 1885)]
[Streptaxis thomasi Pfeiffer already in as Systrophia thomasi Pfeiffer]
Caldas: [Marmato, N. Granada (Tryon, 1885)]
Streptostele musaecola (Morelet, 1860)
Distribution: From Hausdorf & Medina (2003): Western Africa from Guinea to the Congo;
introduced in many countries including Pacific islands, Bermuda, Trinidad, northern South
America and the Netherlands.
Cundinamarca: [Finca Torreblanca near Silvania, 1560 m forest, 4°24’14”N-74°23”12”W, ZMH
2918 (Hausdorf & Medina, 2003)]
Valle del Cauca: [Club Los Farallones, Mpio. Cali, zona de Pance, 1100 m, dead, under rocks,
Coll. F.J. Borrero, June 2004, CMC C13275, C13276] [Club de Golf Los Andes, km. 35 via
Cali-Popayán, dead in leaf-litter in bambu stand, Coll. F.J. Borrero, 1 July 2005, CMC C13293]
Family Strophocheilidae
Strophocheilus pardalis Ferussac, 1821
?: [Coll.Webb, CMC C11035]
Strophocheilus pudicus (Müller, 1774) [Simone (2006) has it as 1773]
Distribution: Simone (2006) reports only Brazilian locations.
[Paratype of S. roseolabris Bequaert, 1948 (a synonym) – MZSP 3224 (52mm) (Simone, 2006)]
[“Colombia”, FLMNH 202986]
Strophochelus pulicarius (Reeve, 1848)
Borrero, F.J. Colombian non-marine mollusks
59
Cundinamarca: [Fusagasugá, Coll. V.Guevara, pre-1935 (Pilsbry, 1935)]
La Guajira: [Coll. B.Quevara, pre-1935 (Pilsbry, 1935)]
[“Colombia” (as Plekocheilus pulicarius Reeve, DMNH 184181; “New Granada”, ANSP 66400;
Coll.: Geale, ANSP 66423]
Strophocheilus thammianus Martyn [(Webb, 1936); “Columbia, 2 ½”, p.205]
Family Megalobulimidae
Megalobulimus auritus (Sowerby, 1838)
Distribution: Simone (2006) reports only Brazilian locations.
?: [As Strophocheilus auritus (Sowerby, 1838), “Colombia”, FMC 27566]
Megalobulimus capillaceus (Pfeiffer, 1855)
Distribution: Peru, Bolivia, Brazil, Paraguay (Simone, 2006).
?: [As Strophocheilus capillaceus Pfeiffer, 1855 (var seneri Jousseaume): U.S. of Colombia
(Pilsbry, 1899)]
Megalobulimus oblongus (Müller, 1774)
Distribution: South America, Western Andes (Simone, 2006).
Arauca: [Coll.: 1974, USNM 853417]
Cesar: [Los Ortizes, Becerril-Socorpa trail, about 10-20 km E of Becerril, Sierra de Perijá, Coll.:
B.Malkin/1969, FMC 163695-696, 173002]
Córdoba: [Caño Juy, Trib.of Río Sinú, near Tierra Alta, Coll.: C.A.Velásquez/1965, FMC
187289]
Cundinamarca: [Villeta, Coll.: J.I.Idrobo, 1961, FMC 114127] [Puerto Salgar, Río Magdalena,
Coll.:H.Dybas/1938, FMC 3376] [Puente de Pandi: 779 m (Pilsbry, 1899); Fusagasugá (Götting,
1978)] [Anapoima-Cuno, Coll. J.A.Ramos, 14Aug/1965, DMNH 066793]
Boyacá: [Muzo: 600-800 m (Pilsbry, 1899); Coll. J.Bequaert/1936, DMNH 128717; (Götting,
1978)]
Tolima: [Ibagué (Götting, 1978)]
Magdalena: [“around Santa Marta and Río Manzanares, Coll.: H.N.Holland/1902, FMC 13588]
[Sierra Nevada de Santa Marta: Pueblo Bello, Coll.: B.Malkin/1968, FMC 159212, 295241;
Pueblobello (southern slope of Sierra), Coll.K.-J.Götting, ~1976-78 (Götting, 1978)] [Socorpa
Mission trail, Becerril, Sierra de Perijá, Coll.: B.Malkin/1968, FMC 159203, 159213]
Meta: [La Macarena, Río Guayabero, Camp.No.2 Selva, Platanillos, Coll.:F.Medem, 1959, FMC
114128] [DMNH 075353]
Valle del Cauca: [Restrepo et al., 1983: Vivero El Vínculo, Buga, UVM 82-237/1; La Paila,
UVM 81-110/2, 81-111/1; Hacienda Formosa, Ansermanuevo UVM 82-060/1]
?: [Río Guaviare, FMC 114163] [Magdalena valley, 500 m (Pilsbry, 1899)] [“Andes of New
Granada, Columbia” CMC C10769]
Megalobulimus oblongus oblongus (Müller, 1774)
Bolívar: [Cartagena, FLMNH 102946]
Boyacá: [Sogamoso, FMC 166268] [Muzo (Boyacá ?), Coll.: J.Bequaert, FLMNH 102949]
Magdalena: [Minca, near Santa Marta, 2500 ft., Collec.: H.H.Smith, FLMNH 102944]
Norte de Santander: [Puerto Santo, NMWZ PU9599, PU9600]
Borrero, F.J. Colombian non-marine mollusks
60
Antioquia(?)/Santander (?): [Concepción , FLMNH 102952 (There is a location named
“Concepción” in both Departments)]
?: [Colombia-Mabi, FLMNH 166269]
Megalobulimus oblongus perellongatus (Bequaert)
Cundinamarca: [La Vega, Coll.: J.Carvajal/1958, FMC 114051] [La Mesa (San Javier), Coll.:
Hernández/1955, FMC 114087]
Huila: [Terraza de Altamira, Coll.: P.F.Acosta/1960, FMC 114050]
Tolima: [Ortega, Coll.: P.E.Acosta/1960, FMC 114052]
?: [“Abajo de San José del Guaviare (Vaupés) Pto.Colombia (Meta). Coll.:1956, FMC 114086]
Megalobulimus popelarianus (Nyst, 1845)
Distribution: Ecuador, Peru, Colombia, Bolivia; Upper Amazon Basin (Simone, 2006).
Putumayo: [Puerto Asís area, Upper Río Putumayo: Coll.: F.Medem/1958, FMC 114094, 147;
Below Puerto Asís, Coll.: J.I.Idrobo/1957, FMC 114044, 114046] [Puerto Leguízamo, Upper
R.Putumayo, Coll.: F.Medem/1958, FMC 114045] [Headwaters of Río San Miguel, Santa Rosa
(Kofan Indian Village), Coll.: B.Malkin & P.Burchard, 1970-71, FMC 168199, 200187291-293]
[Woods near Santa Rosa de Kofanes, Middle course of Río Guamués, right shore, Coll.:
H.LeNestour/1965, FMC 187288]
Meta: [Selva, Platanillos, after Las Quemas, La Macarena, Camp.no.2 ca.300-500 m from border
of Guayabero, Coll.:1959, FMC 114145]
Antioquia: [“SW of town”, DMNH 113711]
?: [Finca “El Salado”, West margin of Upper Río Caquetá, below Finca “Curiplaya”, Coll.:
F.Medem/1958, FMC 114047] [Río Caquetá, Puerto Bay, Coll. 1958, FMC 114146]
[“Colombia-S.Pablo”, FLMNH 237924] [S. popelairianus M., “New Granada”, ANSP 82685]
Strophocheilus popelairianus popelairianus (Nyst, 1849)
Caldas: [Marmato, NMWZ PU 9611]
Strophocheilus popelairianus thamsianus (Martens, 1876)
?: [Upper Magdalena valley (Pilsbry, 1899)]
Strophocheilus popelairianus conectens (Martens, 1876)
?: [Upper course of Magdalena River (Pilsbry, 1899)]
Megalobulimus senezi (Jousseaume, 1884)
Distribution: Only reported from Colombia (Simone, 2006).
?: [Holotype – MNHN (80mm) (Simone, 2006)].
Family Scolodontidae
Hirtudiscus antioquiensis Hausdorf & Medina, 2006
Distribution: Known only from an Andean forest at 2700m altitude near La Ceja, Antioquia,
Colombia (Hausdorf & Medina, 2006).
Antioquia: [Holotype: La Ceja, Finca Guaycoral, Andean forest, 2,720m, 06º 05’20”N, 75º
23’35”W, ZMH 37577; diameter:5.7mm, height:2.4mm (Hausdorf & Medina, 2006)]
[Paratypes: La Ceja, Finca Guaycoral, Andean forest, 2,720m, UMNG-MT 303/4; ZMH
37578/1; 2,700m, UMNG-MT 304/2 (Hausdorf & Mdina, 2006)]
Hirtudiscus comatus Hausdorf, 2003
Borrero, F.J. Colombian non-marine mollusks
61
Cundinamarca (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Holotype ZMH 4359, La Calera,
0.5 km towards Bogotá, farm Las Cabreras, Vereda San Rafael, Andean forest, N slope, 2840m
04°42’30”N-73°58’20”W] [Paratypes: SMF 323673, Distrito Especial: Bogotá, brook above
calle 70-72 2700-2800m; SMF 323674, Bogotá, above Diagonal 72, 2700m; SMF 323675,
Bogotá, above Parque nacional, 2700-2800m; ZMH 4276, Santandercito, Finca el Ermitaño,
primary Andean forest with sandstone boulders, E. slope, 2310m; ICNB, ZMH 4019,4035,4067,
La Calera 0.5 km towards Bogotá, farm Las Cabreras, Vereda San Rafael, 2820-2840m; ZMH
4082, L Calera, S of Entrada Fábrica de Camento Samper, degraded Andean forest, NW slope,
2750m; ZMH 4296, Cachipay 5.5km towards Zipacón, small brook in forest with sandstone
boulders, 220m; SMF 323676, Tibatibá 5km towards Guasca; ZMH 4333, Bosques de Torca,
Andean forest, W slope, 2800m]
Boyacá (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Villa de Leiva: Oak forest near
Manantiales de Iguaque in Vereda Capilla I, W slope, 2600m, ZMH 4201; Cañón de Mamaramo
near Carrizal in Vereda La Capilla II, entrance to Santuario de Flora y Fauna Iguaque, Andean
forest, 2920m, 05°42’24”N-73°27’27”W, ZMH 4227]
Hirtudiscus boyacensis Hausdorf, 2003
Boyacá (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Holotype ZMH 4360, Villa de Leiva
5.8 km towards Gachantiva, dry forest, La Cabrera in Vereda La Sabana, 2170m, 05°40’31”N73°32’15”W] [Paratypes: ZMH 4213, same data; ZMH 4192,Villa de Leiva, 9 km towards
Arcabuco, disturbed oak forest near farm, 2400m; Moniquirá: ZMH 4127, 16.7 km towards
Arcabuco, primary Andean forest, W slope, 2500m; ICNB, ZMH 4166, 5.4 km towards
Arcabuco, disturbed forest rest near quarry Cantera El Porvenir in La Roca, 2150m; ZMH 4141,
13.2 km towards Arcabuco, waterfall above road, 2400m. SMF 323677, ICNB, ZMH 4184,
Barbosa, 5 km towards Arcabuco, forest near quarry El Cairo in Vereda Pueblo Viejo, 1840m]
Hirtudiscus curie Hausdorf, 2003
Cundinamarca (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Holotype ZMH 4361, Cachipay,
Finca El Porvenir, primary forest, 1600m, 04°43’45”N-74°25’45’W] [Paratypes: ICNB, ZMH
4316, La Mesa, 12 km towards Bogotá, Los Alpes near El Rosario, W slope with forest, 1700m;
ZMH 4305, Cachipay, Finca El Porvenit, primary forest, 1600m]
Hirtudiscus hirtus Hylton Scott, 1973
Cundinamarca: [Holotype, MLP 3965, Monte Redondo, Cordillera Oriental, entre Bogotá and
Villavicencio, 3800m]
Hirtudiscus triserialis Hausdorf & Medina, 2006
Distribution: Known only from Andean forests at 2300-2400m altitude in Parque Natural
Chicaque near Soacha, Cundinamarca, Colombia (Hausdorf & Medina, 2006).
Cundinamarca: [Holotype ZMH 37575, Parque Nacional Chicaque near Soacha, Andean Forest
near Cascada, 2400m, 04º 41’31”N, 74º 15’08”W (diameter:11.5mm, height: 4.3mm) (Hausdorf
& Medina, 2006); Paratypes: Andean forest near Cascada, 2400m, UMNG-MT 302/12; ZMH;
37576/4; Andean forest near river, 2300m, UMNG-MT 301/1 (Hausdorf & Medina, 2006)]
Systrophia eudiscus eudiscus (Baker, 1925)
Borrero, F.J. Colombian non-marine mollusks
62
Cesar: [Sierra de Perijá, El Rincón, E of Becerril, 200-300m, Coll. B.Malkin/Aug.1969, FMC
167951 (Solem, 1974), FMC 173000]
Systrophia helicycloides (Orbigny, 1835) [Simone 2006 has it as 1843]
Distribution: Bolivia, Brazil (Simone, 2006).
Syntype – BMNH 1854.12.4.104 (21mm) (Simone, 2006).
Amazonas: [Puerto Nariño, Coll.D.Avella/1858, FMC 114007; probably same record as Helix
(Anchistoma) helicycloides Orbigny, R. Maranon, Columbia (Tryon, 1887)]
Systrophia hondana (Pfeiffer, 1854)
Tolima: [“Honda”, FMC 78856] [as Zonites (Hyalina) hondana Pfeiffer: Honda, New Granada
(Tryon, 1886)]
Systrophia incara (Crawford, 1939)
Valle del Cauca: [6 mi NW Cali, Coll.: E.I.Schlinger & E.S.Ross/1955, FMC 106275]
Systrophia insigna (Orbigny)
Magdalena: [Santa Marta, El Líbano, 6000 ft., Coll.:H.H.Smith, FLMNH 107662]
Systrophia systropha (Albers, 1854)
Amazonas?: [“Upper Maranyon, Colombia” (Baker, 1925); probably same specimens as
“Colombia”, FMC 30514, and as Helix systropha, R.Maranon, Columbia, Tryon (1887)]
Systrophia thomasi (Pfeiffer, 1854) already in as Streptaxis thomasi Pfeiffer
Caldas: [Marmato, FMC 78855]
Systrophia sp.
Cesar: [Sierra de Perijá, El Roncón, 10-12 km E of Becerril, Coll.: B.Malkin/1969, FMC 173862]
Systrophia sp.
Putumayo: [Headwaters of Río San Miguel, Santa Rosa (Kofan Indian Village), Coll.: B.Malkin
& P.Burchard/1970, FMC 168182, 168185]
Happia ammonoceras (Pfeiffer, 1854)
Distribution: Colombia, Brazil (Simone, 2006).
?: [As Drepanostomella ammonoceras (Pfeiffer, 1854): New Grenada, ANSP 23776; “Santa Ana,
New Grenada” (Baker, 1925)]
Happia (Happiella) mediocris (Pfeiffer, 1854)
?: [“Santa Ana, New Granada” (Baker, 1925, citing Tryon’s (1886) Zonites (Hyalina) mediocris
Pfeiffer): “....Four localities with this name are shown on the maps... I suspect that it is actually
some snall town in the Cordillera Oriental near the boundary between Colombia and Venezuela”)
(Baker, 1926: “...I have found a Río Santa Ana near the boundary line of Colombia and
Venezuela (L 9°31’, 72°50’), which seems more probable than...”)]
Happia ponsonbyi ponsonbyi (Pilsbry & Clapp, 1902)
Borrero, F.J. Colombian non-marine mollusks
63
Magdalena: [Sierra Nevada de Santa Marta, 50 km SE Sta.Marta, just W of Río Buritaca along
trail to Ciudad Perdida, 650 m elev., Coll.:D.K.Dorman/1989, FLMNH 137921] [H. posonbyi
(Pilsbry & Clapp, 1902), same gen.loc, 600m, ridge on W bank of R.Buritaca, 1989, FLMNH
137925]
Happia sp.
Magdalena: [Coll: K.-J.Götting, 1976-78 (Götting, 1978): Cerca de Pueblito; San Pedro de la
Sierra]
Drepanostomella excisa (Pfeiffer, 1854)
Caldas: [excisa (Pfeiffer, 1854): Marmato, Coll.: T.Bland, FMC 78864] [Marmato, New Granada,
Coll. T.Bland, ANSP 23774]
?: [as Helix (Patula) excisa Pfeiffer: Santa Ana, New Granada (Tryon, 1887)]
Drepanostomella sp.
Valle del Cauca: [Carmelo, 1000 ft., Coll.: F.G.Thompson/1969, FLMNH 270920]
Family Systrophiidae
Guestieria blandi Pilsbry, 1930c
Caldas: [Type ANSP 23770, Quebrada Arquia, Marmato, 5000ft on the Western Cordillera, Coll.
T.Bland/1851 (Pilsbry, 1930c)]
Guestieria powisiana (Pfeiffer, 1854)
Quindío: [“Mountains of Quendeo” (=Quindío), in the Cordillera Occidental, SW Cartago
(Pilsbry, 1930c)]
Cundinamarca: [Bogotá, FMC 30538]
Caldas: [Marmato (Tryon, 1885)]
Cauca: [Isla Gorgona, Plantain plantation by prison camp, under fallen trees, Coll.:
F.Medem/1961, FMC 114030]
Miradiscops ridicula Pilsbry, 1930
San Andrés y Providencia: [Type ANSP 150163, valley opening westward, north of the High
Peak, among leaves about half way up and at summit of the ridge, Coll.: Pinchot Expedition,
April 1929 (Pilsbry, 1930b] [Providencia Isld., Coll.:K.Emberton/1987: 50-100 ft. below short
ridge between Twin Pinnacles of High Peak, N side (13°30’N- 081°30’W), in wet palmetto litter
in high ferns, ANSP 368428 A12127D, ANSP 368441; Next to small dry stream bed above path,
several hundred ft. below Spansih Saddle on trail from bottom house to High Peak, under leaf
litter beside rocks in 45° slope, ANSP A12127L; Hillside at W end of Manzanille
(=Manchioneal) Bay, S end of Island, virgin forest, leaf litter, ANSP 368438 A12128K]
Family Haplotrematidae
Haplotrema sp. [“New Grenada”, ANSP 23900, 23903]
Family Endodontidae (Family Charopidae FMC)
Borrero, F.J. Colombian non-marine mollusks
64
Radiodiscus mariae (Pilsbry, 1921)
Cundinamarca: [Bogotá, Coll.: Hermano A.Maria, FMC 84702; FMC 132464]
Helix gyroplatys Dohrn (Section Thysanota) [correct family, genus, subgenus?]
Antioquia: [Antioquia, N. Granada (Tryon, 1887)]
Family Polygyridae (?) [these records may belong with Sytrophidae]
Helix (Anchistoma) heligmoidea d’Orbigny (Section Polygyra) [correct family, genus, subgenus?]
?: [Guayaquil, Columbia (Tryon, 1887)]
Helix (Anchistoma) platygyra Albers (Section Polygyra, Subsection Polygyratia) [correct family,
genus, subgenus?]
Amazonas: [R. Maranon, Columbia (Tryon, 1887)]
Family Succineidae
Omalonyx sp.
Magdalena: [Sierra Nevada de Santa Marta, Coll.K.-J.Götting, 1976-78 (Götting, 1978)]
Succinea aequinoctialis (d’Orbigny)
Cauca: [Isla Gorgona, Coll.: F.Medem/1961, FMC 114066]
Succinea andecola (Crawford)
Cundinamarca: [near Bosa, SSW of and near Bogotá, Coll.: R.W.Barker, FMC 53046]
Succinea bogotensis Pfeiffer, 1866
Cundinamarca?: [Bogotá (Baker, 1925)]
??: [“New Grenada”, ANSP 23495]
Succinea cygnorum Pilsbry 1930
San Andrés y Providencia: [Old Providence (=Providencia Isld.), slope below cleft of Split Hill
and valley north of High Peak, Coll. Pinchot Expedition, April 1929 (Pilsbry, 1930b]
Succinea repanda (Pfeiffer, 1854)
Cundinamarca: [Near Tequendama, S of Bogotá, FMC 53045]
Chocó: [Sautatá, Coll.: F.Medem, 1955, FMC 52433]
“Succineidae”
Cundinamarca: [“Mosquerra” (=Mosquera), desierto 10 km E Mosquera, Coll. J.A.Ramos,
20March/1965, DMNH 066958]
Family Sagdidae
Borrero, F.J. Colombian non-marine mollusks
65
Lacteoluna selenina (Gould, 1848)
San Andrés y Providencia: [Providencia Isld.: slope below cleft of Split Hill & valley north of
High Peak, Coll. Pinchot Expedition, April 1929 (Pilsbry, 1930b; Hillside at W end of
Manzanille (=Manchioneal) Bay, S end of Island, from near sea level to top of Ridge, virgin
forest, leaf litter and rocks (13°30’N-081°30’W), Coll.:K.Emberton/1987, ANSP 368440; conch
farm, Coll.: D.Addison/1991, FLMNH 194094] [San Andrés Isld.: Coll.Pinchot Expedition,
April 1929 (Pilsbry, 1930b; Coll.:D.R.Paulson, et al./1966, FLMNH 171003; Coll.: J.Zetek/1931,
FMC 71534, 74101; At base of limestone cliff on N end of island, on or near damp limestone
cobble nested in leaves at base of fig tree (12°30’N-081°60’W), Coll.:K.Emberton/1987, ANSP
368416 A12126C]
Family Milacidae
Milax gagates (Draparnaud, 1801) [Introduced European species]
Antioquia: [Site/Stat.No.75, Coll.:C.Carmona/1954, USNM 575072]
Cundinamarca: [Coll.:R.Ruppel/1957, USNM 576507] [Arid SW slope with sandstone rocks and
Opuntia (cacti) at shore of Laguna de La Herrera, disturbed (quarry, rubbish), 2610m,
04°41’19”N-74°15’57”W, Coll. B.Hausdorf et al., 2000 (Hausdorf, 2002)]
Boyacá (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Villa de Leiva 9 km towards Arcabuco,
disturbed oak forest near farm, 2400 m, 05°42’05”N-73°29’56”W]
Nariño: [Site/Stat.No.82,1955, Coll.:R.Ruppel, USNM 575075] [Site/Stat.No.81,1955, Coll.:
M.Benavides, USNM 575074]
Family Agriolimacidae
Deroceras laeve (Müller, 1774) [introduced European species; agricultural pest]
Magdalena: [San Sebastián de Rabago, Sierra Nevada de Santa Marta, Coll.: B.Malkin, 1968,
FMC 161096] [Socorpa Mission, Siera de Perijá, Coll.: B.Malkin, 1968, FMC 198645]
Cundinamarca (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Bogotá, Distrito Especial: Roadside and forest near brook above Escuela Militar de Caballería, 2790m 04°41’15”N-74°01’33”W]
[Arid S slope with sandstones and Opuntia near the Aso Gas Station SW of Mosquera, disturbed,
2680m; Arid SW slope with sandstone rocks and Opuntia at the shore of Laguna de La Herrera,
SW of Mosquera, disturbed (quarry, rubbish), 2610m; La Calera, 0.5 km. towards Bogotá, farm
Las Cabreras, Vereda San Rafael, sandstones at forest margin, 2820m] [(Simroth, 1910 in
Hausdorf, 2002): Alto de Sibaté, towards Choachí; Páramo near Fusagasugá; Monserrate]
Boyacá (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Villa de Leiva, Pozo de la Vieja, rocks
at the Río Chacón, near metereological station, 2130m]
Antioquia: [Camelia near Angelópolis (Simroth, 1914 in Hausdorf, 2002)]
Valle del Cauca: [Dagua (Malek, 1981 in Hausdorf, 2002)]
?: [“Colombia”, Coll.: M.Benavides,1953, USNM 575076]
Deroceras panormitanum (Lessona & Pollonera, 1882) [introduced species; serious agricultural
pest (Hausdorf, 2002)]
Cundinamarca (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Madrid, Finca Los Cerezos,
garden, 2600m 4°44’N-74°16’W; El Rosal, farm Holanda in Honduras, 2750m 04°54’34”N74°15’44”W]
Borrero, F.J. Colombian non-marine mollusks
66
Deroceras reticulatum (Müller, 1774) [introduced species (Hausdorf, 2002)]
Antioquia: [Coll.:R.Ruppel/1957, USNM 576504]
Cundinamarca: [Coll.: R.Ruppel/1957, USNM 576506] [(Coll. B.Hausdorf et al., 2000, Hausdorf,
2002): Bogotá, Distrito Especial, Calle 100, No.2-15E, rocks and small gorge, 2860m,
04°40’32”N-74°01’52”W] [(Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): Arid SW slope with
sandstone rocks and Opuntia (cacti) at shore of Laguna de La Herrera, SW of Mosquera, 2610m,
04°41’19”N-74°15’57”W; Madrid, Finca Los Cerezos, garden, 2600m 4°44’N-74°16’W; Farm
Holanda in Honduras, grassland, NE slope, 2900m; Subachoque towards radar station at El
Tablazo, grassland above water reservoir, N slope, 3250m]
Boyacá (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Arcabuco 21 km towards Tunja, Alto
del Sote, edge of Andean forest, 3100m, 05°42’05”N-73°29’56”W; Villa de Leiva, Cañón de
Maramano near Carrizal in Vereda La Capilla II, Entrance to Santuario de Flora y Fauna Iguaque,
edge of Andean forest, 2920m, 05°42’24”N-73°27’27”W]
Nariño: [Coll.:R.Ruppel/1957, USNM 576505]
Family Arionidae
Arion intermedius Normand, 1852 [introduced species (Hausdorf, 2002)]
Cundinamarca (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Santandercito, Finca El
Ermitaño, primary Andean forest with sandstone boulders, E slope, 2310m, 04°34’16”N74°20’10”W; El Rosal, Farm Holanda in Honduras, edge of Andean forest, NE slope, 2850m,
04°54’34”N-74°15’44”W; Farm Holanda in Honduras, disturbed Andean forest with bamboo,
NE slope, 2900m; Laguna Guatavita, edge of Andean forest, 3030m; Subachoque towards radar
station at El Tablazo, Andean forest rest above water reservoir, N.slope, 3250m, El Tablazo near
Subachoque, subpáramo near radar station, 3530m]
Boyacá (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Villa de Leiva: oak forest near
Manantiales de Iguaque in Vereda Capilla I, W slope, 2600 m, 05°41’15”N-73°28’53”W; 9 km
towards Arcabuco, farm, 2400m; Cañón de Maramano nearCarrizal in Vereda La Capilla II,
Entrance to santuario de Flora y Fauna Iguaque, Andean forest, 2900m] [Moniquirá 16.7 km
towards Arcabuco, edge of primary Andean forest, W slope, 2500m]
Arion sp.
Cundinamarca: [30 km W of Bogotá, Bosque Robles de Mosquera, under logs, Coll. J.A.
Ramos/Mar.1965, MCZ 323743]
Family Limacidae
Limax marginatus Muller, 1774
Antioquia: [Coll.:R.Ruppel/1957, USNM 576503]
Nariño: [Coll.: R.Ruppel/1957, USNM 576502]
[“Colombia”, Coll.: L.Posada/1955, USNM 575077]
Lehmannia valentiana (Férussac, 1822) [Introduced European species; agricultural pest]
Borrero, F.J. Colombian non-marine mollusks
67
Cundinamarca (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Arid SW slope with sandstone
rocks and Opuntia (cacti) at shore of Laguna de La Herrera, SW of Mosquera, 2610m,
04°41’19”N-74°15’57”W]
Boyacá (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Villa de Leiva 9 km towards Arcabuco,
disturbed oak forest near farm, 2400 m, 05°42’05”N-73°29’56”W]
Antioquia: [Sonsón (Waldén, 1961 in Hausdorf, 2002)]
Family Boettgerillidae
Boettgerilla pallens Simroth, 1912 [introduced species (Hausdorf, 2002)]
Cundinamarca (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Bogotá, Distrito Especial: Calle
100, No.2-15E, rocks and small gorge, 2860m, 04°40’32”N-74°01’52”W] [La Calera, 0.5 km
towards Bogotá, Vereda San Rafael, 2830m, Andean forest and forest margin; La Calera, S of
Entrada Fábrica de Cementos Samper, degraded Andean forest, NW slope, 2750m]
Family Punctidae
Paralaoma servilis (Suttleworth, 1852) [introduced species (Hausdorf, 2002)]
Cundinamarca (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Bogotá, Distrito Especial: Calle
100, No.2-15E, rocks and small gorge, 2860m, 04°40’32”N-74°01’52”W; Road-side and forest
near brook above Escuela Militar de Caballería, 2750-2790m; Bosques de Chicó] [Arid SW
slope with sandstone rocks and Opuntia (cacti) at shore of Laguna de La Herrera, SW of
Mosquera, 2610m; La Calera, 0.5 km towards Bogotá, Vereda San Rafael, 2830m, Andean forest
and forest margin; La Calera, S of Entrada Fábrica de Cementos Samper, degraded Andean
forest]
Family Pristilomatidae (Zonitidae ?? [Pilsbry, 1930])
Hawaiia minuscula (Binney, 1840) [introduced species (Hausdorf, 2002)]
San Andrés y Providencia: [H. minuscula neomexicana (Ckll. & Pilsbry), Old Providence
(=Providence Isld.), valley of the western slope north of High Peak, to the top of the ridge. Coll.:
Pinchot Expedition, April, 1929 (Pilsbry, 1930)] [St.Andrew Isld. (=San Andrés Isld., Coll.:
Pinchot Expedition, April, 1929 (Pilsbry, 1930b]
Boyacá (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Moniquirá, 5.4 km towards Arcabuco,
roadside near quarry Cantera el Porvenir in La Roca, 2150 m, 05°50’58”N-73°32’47”W]
Vitrea contracta (Westerlund, 1831) [introduced species (Hausdorf, 2002)]
Cundinamarca (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Bogotá, Distrito
Especial:Bogotá, Bosques de Chicó, 04°41’N-74°02’W] [La Calera, S of Entrada Fábrica de
Cementos Samper, degraded Andean forest, NW slope, 2750 m]
Zonites (Hyalina) insignis d’Orbigny [correct family, genus, subgenus?]
?: [Columbia (Tryon, 1886)]
Zonites (Hyalina) santanaensis Pfeiffer [correct family, genus, subgenus?]
?: [Santa Ana, New Granada (Tryon, 1886)]
Borrero, F.J. Colombian non-marine mollusks
68
Family Oxychilidae
Oxychilus alliarius (Miller, 1822) [introduced species (Hausdorf, 2002)]
Cundinamarca (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Bogotá, Distrito Especial: Calle
100, No.2-15E, rocks and small gorge, 2860m, 04°40’32”N-74°01’52”W; Road-side and forest
near brook above Escuela Militar de Caballería, 2750-2790m] [Arid SW slope with sandstone
rocks and Opuntia (cacti) at shore of Laguna de La Herrera, SW of Mosquera, 2610m,
04°41’19”N-74°15’57”W]
Boyacá (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Villa de Leiva 9 km towards Arcabuco,
disturbed oak forest near farm, 2400 m, 05°42’05”N-73°29’56”W]
Family Polygyridae
Giffordius corneliae (Pilsbry, 1930)
San Andrés y Providencia: [Old Providencia Isld.: Type ANSP 150864, Valley opening west
north of High Peak, Coll.H.A.Pilsbry, April 1929; Also, below the cleft of Split Hill; near
Southwest Bay] [Old Providencia Isld.:Valley next to High Peak, Coll.: Pinchot Expedition,
April 1929, FMC 72259; Coll. K.Emberton, 16-20Oct.1987, 13°30’N-081°30W: N.Slope of
peak just W of High Peak, About halfway up mountain from Lazy Hill Creek, a few ft below
Station 3, under leaft litter, ANSP A12126F; Top of Gulley between High Peak & next peak
west, , in dead fern fronds on ground, in a stand of ferns interspersed with palmetto, ANSP
368419; 50ft below short ridge between twin pinnacles of High Peak, on north side, in wet
plametto litter, high ferns, ANSP 368421 A12126J, 368427 A12127C; Along trail to High Peak
in Spanish Saddle, ca.100m beyond Boy Scout Camp, under dead palmetto fronds on ground,
ANSP 368425 A12127A; Ridge N of High Peak, ca ¼ way to next peak, under group of
palmettos, ANSP 368426 A12127B; Next to small stream bed above path, several hundred ft
below Spanish Saddle on tral from bottom house to High Peak, under leaf litter and beside rocks,
45° slope, ANSP A12128B]
Giffordius pinchoti (Pilsbry, 1930)
San Andrés y Providencia: [Type ANSP 150865, Old Providence Isld.: below the cleft of Split
Hill, western slope at 3 or 4 hundred feet above the sea; valley opening west, north of High Peak;
near Southwestern Bay. Pinchot Expedition, 1929, Coll. H.A.Pilsbry] [Old Providencia Isld.,
below cleft of Split Mountain, Coll.: H.Pilsbry/1929, FMC 72260; Providencia Islnd, Coll.
K.Emberton, 16-19Oct.1987, 13°30’N-081°30W: N slope of peak just west of High Peak, about
halfway up mountain from Lazy Hill Creek, a few hundred ft below Station 3, under leaf litter,
ANSP A12126E; Above station 2, under dead grass on ground in grassy area, ANSP A12126G;
Ca. 50 ft below short ridge between twin pinnacles of High Peak on North side, in wet palmetto
litter, high ferns, ANSP 368423]
Family Thysanophoridae
Thysanophora crinita (Fulton, 1917)
Bolívar: [(Trichodiscina crinita Fulton), Cartagena, FLMNH 103132]
??: [“Colombia” (Pilsbry, 1926)]
Borrero, F.J. Colombian non-marine mollusks
69
Thysanophora plagioptycha (Shuttleworth, 1854)
San Andrés y Providencia: [St.Andrew Isld. (=San Andrés Isld.), Coll.Pinchot Expedition, April
1929 (Pilsbry, 1930b]
?: [“Specimens are at hand from....the U.S.of Colombia (R.Swift coll.) (Pilsbry, 1920)]
Thysanophora santanensis (Pfeiffer, 1854)
?: [“Santa Ana, New Granada” (Baker, 1926: “...I have found a Río Santa Ana near the boundary
line of Colombia and Venezuela (L 9°31’, 72°50’), which seems more probable than...”)]
Thysanophora sp. [“New Granada”, ANSP 28271, 28272]
Miroconus sp.
San Andrés y Providencia: [Providencia Isld.: 50-100 ft. below short ridge between Twin
Pinnacles of High Peack, on N side (13°30’N-081°30’W), in wet palmetto litter in high ferns,
Coll.: K.Emberton/1987, ANSP A12127H]
Family Camaenidae
Solaropsis andicola Pfeiffer (? Colombia)
?: [“Eastern Cordilleras” (Pilsbry, 1889)]
Solaropsis gibboni (Pfeiffer, 1846)
Cundinamarca/Meta?: [Type ANSP 203477-478 (S.gibbona stictoraphe Pilsbry), Monteredondo
area: km 73 on Road Bogotá-Villavicencio, Coll.:F.Medem/1956; Coll.:1956, FMC 114125,
114005; Coll.:P.I. Scheible/1959, FMC 114101; km 75 on road Bogotá-Villavicerscio
(=Villavicencio), Coll.: P.I.Scheible & H.Frohmund/1961, FMC 114151; Monteredondo &
Manzanares, Coll.: P.I.Scheible/1960, FMC, 114102]
Cundinamarca: [Bogotá: Coll.: M.Smith, FLMNH 209170; CMC C10711, C10712]
Caldas: [“Marmato, etc., New Granada” (Pilsbry, 1889)] [Manizales, 1340 m, FLMNH 113815]
Meta: [El Mico, near Río Guejar, Biotopo: Humid Forest, Coll.: J.Idrobo, FMC 114152]
[“Bogotá”, FLMNH 113810, 113814]
Santander: [San Andrés, Quebrada de Pangote, 1400 m, Coll.: PE.Acosta/ June 26, 1960, FMC
115800]
Solaropsis gibboni gibboni (Pfeiffer)
Antioquia: [“Antioquia”, FLMNH 165512]
Solaropsis gibboni amori (Hidalgo, 1867).
Tolima: [Causon (??) de Honda, FLMNH 165513]
Solaropsis hians Pfeiffer
Caldas: [Marmato, Colombia (Pilsbry, 1889)]
(?) Solaropsis marmatensis Pfeiffer [same as S. gibboni from Marmato?]
Caldas: [Marmato, New Granada (Pilsbry, 1889)]
Solaropsis praestans (Pfeiffer, 1853)
Borrero, F.J. Colombian non-marine mollusks
70
Norte de Santander: [Ocaña, Coll.:Wallis, FLMNH 165718]
Solaropsis rosarium Pfeiffer 1849 [“New Granada”, ANSP 33043]
Psadara catenifera Pfeiffer, 1852
?: [(Helix catenifera Pfeiffer, 1852) “Andes, New Granada”, Coll. M.Bourcier, MCZ 222360,
Pilsbry (1889)] [“(P. pizarro Pilsbry, 1944 from Peru) belongs to the group of P. catenifera
(Pfeiffer) of Colombia....” (Pilsbry, 1944)]
Psadara colombiana Pilsbry, 1939
Norte de Santander?/Cundinamarca?: [Type ANSP 170698, “Santa Librada, Colombia”, Coll.:
Brother Nicéforo María (Pilsbry, 1939)] [Santa Librada is a town in N.de Santander, and a
Quebrada (creek) in Cundinamarca]
[Psadara monile] Solaropsis (Psadara) monile Broderip, 1832
Cauca ?/ Caldas?: [Salango and Marmato, Colombia (Pilsbry, 1889). Note that Salango is an
island in Ecuador]
[Psadara undata] Solaropsis (Psadara) undata Haas, 1966
Meta: [Holotype FMC 114100, N of coulee No.3 of Río Guayabero, in the direction of La
Macarena (circa 74°W-2°30’S), Coll.F.Medem, Jan.1957]
Isomeria awa Cuezzo, 2006
Distribution: Known only from Type Locality (Reserva Natural La Planada, Nariño).
Nariño: [Holotype AvH Unnumbered, Reserva Natural La Planada, Nariño, 1730m, 1º 50 N, 77º
43’ W, Coll. J.E.Orejuela & G.Cantillo; Paratypes: FML 14796 A, from type locality (Cuezzo,
2006)] [Reserva Natural La Planada, Hacienda Buenos Aires, UV 81-225 (1, 10 Aug 1981)]
Isomeria basidens (Mousson, 1873) (Separation of material into 2 subspecies after Solem, 1966)
Isomeria basidens basidens (Mousson, 1873)
Distribution: Known only from type collection (Solem, 1966) [But see range extension below].
Cundinamarca: [Holotype, Bogotá, Zurich (Solem, 1966)]
Valle del Cauca: [Parque Nacional Farallones de Cali, Corregimiento Pance, ~3000m: UV087 (1,
Coll. 5 Jan.1980); Sendero La Cuchilla, UV086 (1, Coll.17 Aug.1980)]
Isomeria basidens gudeana (Ancey, 1904)
Distribution: Probably Central Cauca, Colombia (Solem, 1966) (but see Valle del Cauca record).
Valle del Cauca: [“La Nevera”, 12 km from Potreritos, road to Ataco, <100m, UV 83-048 (1,
Coll. R.Contreras 1 Feb 1983)]
Cauca: (Solem, 1966): [Popayán: ANSP 13249; Coll.: Pennell & Killip/1922, FMC 78831]
[Volcán de Puracé, Paletará, Coll.: R.Wright Barker, FMC 132553]
Isomeria bourcieri (Pfeiffer, 1853)
Distribution: Central Ecuador in Upper Esmeraldas, Napo and Pastaza Rivers (Solem, 1966).
[But see Colombian Records below]
Borrero, F.J. Colombian non-marine mollusks
71
Nariño: [Reserva La Planada, Ricaurte, 1770-1800m: UV81-235 (1, Coll.G.Cantillo 1 July 1981;
UV82-183 (1, Coll. G.Cantillo, 1 July 1982); UV99-147 (1, 1 June 1979); UV99-149, UV99-150
(1 each, Coll. C.Restrepo, 1 Sep 1987)]
Isomeria continua (Pfeiffer, 1854)
Distribution: Central Colombia between Manizales, Caldas and Villavicencio, Meta. Only
latitudinal data available: 3375-4390 feet (Solem, 1966).
Norte de Santander: [Province of Ocana (Ocaña) (Pilsbry, 1889)]
Cundinamarca: [Bogotá: FLMNH 208228; Edinburgh; SMF 172167; ANSP 63585, 45555,
ANSP 33134 (Types of I. peritropis Pilsbry, 1889, a synonym)]
Caldas: [Manzanares, 1340 m, MCZ 145206 (Solem, 1966)]
Meta/Cundinamarca: [(Solem, 1966): Guayabetal, 1030 m., MCZ 143365; Pipiral on road
Bogotá-Villavicencio (Jackson 6105)] [Region of Villavicencio, Coll.H.Apolinar María, pre1935 ANSP 164559 (Pilsbry, 1935)]
Isomeria fauna (Philippi, 1851) [Uncertain status; could be mislabeled I. kolbergi, as they differ
only in insignificant details (Solem, 1966)]
Distribution: Known only from the type locality (Solem, 1966).
Cundinamarca: [“Santa Fé de Bogotá, ZMA, Zurich (Solem, 1966)]
Isomeria fordiana (Pilsbry, 1889)
Distribution: Colombia: Colombia- One location each on the Cauca (Marmato, Caldas) and
Magdalena River Basins (?Ocana, Norte de Santander) (Solem, 1966).
Caldas: [Marmato, Types ANSP 33133 (Solem, 1966)]
Norte de Santander: [?Ocaña, Zurich (Solem, 1966)]
Valle del Cauca: [As Isomeria sp. cf fordiana (open umbilicus & too large a basal tooth): Lago
Calima, 1 km towards Campoalegre, UV81-48 (1, 1 Feb 1980)]
Isomeria globosa (Broderip, 1852) [=I. subcastanea (Pfeiffer)]
Distribution: Pacific lowlands from Tumaco Island, Nariño, Colombia south to Rio Jubones
basin, up to at least 3,500 feet at Paramba and Huigra, probably 7500 feet at San Jose de Minas
and Ibarra. Also from Mera, Topo and Napo on the Atlantic slopes (Solem, 1966).
Nariño: [Tumaco Area: Rio Rosario, E of Tumaco, FMC 133262 (Solem, 1966); Finca Vaquería,
on arm of Estero Agua Clara, <100m (problably less than 50m), UV-83-069 (1, Coll. H.von
Prahl 11 Sep 1983); Estero de La Varas, Finca Flor de la Brisa, Municipio Robles, 2 m, UV84067 (1, Coll. H.von Prahl 17 June, 1984)]
Cauca: [Quebrada Huanqui, Río Saija area, ca.100 m, Coll.: B.Malquin & A.Granja/1971, FMC
170629, 174891]
?: [“New Grenada”, ANSP 30645] [“S.W. Colombia; N.W. Ecuador” (Pilsbry, 1889]
Isomeria inexpectata Solem, 1966
Distribution: Previously known only from the type locality (Solem, 1966). The record from San
Lorenzo (Götting, 1978) extends the range by about 80 km northward.
Magdalena: [(Solem, 1966): Holotype USNM 599522 and Paratype FMC 132554, next to
Siminchucua River, between San Sebastián de Rábago and Mamancanáca, 8500-9000 ft,
Borrero, F.J. Colombian non-marine mollusks
72
headwaters of Rio Fundación, south slope of Sierra Nevada de Santa Marta, Coll.:
M.R.Carriker/1946] [San Lorenzo, 2600m (Götting, 1978). The photos of this specimen do not
show attachment of parietal lamella and parietal tooth, although the tooth is in the correct
position. May be a juvenile specimen, a variation of I. inexpectata, or a new species]
Isomeria jacksoni Solem, 1966 [?Colombia]
Distribution: Upper Esmeraldas and Amazonian drainages in Central Ecuador (Solem, 1966).
?: [“Mountains of Colombia and Ecuador” (Pilsbry, 1889)].
Isomeria juno (Pfeiffer, 1850) [?Colombia: probably not]
Distribution: Ecuador, sites near Rio Napo, Rio Pastaza, Mindo, Nanegal, and
Valle de Pilatón (Solem, 1966).
?: [“Andes of Colombia, Ecuador and Peru” (Pilsbry, 1889].
Isomeria kolbergi Miller, 1878
Distribution: Upper Río Esmeraldas drainage; Central Ecuador from Ibarra to Cayandeled;
Amazonian grainage from Río Quijos to Upper Río Pataza (Solem, 1966). Some records from
“Colombia”. Possibly I. fauna is a synonym (Solem, 1966).
?: [“Colombia”, CMC C10700, C10701] These are new records !!!
Isomeria medemi Solem, 1966
Distribution: Known only from type locality (Solem, 1966).
Magdalena: [(Solem, 1966): Holotype FMC 133261 and Paratype FMC 145160, Finca La
Granja, San Lorenzo Ridge, Sierra Nevada de Santa Marta, 8200-8850 ft, Coll.: C.A.Velásquez
& F.Medem, Feb. 19-25, 1964]
Isomeria minuta Solem, 1966
Distribution: Only known from a single specimen from the type locality (Solem, 1966).
Caldas?/Tolima: [(Solem, 1966): Holotype, SMF 172174, Sierra de Mariquita, 2170 m. Probably
this is a portion of Cordillera Central near de Caldas-Tolima border, above the town of Mariquita,
Tolima]
Isomeria neogranadensis (Pfeiffer, 1845)
Distribution: Known only from the type locality (Solem, 1966).
Caldas?/Tolima?: [(Solem, 1966): “Quendeu Mt”(=Quindío Mt.?; probably area of Central
Cordillera between Páramo de Barragán and Nevado de Quindío in the Caldas-Tolima border
area); “New Granada” (Zurich), probably a paratype]
Isomeria oreas (Koch, 1844)
Distribution: Colombia, Valle of the Rio Cauca and Cordillera Central from Frontino or
Angelopolis, Antioquia south to Popayán, Cauca. Also at Villavicencio, Meta, and “Ecuador”.
3,900-9,500 feet (Solem, 1966).
Valle del Cauca: [Type [unconfirmed?] MCZ 216984, Bitaco, Coll.: E.N. Kozloff, July 1954]
[Rio Paila Valley, 1800 m., Central Cordillera, USNM 251172 (Solem, 1966); Las Frias, Corr.
La Habana, Mpio. Buga, 1750m, UV84-063 (4, Coll. J.H.Restrepo 6 June 1984); Quebrada
Honda, near Hacienda El Paraíso, UV99-084 (1, Coll. C.Restrepo, M. Giraldo 1 Apr 1977);
Borrero, F.J. Colombian non-marine mollusks
73
Hacienda Chachafruto, Ver.La Morena, Corr.Galicia, Mpio.Bugalagrande, 1800m, UV99-093 (4,
Coll. V.Rojas 23-28 Feb 1999); Hacienda Los Cristales, Mpio. Sevilla, road Barragán-Sevilla,
2100m, UV99-105 (1, Coll. V.Rojas 20 Jan 1999); Finca la Esmeralda, Corr.San José, Mpio.
Obando, 1600m, UV99-110 (1, Coll. V.Rojas 16 Feb 1999)] [Miranda, UV060; Villa Carmelo,
1300m, 25 July 1980, UV074] [Pichindé, 5100 ft, Coll.: F.G.Thompson/1969, FLMNH 36101]
[3 km W Atuncelo, 4500 ft., Coll.: F.G.Thomson/1969, FLMNH 36102, 257926] [Cordillera
Occidental, road Cali-Buenaventura: Alto del Oso, Mpio. Restrepo, 2100m, Uv84-046 (1, Coll.
R.Neira 7 Mar 1984); Bosque San Antonio, km 14, 2140m, UV82-107 (1, Coll. I.Velasco/
R.Contreras 20 Mar 1982), UV82-149 (1, Coll. J.García 19 May 1982), UV82-180 (Coll.
C.Restrepo/ M.Giraldo 23 June 1982), UV82-200 (1, Coll. M.Giraldo 1 June 1982); Bosque San
Pablo, km 16, 1800m, UV82-070 (1, Coll. G.Kattan, 10 Nov 1981); Finca Zingara, Km 18, 4km
from Pavas, 1800-2000m, UV81-363 (1, Coll.M.Giraldo 28 Dec 1981), UV82-201 (1, Coll.
M.Giraldo, 1 June 1982); km 26, Parcelación El Ensueño, 1850m UV-no number (1); Quebrada
Los Indios, between Loboguerrero and Cisneros, UV81-256 (1, 1 Nov 1981)] [Palmaseca,
Mpio.Palmira, ~1000m, UV 83-066 (2, Coll. J.C.Cubillos 1 Jan 1966); El Arenillo, Mpio.
Palmira, UV84-025 (1, Coll. R.Neira 19 Mar 1984)] [Villa Colombia, Munic. Jamundí, UV 161
(1, 25 Jan 1981); UV81-292 (1, Coll. C.Murcia 13 Dec 1980)] [El Topacio, Corregimiento de
Pance: UV062; Coll. F.J. Borrero: CMC C13023 (1, 17 Jan 2006), C13024 (1, 15 Aug 2006),
C13283 (1, 30 July 2005), C13261 (1, March 2007), C13225 (4 juveniles, 18 Dec 2005)] [Lago
Calima, trail to Campoalegre: UV059, (1, 1Feb 1980), Finca Las Campanas (=Las Carmelitas),
1km between Lake and Campoalegre, 1600m, UV82-269 (2, Coll. E.Velasco. I need to check
them !)] [Vereda Portobello, Corr.Fenicia, Mpio. Riofrío, 1700m, UV-no number (2, Coll.
L.Neira 4 Mar 2004); Finca Lucitania, Mpio.Alcalá, near Río La Vieja UV-no number (1, Coll.
L.Neira Aug 2004)][Yotoco: 28 Oct 1980, UV064, 1 Nov.1980, UV065; Bosque de Yotoco
UV163 (1, 16 Dec 1980)] [Cali, city limits, UV164 (? specs. 1 Sep 1978)] [Municipio de Bolívar:
Finca La Cristalina, 3 km W of Corr. Betania, 2000m, UV84-110 (1, Coll. J.H.Restrepo/
C.Restrepo 15 Dec 1984); Finca El Manzano, 1.5km NE Betania, 1700m, UV84-113 (2, Coll.
J.H.Restrepo/ C.Restrepo, 16 Dec 1984)] [La Leona, Corr. Barragán, Tuluá-Morales Drainage,
UV99-006 (4, 26 Jan 1995)] [Bosque El Medio, Mpio. Zarzal, 990m, UV99-008 (1, Coll.
P.Silverstone 13 Jan 1986)]
Valle del Cauca/ Chocó: [Alto de Galápagos, road Ansermanuevo (Valle)- San Juán del Palmar
(Chocó), 2000m, UV85-027 (1, Coll. J.H.Restrepo 17 May 1985) First Chocó record!]
Cauca: [“Cauca”: ANSP 91219, Edinburgh (Solem, 1966); DMNH 083625] [Central Cordillera:
Rio Palo headwaters, USNM 251174 (Solem, 1966); El Silencio, Timba, Coll. 1 Jan 1979,
UV063; Cord. Occidental, Cuchilla del Timba, 2300m, UV81-188 (1, Coll. C.Murcia 28 Apr
1981)] [(Solem, 1966): Popayán, SMF; La Cumbre, near Popayán, ANSP 132448; Quilichao,
Edinburgh] [FLMNH 92427] [Coll. Fulton, CMC C11736] [Reserva El Guayabo, UV061] [El
Yarumal, Buenos Aires, 4 km on road to Esmeraldas, UV81-183 (1, Coll.R.Morales, 19 Apr
1981)]
Antioquia?: [Frontino, Colombia (Zilch, 1960)] [(From Solem, 1966): Brit.Mus., types of
procera Pfeiffer- Támesis (Jackson 6311)]
Cundinamarca: [?Bogotá, Zurich; Paris (Solem, 1966)] [Bogotá, FLMNH 159839]
Meta: [(Solem, 1966): ?Sonora, Zurich; Caño Grande, Villavicencio, MCZ 146366]
Risaralda: [Pereira, Carica, FLMNH 159837]
Antioquia: [(Solem, 1966): Támesis, Jackson 6311 (DMNH 141729); ?Frontino, Zurich]
Tolima: [Mt. Tolima, SMF (Solem, 1966)]
Borrero, F.J. Colombian non-marine mollusks
74
Caldas: [(Solem, 1966): Salamina, Jackson 6305 (DMNH 141727); Mt. Quindiu (=Quindío?),
Paris] [Marmato, ANSP 9272, 30642] [Calarcá, Coll.H.Apolinar María, pre-1935 ANSP 164560
(Pilsbry, 1935)] [Valle de Magdalena, 1200 m., MCZ 145205]
Huila: [Rio Páez Valley, 2900 m., Central Cordillera, USNM 251173 (Solem, 1966)]
Norte de Santander: [Ocaña, FMC 40636]
[“Colombia”: FMC 245, 40636; ANSP 30641; USNM 307678, 316112-316115; ZMA; Zurich]
[“New Grenada” DMNH 183750]
Isomeria scalena (Martens, 1881)
Distribution: La Plata Drainage of Huila, Colombia at 7550-7875 feet (Solem, 1966).
Huila: [Cerro Pelado, La Plata, New Granada, 2300 m (Pilsbry, 1889)] [(Solem, 1966): Rio Páez
Valley, 2400 m, USNM 251175; Hacienda Medersenberg, La Plata, SMF 172173, FMC 133152]
[Finca Merenberg: Santa Leticia, Municipio La Plata, 2300m, UV82-097 (1, 4 June 1982);
2300m, UV82-299 (1, Coll. G.Kattan, 1 Nov.1982)]
Isomeria subelliptica (Mousson, 1869)
Distribution: Cauca River Valley in Valle del Cauca, Colombia (Solem, 1966)
Valle del Cauca: (Solem, 1969): [gardens of Cali at 1100 m, USNM 251176] [Holotype Zurich,
“Bugua” (=Buga?), 4000 ft.]
Antioquia?: [Ocanna (Ocaña?), Zurich (Solem, 1966)]
Isomeria sp. new Species1 !!! [Looks like a Pleurodonte: Deep & lamella-like lower teeth as
palatal barriers; no other teeth, completely closed umbilicus; flared/deflected lip]
Boyacá: Finca El Bambú, Mpio. Puerto Boyacá, near Río Magdalena, <200m, UV82-029 (1,
Coll.H.vonPrahl 2 Oct 1978)]
Isomeria sp. new species2 !!! [1 basal tooth, no other teeth, umbilicus nearly closed. Need
better photographs of single, worn specimen missing spire]
Valle del Cauca: [Mpio. Buga, Road Buga-El Crisol, Río Guadalajara Basin, 1800-2200m,
UV85-011 (1, Coll. I.Cabrera, 23 Feb 1985)]
Isomeria sp. new species3 !!! [Large, 41mm diam; Large parietal tooth; very slight
engrossment of of lip in basal area (no tooth); no labial teeth; completely closed umbilicus;
lip edge brown-tinged, white see-thru band around periphery extending to edge of lip.]
Chocó: [Cerro del Torrá, Mpio. San José del Palmar (<2000 m ??), UV-no number (1, Coll.
R.T.Gonzalez Aug 7-Sep 5 1988)]
Labyrinthus aenigmus (Dohrn, 1875)
Distribution: Known only from type locality (Solem, 1066).
Antioquia: [Paratypes (Zurich) Frontino, at 6000 ft (Solem, 1966)]
[“New Granada”: Pleurodonte aenigma Dohrn, ANSP 30647; Solem (1966): MCZ 74568;
USNM 354647; Zurich] [“Colombia”, FLMNH 208145]
?Labyrinthus bifurcatus Deshayes, 1838 (Colombia?).
Borrero, F.J. Colombian non-marine mollusks
75
Distribution: Unknown range. Solem (1966) includes material from Ecuador and “New
Granada”, but not from Colombia.
“New Grenada”: [Edinburgh, Zurich (Solem, 1966)] (as L. bifurcata Deshayes), Coll.: Seymour,
ANSP 5330]
Labyrinthus clappi Pilsbry, 1901
Distribution: Colombia, Sierra Nevada de Santa Marta, probably at higher elevations only
(Solem, 1966).
Magdalena: [Sierra Nevada de Santa Marta: Alto del Cielo, about 5000 ft., in forest among
rotting leaves on the ground, Coll.:H.H.Smith (prior to 1901), ANSP 87392; “Santa Marta”,
FMC 40771, USNM 307449; CM 4091] [Don Amo State, 4000 ft ANSP 83168, CM 4530
(Solem, 1966)]
La Guajira: [Las Nubes State, Sierra Nevada de Santa Marta, 4000 ft, FLMNH 92433; CM
62.16585; USNM 171917; FMC 132086 (Solem, 1966)]
?: [“Colombia”, FLMNH 208213]
Labyrinthus creveauxianus (Ancey, 1890)
Distribution: The only definite locality is Frontino, Antioquia, Colombia (Solem, 1966).
However, a specimen at Carnegie Museum in Pittsburgh has a locality as “Panama”, suggesting
the species may be more widely distributed (CM 62.13580 Panama, as Labyrinthus uncigera
creveauxiana (Ancey)).
Antioquia: [Frontino: Basel 432a-Lectotype and Paratype; Edinburgh; Paratypes, FMC
132551 (Solem, 1966); CMC C11709]
[“Colombia”: ANSP 8915, MCZ, Zurich (Solem, 1966); FLMNH 208276]
Labyrinthus dunkeri (Pfeiffer, 1852)
Distribution: Colombia, from Norte de Santander to Guajira along the Sierra de Perijá, probably
in Estado Zulia, Venezuela (Solem, 1966).
Antioquia: [From Solem (1966): Frontino (Edinburgh); Sonson (?) (Zurich)]
Cesar: [Foothill of Sierra de Perijá, El Roncón, 10-25 km E of Becerril, 9° 42’ N-73° 11’W, 200300 m., Coll.: B.Malkin/1969, FMC 163698, 163700, 163704, 163709, 173867, 173871]
La Guajira: [5-6 km above Manuare (=Manaure), ANSP] [Hills south of Carraipia, USNM
599497]
Magdalena: [Sierra de Perijá: Western foothills at 200-500 meters, USNM 473933, FMC 117644;
Socorpa Mission, Finca San José, Coll.: B.Malkin/1968, FMC 159190 159192-159197; 12 km E
of Becerril, Finca Pedro Ávila, Coll.: B.Balkin & A.Granja, FMC 159191; Rio Maraca, ANSP
180017, FMC 117648; Airoca, 4000-6000 ft, USNM 488784, 488787] [Tierra Nueva, E of
Fonseca, 3700-5000 feet, Sierra Negra, USNM 533877] [La Palmita, trail from Loma Corredor
to Ocaña, ANSP 137825] [Near Codazzi, 600 m, USNM 599518] [Rio Cesar Valley, USNM
534075]
Norte de Santander: [Guamilito at 1500-2000 ft., USNM 517810]
Labyrinthus ellipsostomus (Pfeiffer, 1854)
Distribution: Upper Orinoco drainage in Colombia (Solem, 1966); Upper Amazonian Basin,
Brazil and Peru (Solem, 1966; Simone, 2006).
Holotype – BMNH 1966599 (24mm) (Simone, 2006).
Borrero, F.J. Colombian non-marine mollusks
76
Cundinamarca: [Santa Fé de Bogotá, ANSP 109259, FMC 127743, Edinburgh (Solem, 1966)]
Labyrinthus euclausus Beddome, 1908
Only known from one record, with uncertain locality (Solem, 1966).
Antioquia?: [(Beddome, 1908; Solem, 1966): Near Zaragoza at 800 ft in dense forest, Coll.:
E.Bell] [It could also be from the towns called by the same name in Cauca or Valle del Cauca]
Labyrinthus furcillatus (Hupé, 1853) (No Colombia material in Solem, 1966)
Distribution: Perú, San Martin and Huancavelica (Solem, 1966). Record below is uncertain, but
unique if confirmed.
[“Colombia”, FLMNH 208275]
? Labyrinthus leucodon (Pfeiffer, 1847) (?Colombia)
Distribution: Venezuela (Mérida, Yaracuy, Carabobo, and possibly Distrito Federal) (Solem,
1966).
?: A single record exists from “Colombia” (Zurich ex von Martens in 1854) (Solem, op.cit.),
which was not examined by this author.
Labyrinthus isodon (Pfeiffer, 1853)
Distribution: Known only from Ocaña, Norte de Santander, Colombia (Solem, 1966).
Norte de Santander: [Ocaña: ANSP 104823, FMC 127742, Zurich, Edinburgh (Solem, 1966);
FLMNH 208334] [Mt.Lucero (Zurich) (Solem, 1966. This author states that “Mt. Lucero” is
unknown, but it is probably either the Corregimiento of El Lucero, or the Filo El Lucero, both
close to one another in Norte de Santander)]
?: [“Western Colombia” (Pilsbry, 1889), although Norte de Santander is on Eastern Colombia]
Labyrinthus magdalenensis Solem, 1966
Distribution: Known only from Type locality (Solem, 1966).
Guajira: [Holotype USNM 533878, Tierra Nueva, E of Fonseca, 3700-5000 feet, Sierra Negra,
Coll.:M.R.Carriker; Paratypes: USNM 533878, FMC 117645 (Solem, 1966)]
Labyrinthus marmatensis Pilsbry, 1910
Distribution: Cauca River Valley in Colombia from Marmato to Angelopolis (Solem, 1966).
Caldas: [Marmato: Types of marmatensis, ANSP 8913, MCZ 154017 (Solem, 1966)]
Antioquia: [(from Solem, 1966): Jericó, USNM 424726; near Medellín, USNM 518435;
Angelópolis, Neuchatel (Types of angelopolites (Piaget, 1912))] [Jericó, Coll.15 Mar.1980,
UV072] [Angelópolis, near Titiribí, 1970 m (reference ?)]
?: [(Solem, 1966): Edinburgh, Zurich, FMC 132550]
Labyrinthus otis (Lightfoot, 1786) (Separation of material into 2 subspecies mostly after Solem,
1966)
Labyrinthus otis otis (Lightfoot, 1786)
Distribution: Colombia: Cundinamarca, Santander, Bolívar, Antioquia, Chocó (Solem, 1966).
Antioquia: [Turbo & León Valley to the south, Coll.: Haught, FMC 71045; USNM 543479]
[Golfo de Urabá: Region of Nicolí & Río Caimán, Punta Piedras, E shore of Gulf,
Coll.:G.R.Dolmatoff/1961, FMC 113925; R.Wright Walker (Basel 5432a) (Solem, 1966)]
Borrero, F.J. Colombian non-marine mollusks
77
Chocó: [Between Punta Ardita and Coredó, S.of Panamá, Coll.: G.R.Dolmatoff/1961, FMC
113888] [15 km inland, Rio Nuquí, USNM 488859]
Bolívar: [Quebrada Salvajín, Upper Rio Esmerelda, Upper Rio Sinú, USNM 601776] [Upper Rio
Uré, Rio San Jorge FMC 133263] [Limón, W of Montería, Rio Sinú, USNM 364339] [Road
between Rio San Jorge and Palmavirál, ANSP 160144, MCZ 95722] [La Burra (R.Wright
Barker)] [La Unión, FMC 117527] [Rio Canalete, ½ mile above La Unión, USNM 364338]
[Near Cartagena, ANSP 111651, CM 62.39790] [Beach of Cartagena-Bocachica (beach-worn
shell), Coll.K.-J.Götting, 1976-78 (Götting, 1978: reported as L. otis orthorrhinus, but the photos
seem closer to L. otis otis)]
Cundinamarca: [Monteredondo, km.76 carretera Bogotá-Villavicencio, Coll.: P.I.Scheible/1956,
FMC 113995; Coll.: L.Richter/1942, FMC 113996]
Santander: [45 miles SE of Barrancabermeja, USNM 341476]
Cauca (subspecies affiliation unknown, probably L. otis otis on the basis of its more southern
distribution): [Quebrada Huanqui, Río Saija area, Coll.: B.Malkin & A.Granja/1971, FMC
174893, 174895]
?: [“Colombia”, CMC C10788]
Labyrinthus otis orthorhinus (Pilsbry, 1910)
Distribution: Vecinity of Panamanian-Colombian border north to Golfo Dulce region of Costa
Rica. Possibly absent from Chiriquí and Bocas del Toro, the Costa Rican populations being
relicts (Solem, 1966).
Chocó: [Acandí: ANSP 150244 (Solem, 1966); FLMNH 92439] [Mountains near mouth of
Atrato River, USNM 206288 (Solem, 1966)] [Pangala, about 40 km N of Palestina, Río San Juán,
4°30’N-77°5’W, Coll.: B.Malkin/1972, FMC 174880]
Bolívar:
Labyrinthus otostomus otostomus (Pfeiffer, 1852)
Distribution: Unknown (Solem, 1966). Only record known below.
[Colombia, Edinburgh (Solem, 1966)]
Labyrinthus otostomus bogotensis (Pfeiffer, 1854)
Distribution: Known only from type locality, Santa Fé de Bogotá (Solem, 1966) [However, this
author also has records from Muzo, and there is an unconfirmed record from Meta (see below)].
Cundinamarca: [Santa Fé de Bogotá, ANSP 33168, SMF 82536, ZMA, Edinburgh, Paratypes
(Solem, 1966)]
Boyacá: [Muzo, ANSP 174437, MCZ 64931 (Solem, 1966)]
Meta: [“Bogotá”, FLMNH 92432]
[L. otostoma bogotensis (Reeve, 1854), “Colombia”, FLMNH 209387]
Labyrinthus otostomus assimilans E.A.Smith, 1897
Distribution: Known only from type location, Cauca River, Colombia (Solem, 1966).
?: [Types, “Cauca River, Colombia”, BMNH 97.7.26.7-8 (Solem, 1966)]
Huila: [Finca Meremberg, 2300m: 1 Aug.1979; UV085; 1 Dec.1979, UV068]
Labyrinthus plicatus (Born, 1780) (?Colombia)
Distribution: Venezuela: Aragua, Carabobo, Yaracuy, Falcón (Solem, 1966); Panama (Abbott,
1989).
Borrero, F.J. Colombian non-marine mollusks
78
Chocó?: [“Near the Atrato River, in the Sierra Darién”, Coll:”presented to the U.S.Nat.Museum
by A.E.Heighway” (Dall, 1909). Must find this specimen in USNM database!. Solem (1966)
considers this location within modern Panamá, but states the range of this species as restricted to
Venezuela]
?: [“Venezuela; Panamá” (Abbott, 1989) (Thus Colombia??)] [Thompson (1957) reports L.
plicatus from Rancho Grande & Turiano Pass, Venezuela, in leaf litter on the forest floor]
Labyrinthus raimondii (Philippi, 1867)
Distribution: Amazonian Basin, from Amazonas and Caquetá in Colombia to southern Loreto in
Peru and the Rio Tapajóz in Pará, Brazil (Solem, 1966; Simone, 2006).
Amazonas: [Caño La Duche, below the colony of Araracuara, FMC 114107]
Caquetá: [El Salado, below Curiplaya, Upper Río Caquetá, Coll.: F.Medem/1958, FMC 114143]
Putumayo: [Sta.Rosa de Sucumbios, Río San Miguel, Coll.:B.Malkin/1971, FMC 170617] [La
Tagua, SMF 162666] [Río Mecaya, tributary of Upper Río Caquetá, Coll.: H.Granados &
H.Arévalo, 1957, FMC 114000] [Las Lomitas, Río Sencella, affluent of Upper Caquetá,
Coll.:F.Medem/1958, FMC 114001] [Puerto Leguízamo, 1958, FMC 114002]
Labyrinthus sharmani (Gude, 1912)
Distribution: Only the Type specimen is known (Solem, 1966).
Antioquia: [Type BMNH 1912.1.27.1, Alejandría, 50 miles from Medellín, 5600 ft, Coll.:
T.P.Sharman (Gude, 1912; Solem, 1966)]
Labyrinthus sieversi (von Martens, 1889)
Distribution: Colombia, Sierra Nevada de Santa Marta, Magdalena, from at least 300-5500 feet
of elevation (Solem, 1966).
Magdalena: [Sierra Nevada de Santa Marta: Below Oblo, 100-300 m (von Martens (1889) in
Solem (1966); 50 km SE of Sta.Marta, near trail beside Río Buritaca, 550m elev., 11° 2’N73°55’W, Coll.:D.K.Dorman/1989, FLMNH 137929; San Miguel, 5500 feet, ANSP 140048;
Ecoandes Transect Alto Buritaca, 700 m, Coll. T.van der Hammen et al./Aug.1977, RMNH,
ICNB (Breure, 1977b)] [Los Gorros, old trail from Fonseca to Riohacha, northeast spur of Sierra
Nevada de Santa Marta, 2500 feet, USNM 488812] [Buritaca 200, Rio Buritaca Canyon, 1100m,
Coll. C. Restrepo, 15-18 July 1981: UV81-203 (2), UV81-204 (1)]
Labyrinthus subplanatus (Petit, 1843) (Separation of material into 2 subspecies mostly after
Solem, 1966)
Cauca: (subspecies affiliation unknown) [Quebrada Huanqui, Río Saija area,
Coll.B.Malquin/1971, FMC 175445]
Valle del Cauca: (subspecies affiliation unknown) [Chigüedó, arm of Río Cajambre, 10m,
UV83-097 (1, Coll. Y.Solarte 28 July 1983)]
Chocó: (subspecies affiliation unknown) [Quebrada Taparral, Río San Juán, 15-20 km N of
Palestina, Coll.: B.Malkin/1968-69, FMC 163705-163707; Coll.: B.Malkin & P.Burchard/197172, FMC 169396, 170624, 223565] [Pangala, about 40 km N of Palestina, Río San Juán, Coll.:
B.Malkin& P.Burchard/1971-72, FMC 170622-623, 223566] [Caño Docordo: Between
Cucurrupi & Noamana, Río San Juán, Coll.: B.Malkin & A.Granja/1968, FMC 173870, 173872;
ca. 110 km N of Palestina, Río San Juán 4° 55’N-76° 55’W, Coll.: B.Malkin & P.Burchard/1971,
FMC 223561-562]
Borrero, F.J. Colombian non-marine mollusks
79
Labyrinthus subplanatus subplanatus (Petit, 1843)
Distribution: Cordillera Oriental of Colombia in Santander and Cundinamarca; Cordillera
Central in Caldas. Possibly Sierra de Perija in Norte de Santander (Solem, 1966).
Norte de Santander: [(Solem, 1966): Ocaña, Zurich; Mt. Alto de la Cruz, Ocaña, Zurich; 13 S of
Puerto Santos, dense jungle, USNM 380794]
Santander: [Near Río Sucio at 600-800 ft, USNM 363924] [Río Lebrija Valley, 1000 ft, USNM
601777] [Rios San Juan de Dios and San Juan del Silencio, Carere, R.Wright Barker] [South part
of Santander: Coll.: Haught, FMC 71044; USNM 472800] [Landázuri at 830 m. MCZ 179595]
Caldas: [Victoria, 820 m, Coll.: G.R.Dolmatoff/1960, FMC 113997]
Labyrinthus subplanatus erectus (Mousson, 1873)
Distribution: Chocó and Valle del Cauca in Western Colombia, possibly Cundinamarca (Solem,
1966).
Cundinamarca: [Bogotá, Zurich-Types (Solem, 1966)]
Antioquia: [Frontino, Zurich (op.cit)]
Valle del Cauca: [Region between Buenaventura & Río Calima, Coll.: I.Cabrera, 1959, FMC
113999, 117525; Reserva Escalarete, UV-no number (1, Coll. G.Kattan 29Aug 1995)]
[Anchicayá, between Cali & Buenaventura: 300-500 m, SMF 162665; in front of Escuela El
Chidral, 980m, UV81-258 (1, Coll. R.Ríos 25 Feb 1981); “Bajo Anchicayá”, 800m, UV85-001
(1, Coll. F.J.Borrero 10 Apr 1980); near dam, UV 069 (1, 22 Dec 1979); km 89, UV071 (1,
Coll.R.Contreras, 18 Oct.1980); Estación Agroforestal (1, Coll.G.Cantillo 1 July 1981); Estación
Cartón de Colombia, UV83-114 (1, Coll. R.Contreras 1 Jan 1983), UV85-019 (1, Coll.
J.H.Restrepo 24 Mar 1985)] [La Brea, Calima, Municipio Buenaventura, MCZ 177962] [Rio
Cajambre: Chigüedó, arm of Río Cajambre, 10m, UV83-098 (1, Coll. J.Rojas, 1 Aug 1983),
Barco, 37m, UV83-100 (1, Coll. A.Gonzalez, 4 Aug 1983); El Piñuelal, 350m, UV83-107 (1,
Coll. R.Ríos 23 Aug 1983)] [Bahía Málaga, Naval Base, 0-30m, UV85-032 (3, Coll. R.Neira 31
Dec 1985)]
Chocó: [“Río Atrato”: R. Wright Walker; Coll.: Diepetrich, FMC 117526] [Rio Nuquí, 15 km
inland, USNM 488860] [Quebrada Docordo, Río San Juán, Coll.: B. Malkin/1969, FMC 173866]
Labyrinthus subplanatus sipunculatus (Forbes, 1850)
Distribution: Río Baudó and Río Nuquí, Chocó, Colombia north to Cerro Campana, Panamá
Province, Panamá (Solem, 1966).
Chocó: [Finca “La Victoria”, Boca del Pepé, Río Baudó, Coll.: F.Medem/1959, FMC 113994]
[Nuquí, in forest, USNM 596996] [Mountains near mouth of Atrato River, USNM 206289
(probably material from “near the Atrato River, in the Sierra Darién, Coll.A.E.Heighway”
mentioned in Dall (1909). Solem (1966) considers this location within modern Panamá)]
[Quebrada Docordo, Río San Juán, Coll.: B. Malkin/1969, FMC 168536] [Ensenada de Utría:
UV073 (1, Coll. G.Bolívar 1 Nov 1976); Forest around La Aguara Creek, UV81-178 (1, 14 Apr
1981), 5m, UV83-089 (2, Coll. G.Cantillo/ E.Velasco 17 Oct 1983)]
Borrero, F.J. Colombian non-marine mollusks
80
Labyrinthus unciger (Petit, 1838) (Separation of material into 3 forms mostly after Solem (1966).
Distribution (all 3 forms combined): Atlantic drainage of Panama from Costa Rica border to the
lower reaches of the Atrato River Basin in Colombia (Solem, 1966).
- Form undetermined:
Valle del Cauca: [Río Azul, Munic. El Darién, Río Calima Drainage, 560m, UV82-255 (3,
Coll. G.Cantillo, 18 Aug 1982). This is a new record for Valle del Cauca, regardless of
which form it is]
- Chocó: [Bahía Solano, Finca El Requinteadero, km 8 road to airport, 8m, UV83-093 (1, Coll.
E.Velasco 1 Nov 1983)] [Pizarro, 2.5 km from airport, UV-no number (1, 27 Aug 1996)]
- Form chiriquiensis has not been reported from Colombia
Chocó (form affiliation unknown): [Acandí, FLMNH 92451, 159879]
??: (form affiliation unknown): [New Grenada-Colombia, FMC 40760] [“Colombia”, ANSP
8915]
- L. unciger form unciger (Solem, 1966):
Chocó: [Arquía, 1 ½ hours on foot above Unguía, Lower Río Atrato, Coll.:F.Medem, 1958, FMC
113998, 114110] [Acandí, ANSP 164970] [Golfo de Urabá (R.Wright Barker) (Solem, 1966)]
[Mountains near mouth of Atrato River, USNM 111073 (Solem, 1966). This is the material from
Dall (1909), as Pleurodonte (L.) tenaculum Dall, 1909]
- L. unciger form tau (Solem, 1966):
Chocó: [Acandí, USNM 341768 (Solem, 1966)]
Labyrinthus vexans (Dohrn, 1875)
Distribution: Known only from the type collection (Solem, 1966).
Antioquia: [Lectotype, Cañas Gordas] [Frontino (Zurich) (Solem, 1966)]
[“Colombia”, FLMNH 209152] [“New Granada”: ANSP 63598, MCZ 79494]
Labyrinthus sp. new species [or Isomeria sp. new species ?]
(Just like Labyrinthus unciger, but no teeth at all)
Valle del Cauca: [Bajo Anchicayá: In front of El Chidral school, 980m, UV080 (2, Coll. 23
Nov.1979), UV81-259, UV81-267, UV81-269 (1 each, 25-26 Sep 1981); forest in from of dam,
UV079 (3, Coll. 1 May 1980)]
Family Helminthoglyptidae
Epiphragmophora pilsbryi Haas, 1934
Distribution: Only reported from Colombia (Cuezzo, 2006; Simone, 2006).
?: [Holotype – SMF 7752 (18mm) (Simone, 2006)].
Family Helicidae
Cornu aspersum (Müller, 1774) (=Helix aspersa (Muller, 1774) [Introduced European species;
commercially grown and “wild-ranging”; potential agricultural pest].
Cundinamarca (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Bogotá, Distrito Especial:
Universidad Militar Nueva Granada, 04°41’N-74°02’W] [Arid S slope with sandstone rocks and
Borrero, F.J. Colombian non-marine mollusks
81
Opuntia near Aso Gas Station SW of Mosquera, disturbed (rubbish), 2680m; Arid SW slope with
sandstone rocks and Opuntia at the Shore of Laguna de La Herrera SW of Mosquera, disturbed,
2610m]
Boyacá (Coll. B.Hausdorf et al., 2000, Hausdorf, 2002): [Villa de Leiva, market place, 5°38’N73°34’W]
Magdalena: [Bahía de Chengue, Parque Nacional Tayrona (Hausdorf, 2002)]
Leptarionta maxwellsmithi Pilsbry, 1930 (Family Helminthoglyptidae FMC, Family
Xanthonycinae ANSP)
Chocó: [Type ANSP 150243, Acandí, on leaves of bananas, Coll.: M.Smith,1950, (Pilsbry,
1930)]
Leptarionta venusta (Gude, 1926)
Chocó?: [Chirigu, FMC 146593]
Family Bradybaenidae (Xanthonychidae, FLMNH)
Hemitrochus gallopavonis (Pfeiffer)
San Andrés y Providencia: [Providencia Isld.: back of Third Turtle Marina, Coll.:
H.Parkington,1981, FLMNH 48988; Airport, Coll.:D.G.Robinson/1982, FLMNH 49007]
Plagioptycha bahamensis Pfeiffer
San Andrés y Providencia: [Providencia Isld.: Airport , Coll.:D.G.Robinson/1982, FLMNH
49008; conch farm, Coll.: D.Addison/1991, FLMNH 194098]
Trichodiscina coactiliata (Férussac)
Magdalena: [Cacagualito, 15 mi ESE of Santa Marta, Coll.:H.H.Smith, FLMNH 103130]
Family Helicarionidae
Habroconus lividus (Goilding)
Cundinamarca: [Above Bogotá, Coll.: F.Medem/1956, FMC 11084, 085]
Habroconus (Habroconus) selenkai (Pfeiffer, 1866) [?Colombia. Widely distributed in the
Americas (Baker, 1928 in Pérez & López, 2003)]
Habroconus semen-lini (Moricand)
Valle del Cauca: [6 mi NW of Cali, Coll.: E.I. Schlinger & E.S.Ross/1955, FMC 106274]
Habroconus sp.
Magdalena: [Sierra Nevada de Santa Marta: 50 km SE Sta.Marta, just W of Río Buritaca along
trail to Ciudad Perdida, 360 m elev., Coll.:D.K.Dorman/1989, FLMNH 137923; 750 m,1989,
FLMNH 137924; 550m, near trail beside Río Buritaca,1989, FLMNH 137928]
Habroconus sp.
Borrero, F.J. Colombian non-marine mollusks
82
Valle del Cauca: [Cali, Balneario Santa Rita, 1000 ft., Coll.: F.G.Thompson/1969, FLMNH
270850]
Habroconus sp.
Putumayo:[Headwaters of Río San Miguel, Santa Rosa (Kofan Indian Village), Coll.: B.Malkin
& P.BUrchard/1970, FMC 168181]
Guppya gundlachi (Pfeiffer, 1839) [?Colombia]
?: [Considering its extensive distribution in North, Central and South America (Venezuela,
Brazil)and the Caribbean (Correa, 1999), this species is likely present in the Colombia]
Family Ancylidae
Velletia fuhrmanni Piaget, 1914
?: [“Colombia”, Wurtz, 1951]
Ferrissia sp. (Roldán, 2001)
Gundlachia radiata (Güilding, 1828)
Distribution: USA, Mexico, Central America, Bolivia, Brazil (Simone, 2006) and Colombia.
Magdalena: [Río Sevilla and Boca de López, Ciénaga Grande de Santa Marta (Cosel, 1986)]
Magdalena/La Guajira: [Rivers Buritaca, Don Diego, Palomino, and other rivers East of Santa
Marta (Gotting in Cosel, 1986)].
Simone (2006) cites Hebetancylus providentialis Wurtz, 1951 as a synonym of G. radiata
(Guliding, 1828)]
San Andrés & Providencia: [(H. providentialis Wurtz, 1951): Type ANSP ?????, Old
Providencia Isld., Huffington Creek, Coll.C.B.Wurtz, 30April/1948]
Uncancylus concentricus (Orbigny, 1835)
Distribution: Central and South Americas (Simone, 2006)
Simone (2006) treats Hebetancylus culicoides (Orbigny, 1835) as a synonym of U. concentricus
(Orbigny, 1835).
?: [as H. culicoides: “Colombia: (Wurtz, 1951)]
Gundlachia sp. (Roldán, 2001)
Uncancylus sp. (Roldán, 2001)
FAMILY ??????? [ALL FROM PEALE’S COLOMBIA COLLECTION IN
MAGDALENA REGION, DESCRIBED BY ISAAC LEA (Abbott, 1955), including the
following holotypes:]
Bulimus pealianus
B. lacteus
B. corneus
B. columbianus
Caracolla hydiana
Borrero, F.J. Colombian non-marine mollusks
83
ANSP 199448 Bulimus malleatus Jay 1842 Type ?
Counts: dry: 3, alc: Original number: 984
Locality: New Grenada
Depth: Elevation: Station number:
Collector(s): Jay, J.C.
Date Collected:
Collection: Brooklyn Collection
Date Cataloged: 01 apr 1956
Family Ferussaciidae
Cecilioides (Karolus) consobrina (d’Orbigny, 1855) [Simone (2006) has it as C. consobrina
(Orbigny, 1841)
Distribution: Northen South America (Simone, 2006).
San Andrés y Providencia: [Old Providence (=Providence Isld.), ridge north of High Peak,
Coll.Pinchot Expedition, April 1929 (Pilsbry, 1930b] [Providencia Isld., 50-100 ft. below short
ridge between Twin Pinnacles of High Peak, on North side, 13°30’N-081°30W, Coll.:
K.Emberton/1987, ANSP A12127F] [St.Andrew Isld. (=San Andrés Isld., Coll.: Pinchot
Expedition, April, 1929 (Pilsbry, 1930b]
Family Oleacinidae
Pseudosubulina insularis Pilsbry, 1930
San Andrés y Providencia: [Type ANSP 150172, Old Providence (=Providencia Isld.): western
slope below the cleft of Split Hill; also in the valley north of High Peak. In woods among leaven
on the ground. Coll.: Pinchot Expedition, April 1929 (Pilsbry, 1930b] [Providencia Isld., Coll.:
K.Emberton/1987: Next to small dry stream bed above path, below Spanish Saddle on trail from
bottom house to High Peak (13°30’N-081°30W), under leaf liter, besides rocks and large twigs
on 45° slope, ANSP A12127K; Hillside at W end of Manzanille (=Manchioneal) Bay, S end of
Isld., from near Sea level to the top of the Ridge, Virgin forest, leaf litter and rocks, ANSP
A12128M]
Family Truncatellidae
Truncatella caribbaensis Reeve
San Andrés y Providencia: [Old Providence (=Providencia ), on beach debris a short
distance S of Isabel, Coll. Pinchot Expedition, April 1929 (Pilsbry, 1930b]
Truncatella bilabiata Pfeiffer
San Andrés y Providencia: [Old Providence (=Providencia), on beach debris a short
distance S of Isabel, Coll. Pinchot Expedition, April 1929 (Pilsbry, 1930b]
CLASS BIVALVIA
Systematic arrangement follows Cummins & Meyer (1997), Cummins (1999) and the SimpsonHaas Index (Graf & Cummins, 2004).
Family Hyriidae
Borrero, F.J. Colombian non-marine mollusks
84
Acostaea (Acostaea) rivolii (Deshayes, 1827)
Tolima: [Rio Opia, Coll. A.Dueñas/1995, CMC C11692] [Granados (1973) in Hawkins et al.
(1993): Magdalena River Basin, in Rios Cunday and Opía, and in the Hidroprado reservoir
(towns of Cunday, Prado, Piedras, Remolino, La Fragua, Caracolí, El Guadual and Doima),
generally attached to rocky substrata]
Santander: [“Santander”: Coll.: C.Cooke,1920, USNM 34471; Coll.: T. Link, USNM 363769]
Cundinamarca, Huila, Caldas, Santander: [Granados (1973) in Hawkins et al. (1993)]
?: [“Bogotá, Colombia”: Coll.: Fulton ,USNM 120889; “Bogotá, Colombia”, USNM 102027;
“Colombia”, OSUM 55648] [“....A. lobata Ferrussac (the type of the genus), which occurs in the
Republic of Colombia” (Marshall, 1927)] [Etheria labata (Férussac)
[“Magdalena River Drainage”, FLMNH 18250: Very possibly A. rivolii since Etheria is an
African genus of similarly attached mussels]
Diplodon (Diplodon) dunkerianus Lea, 1856
?: [(Unio dunkerianus Lea, 1856) USNM 84155, Holotype, “Colombia”, 2 valves,dry, Coll.: E.
Verreaux]
Diplodon (Diplodon) flucki Morrison, 1943
Vichada: [Paratype USNM 521999, Rio Orinoco at Maipures, Collec.: W.Fluck]
Diplodon sp. [Diplodon ???? / Diplodontites ????] NOT in SIMPSON-HAAS
?: [“Columbia”, D. graniferus Deshayes ANSP 327084, “Columbia”]
Diplodon sp.
Meta: [(Diplodon (Diplodon) losadae Haas, 1966 (Cummins, 1997 does not list it, even as a
synonym. In addition, Cummings, op.cit. states that there are “only 3 species of Diplodon in the
Orinoco Ricer drainage...., one undescribed, D. flucki Morrison, 1943, and D. granosus
(Bruguière, 1792)”) OMITTED in HAAS, 1969) Holotype FMC 114042/Paratype FMC 114043,
Caño Losada, Upper Río Guayabero, Orinoco System (circa 74°9’W-2°9’N), Coll.G.Dahl &
F.Medem, March 1959; Paratype FMC 114115, mouth of Caño Losada]
Diplodontites cookei Marshall, 1922
Distribution: Upper Amazon Basin, Colombia, Peru (Simone, 2006).
Santander: [Holotype USNM 341473 (53mm), Rio Colorado, Magdalena drainage system, Coll.:
C.Cooke; Paratype, USNM 341473 (65mm)]
Bolívar: [Quebrada Perro, Tributary of Rio San Jorge, Coll.: T Linke/1924, USNM 362862
(Marshall, 1926)]
Magdalena: [Coll.: O.Haught, USNM 534048] [Rio Maraca, near Becerril (Olsson & Wurtz,
1951)]
Diplodontites pilsbryana Olsson &Wurtz, 1951
Magdalena: [Holotype ANSP 187180a, Caño Las Animas, near Chiriguana, under 100 m (from
a fairly rapid stream with a bed of sand, 1973); Syntype USNM 187180 (60mm)]
Castalia multisulcata Hupé, 1857
Borrero, F.J. Colombian non-marine mollusks
85
Distribution: Venezuela, Guyana, Colombia, Brazil (Simone, 2006).
Syntypes – MNHN (42mm) (Simone, 2006)
[Johnson & Baker (1973) list Holotype ANSP 125553a (Rio Yuruari, Essequibo drainage,
Venezuela) as type of Tetraplodon stevensi H.B. Baker, 1930, which is a synonym of C.
multisulcata Hupé, 1857]
Bolívar: [As Tetraplodon linki (Marshall, 1926): Holotype USNM 362860, 2 valves, dry, Coll.:
T.A. Link, Sinu River at Lorica, near Gulf of Morrosquillo; Paratype USNM 3628861, 10
valves, dry, Coll: T.A.Linke, same locality]
Magdalena: [(As Castalia ecarinata Mousson, 1869): Type locality: Magdalena River, Puerto
Nuevo, Colombia (Cummings & Mayer, 1997); Coll.: O.Haught, USNM 473934]
Castalia orinocensis Morrison, 1943
Distribution: Orinoco River (Simone, 2006)
Vichada: [Holotype, USNM 522000?, Orinoco River at Maipures, Coll.: W.Fluck; Paratypes,
Maipures, Orinoco River, Coll.: G.K.Cherrie,1899, MCZ 167992 (Cummings & Mayer, 1997)]
Paxyodon syrmatophorus (Meuschen in Gonovius, 1781) [?Colombia]
Distribution: Amazon Basin (Simone, 2006).
?: [Rio Orinoco, ANSP 125546 (Cummins, 1999)]
Prisodon corrugatus (Lamarck, 1819)
Distribution: Amazon Basin (Simone, 2006).
Amazonas?/Caquetá?: [(As Prisodon (Triplodon) rugosissima savillei Olsson & Wurtz, 1951)
Holotype ANSP 187179a, “from the general Rio Mecaya area of the Caquetá Basin, Coll.:
C.W.Saville (Olsson & Wurtz, 1951; Jonson & Baker, 1973)] [As Triplodon (Triquetrana)
jamauchimensis (Baker, 1913), and Prisodon (Triplodon) amazonia (Frierson, 1914): Rio
Mecaya area of the Rio Caquetá, Coll.: C.W.Saville/1944, ANSP 187178 (Olsson & Wurtz,
1951)]
Prisodon obliquus (Schumacher, 1817) [? Colombia]
Distribution: Amazon Basin (Simone, 2006).
?: [Orinoco River, BMNH (76mm) (Simone, 2006)]
Family Mycetopodidae
Subfamily Anodontinae
Anodontites (Anodontites) carinatus carinatus (Dunker, 1858)
??: [“known ....from the Magdalena River System” (Haas, 1951)]
Norte de Santander: [(Anodontites palmeri Marshall, 1930). From Marshall (1930): Type
USNM 380695, 75 mmL, 42Hm 24Diam; Paratype USNM 380702, younger specimen; from
small stream, dense jungle, 13 km S of Puerto Santos, Coll.: R.H. Palmer; USNM 380695, 2
valve fragments, dry, Holotype, Coll.: R.Palmer]
Anodontites (Anodontites) elongatus (Swainson, 1823)
Distribution: Reported only from Amazon and Paraná Basins (Simone, 2006)
Valle del Cauca: [Holotype (Anodonta holtonis Lea, 1857) USNM 86519 , Near Cartago, New
Granada, 2 valves,dry, Coll.: J.Holton/1857 (Johnson & Baker, 1973)]
Borrero, F.J. Colombian non-marine mollusks
86
Anodontites (Anodontites) patagonicus (Lamarck, 1819)
Distribution: All South American Basins, west of Andes (Simone, 2006).
?: [As A. patagonicus patagonicus (Lamarck, 1819): “Colombia”, USNM 809472, Coll.:
G.Sarmiento,1980]
Anodontites (Anodontites) tortilis (Lea, 1852)
?: [(Anodonta tortilis Lea, 1852) Holotype USNM 86405, 2 valves,dry, Coll.: J. Trautwine,
“Colombia”]
Bolívar: [Coll.: T. Link/1924, USNM 364263]
La Guajira: [Coll.: A.Wetmore & M.Carriker, Jr./1941, USNM 533871]
Magdalena: [Coll.: O.Haught, USNM 473935]
Valle del Cauca: [Florida (nr.Palmira), Hacienda Atenas, 930 m, Coll.: G.C.Sarmiento, 1980,
FLMNH 29685]
Anodontites (Anodontites) trapesialis (Lamarck, 1819)
Distribution: All South American Basins, west of Andes (Simone, 2006).
Meta: [As A. trapesialis trapesialis (Lamarck, 1819): USNM 636121]
Vichada(?): [As A. trapezialis trapezialis (Lamarck, 1819), Metica River, Meta River, Orinoco
River, Coll.: J.D.Williams et al./1973, OSUM 33684]
?: [“Colombia”, Coll.: Gibbon, USNM 86610] [As Anodontites (Anodontites) trapesialis cf.
forbesianus (Lea, 1860)] [As A. trapesialis : “Colombia”, Coll.: Trautwine, USNM 86611]
Anodontites (A.) trapesialis glaucus (Valenciennes, 1833)
Bolívar: [“Cartagena”, Coll.: L.H.Strong, FLMNH 270533]Casanare(?): [A.trapezialis glauca
(Valenciennes, 1827), Río Tauto (lake), Río Magdalena, Coll.: F.Vidal/1982, 1983, OSUM
52966, 54084]
?: [As Pleiodon glauca Valenciennes, “New Grenada”, Coll.: I.Lea, ANSP 41892]
Anodontites (Anodontites) trigonus (Spix, 1827)
Distribution: Amazon and Paraná Basins, Patagonia (Simone, 2006).
?: [As Anodonta trautwiniana Lea, 1852, Holotype USNM 86684, Coll.: J.Trautwine, Cartagena,
Colombia, 1852; Paratypes USNM 86684 00A, and ANSP 41841]
Valle del Cauca: [As A. trautwiniana (Lea): Cali, in W Colombia, Coll.pre-1935 (Pilsbry, 1935)]
Antioquia: [Site/Sta.No.: 5B, USNM 517848]
Bolívar: [Coll.: M.Carriker, Jr./1947, USNM 590652]
Magdalena: [Coll.: O Haught/1944, USNM 543481]
?: [“Colombia”: [USNM 26100; Coll.: O.Hauht/1943, USNM 543460; As A. trautwiniana (Lea,
1852), OSUM 55650]
Anodontites (Ruganodontites) columbiensis Marshall, 1922
Santander: [Type & 3 Paratypes USNM 341472, Rio Colorado (tributary of Rio Magdalena),
Santander, Coll.: C.Cooke] [Rio Sucio, near Dos Bocas, Coll.: T.A.Link,USNM 365313]
[Barrancabermeja (Olsson & Wurtz, 1951]
Bolívar: [Coll.: M.Carriker, Jr./1948, USNM 590665]
Boyacá: [Coll.: O.Haught,1939, USNM 534056,]
Borrero, F.J. Colombian non-marine mollusks
87
Magdalena: [Coll.: C.Eigenmann, USNM 381553] [Rio Maraca, near Becerril (Olsson & Wurtz,
1951)] [Caño Las Animas, N of Chiriguana (Olsson & Wurtz, 1951)]
Valle del Cauca?: [“from Rio de la Paila, trib.of Upper Cauca River, Magdalena drainage”,
Coll.Eigenmann/1912, USNM 381553 (Marshall, 1930)]
Anodontites sp.
Meta: [Rio Meta, Rio Metica overflow pools at Hwy.bridge, 4.3 km NE Puerto López, Coll.:
J.Williams/1973, FLMNH 22109]
Subfamily Mycetopodinae
Mycetopoda siliquosa (Spix, 1827)
Distribution: Amazon, São Francisco and Paraná Basins (Simone, 2006).
Bolívar: [“Cartagena”, Coll.: W.D.Averill, FLMNH 270634]
Valle del Cauca: [Florida (nr.Palmira), Hacienda Atenas, 930 m, Coll.: G.C.Sarmiento, 1980]
?: [“Colombia”: Coll.:G. Sarmiento,1980, USNM 809471; Coll.: Totten & Trautwine, USNM
86785]
Mycetopodella falcata (Higgins, 1868)
Distribution: Upper Amazon River (Simone, 2006)
Holotype – BMNH 684436 (90mm), Paratype – USNM 86788 (95mm) (Simone, 2006).
Meta: [(Mycetopoda falcata Higgins, 1868), Upper Río Guayabero, Colombia, FLMNH 114133]
Chevronais colombiana Olsson &Wurtz, 1951 [Family/Subfamily ??]
Caquetá: [Holotype ANSP 187181, Rio Caquetá above Tres Esquinas (Olsson &Wurtz,1951;
Johnson & Baker, 1973)]
Nephronaias (Nephronaias) rowelli (Lea, 1859) [Family/Subfamily ??]
?: [(Unio rowelli Lea, 1859) “Colombia”, USNM 151555, Coll.: W.Cooper]
Haasica balzani (Ihering, 1893) [? Doubtful Colombian record]
Distribution: Paraguay River Basin (Simone, 2006).
?: [“Columbia rivers”, Freshwater shells Web site]
Family Sphaeriidae
Sphaerium forbesi (Philippi, 1869)
Distribution: Localities in Bolivia, Peru and Colombia (Kuiper & Hinz, 1084).
Cundinamarca: [Hacienda Portugal, Laguna de la Bersia, San Cayetano, 2830 m, ZMA/K 19504
(Kuiper & Hinz, 1984)]
Boyaca: [Alto Valle de Lagunillas, Sierra Nevada del Cocuy, 3950m (ZMA/K 19505) Kuipern &
Hinz, 1984)]
?: [“Tierras altas de Colombia” (Programa de biodiversidad, Chile)] [as Pisidium boliviense
(Philippi, 1869): “Colombia” Parada & Peredo (2002)]
Musculium (Clench, ????) [Clench, 1939]
Borrero, F.J. Colombian non-marine mollusks
88
Pisidium (Cyclocalyx) casertanum (Poli, 1795)
Distribution: Nearly cosmopolitan (Burch, 1972; Kuiper & Hinz, 1984)
Pisidium sp. (Roldán, 2001)
Eupera sp. (Roldán, 2001)
Sphaerium sp. (Roldán, 2001)
Family Dreissenidae
Mytilopsis leucophaeta (Conrad)
Magdalena: [Ciénaga Grande, 2 km NW of mouth of Río Sevilla, Coll.: F.G.Thompson/ 1990,
FLMNH 174344]
Mytilopsis sallei (Récluz, 1949)
La Guajira: [Riohacha, Coll.: A.A.Olsson, FLMNH 148060]
Magdalena/Atlántico/Bolívar: [Mytilopsis sp., probably M. sallei (Recluz): Ciénaga Grande,
Magdalena, near center, 10°50’N-74°23’W, Coll.:D.K.Dormar/1989, FLMNH 137909, 137915,
137918] [Attached by byssus to hard substrata in estuarine areas, as well as fresh and brackish
water lagoons and ponds along the road between Tolú (Morrosquillo Gulf, up to Riohacha
(Guajira). Also present in fully marine areas seasonally exposed to freshwater inflows in this
region (Cosel 1986); author’s own observations]
“Dreissenidae”
Nariño: “Nariño dept.”, DMNH 086823]
Family Corbiculidae
Polymesoda arctata (Deshayes, 1854)
Magdalena: [Ciénaga Grande, near center, 10° 50’N-74° 23’W, Coll.:D.K.Dormar/1989.
FLMNH 137914, 137917] [dead specimens common on beaches along the road between Santa
Marta and Riohacha (Guajira) (Cosel, 1978] [fresh and brackish water lagoons and ponds along
the road between Santa Marta and Barranquilla (Atlántico) (author’s own unpubl.obs.)]
Atlántico: [Mouth of Magdalena River (Cosel, 1986)] [Poza Verde, brackish, freshwater, and
changing salinity (author’s own unpubl.obs.)]
Polymesoda (Egeta) inflata Philippi, 1851
Pacific Coast of Colombia, including freshwater (deltaic) habitats.
? Cyanocyclas cuneata (Jonas, 1844) [? Colombia]
Distribution: Orinoco River, Venezuela (Simone, 2006).
Cyanocyclas isocardiodes (Deshayes, 1854) [Considered a doubtful species by Simone (2006)]
West Colombia, 58mm (Simone, 2006).
Cyanocyclas obscura (Prime, 1860) [Considered a doubtful species by Simone (2006)]
Borrero, F.J. Colombian non-marine mollusks
89
Colombia. Syntype – BMNH 1852.8.20.29 (49mm) (Simone, 2006)
Cyanocyclas regalis (Prime, 1865) [Considered a doubtful species by Simone (2006)]
Gulf of Uraba, Colombia. Holotype – MCZ 176894 (26mm).
Family Pisidiidae
Eupera sp.
Magdalena: [Rio Fundación, 1 km upstream from mouth at Ciénaga Grande, 10°43’N74°25.5’W, Coll.: D.K.Dorman/1989, FLMNH 137930]
XXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXXX
Acknowledgements
1- Dr. B. Hausdorf, Germany (references, discussion)
2- Cincinnati Museum Center: Steve Matter, Glenn Storrs, Jane MacKnight
3- Field Museum Nat.History Curator, Rüdiger Bieler and Jochen Gerber (info on Walter
Freeman Webb (collector): dates, places, taxa, whether samples were split, etc. Loan I.
inexpectata
4- G.T.Watters, OSUM
5- Leslie Skibinski, DMNH- doing database queries
6- Tim Pearce-Carnegie Museum-doing database queries/info on Rina Ramirez/access to
collection & host
6- Gary Rosenberg-ANSP (Suggestions where/how to publish)
7- Gene Coan- NHMLAC (Suggestions where/how to publish)
8- Janice Voltzow-AMS (Suggestions how to publish)
9- G. Kattan & Carolina Murcia, WCS-Cali – Hosts, field help.
10- Manuel Giraldo, UniValle, Cali. Access to Collection
11- Maria Gabriela Cuezzo, Tucumán, Argentina. Discussion, pictures, references
12- Hernando & Felix Montoya, Instituto del Medio Ambiente, El Topacio, Pance. Collection,
guidance, field help.
13- Ronald Janssen, Senkenberg. Loan I. inexpectata
14- Klaus Götting – Data & photographs of I. inexpectata
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[Pleurodonte (Labyrinthus) clappi n.sp., pl.II, figs. 3,4, from Sierra Nevada de Santa Marta][“it
is interesting that Labyrinthus is absent from southern Colombia and Ecuador, where the almost
toothless Isomeria occur”] [“It is most nearly allied to Helix sieversi v.Martens (Conch.Mittheil.,
iii,p.7, 1989; Man.Conch., viii, p.263.ix,pl.22.f.7,8, also from the Santa Marta mountains]
Pilsbry, H.A. 1901. A new Colombian Clausilia. The Nautilus, XV(4):39-40.
[Nenia smithiae, n.sp. from Sierra Nevada de Santa Marta]
Pilsbry, H.A. and G.H. Clapp.1902. New land shells from the Santa Marta Mountains, Colombia.
The Nautilus, XV(12):133-137, pl.vii
[Glandina callista, Circinaria ponsonbyi, Aperostoma sanctamarthae, Aperostoma smithi,
Helicina sanctamarthae, Helicina cacaguelita, all described as n.sp.]
Pilsbry, H.A. 1919. Nenia cooki N.SP. (Plate VII, Figs.11,12,13). The Nautilus, XXXII32(3):80-81.
[from the Peruvian Andes… “Dr. Paul Ehrmann has remarked of the genus Nenia that Ecuador
and northern Peru are its distribution center; the group here reaches its distribution acme of
differentiation, and is most numerous in species”]
Pilsbry, H.A. 1920. Some Auricularidae and Planorbidae from Panamá. The Nautilus,
XXXIII(3):76-79.
[Detracia zeteki, Phytia brevispira, Planorbis isthmicus from Panamá. Could they be in
Colombia ????]
Pilsbry, H.A. 1920. Review of the Thysanophora plagioptycha group. The Nautilus,
XXXIII(3):93-96.
Pilsbry, H.A. 1921. A Radiodiscus from Bogotá, Colombia. The Nautilus, XXXV(2):49.
[Radiodiscus mariae, n.sp., 1mm x 1.85mm, no photo.]
Pilsbry, H.A. 1926. The land mollusks of the Republic of Panama and the Canal Zone.
Proceed.Acad.Nat.Sci.Philadelphia, 78:57-126, pls.9,10.
Pilsbry, H.A. 1926b. South American Land and Freshwater Mollusks: Notes and descriptions,VI. Proceed.Acad.Nat.Sci.Philadelphia, 78:1-15, pls.I, II.
Pilsbry, H.A. 1930a A new Colombian Helicid snail of the Genus Leptarionta. The Nautilus,
43(4):116-117.
[Leptarionta maxwellsmithi, n.sp., from Acandí, Colombia, on leaves of bananas, Type
No.150243 ANSP]
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Pilsbry, H.A. 1930b. Results of the Pinchot South Sea Expedition, - I. Land mollusks of the
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Proceed.Acad.Nat.Sci.Philadelphia, 82:221-261, pls.15-19.
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Pilsbry, H.A. 1935. South American land and freshwater mollusks, IX- Colombian species.
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Pilsbry, H.A. 1939. South American land and freshwater mollusks, X- Species from Colombia
and Ecuador. Notulae Naturae, no.19, 7p.
Pilsbry, H.A. 1944a. Peruvian land Mollusca.- II. The Nautilus, 57(4):118-126 + pl.11
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Pilsbry, H.A. 1950. Freshwater mollusks from Colombia and Guatemala. The Nautilus, 63(3):8285, pl.5.
[Lithococcus venustus, n.sp., from Nariño, Colombia].
Pilsbry, H.A. & Axel A. Olsson. 1953. A Colombian Pomacea of the effusa Group. The Nautilus,
66(3):98-99, Pl.6
[Pomacea (effusa) oligista, new species from Cienaga de Luruaco (In the picture it appears with
a very low spire, almost as a Marisa)].
Pilsbry, H.A. 1956. Southern record for Pomacea cumingi King. The Nautilus, 69(4):140.
Pilsbry, H.A. 1957. Notes on land snails of Genera Solaropsis and Nenia. The Nautilus,
71(2):47-52, pl.3.
[mentions his work on Colombian Solaropsis (Pleurodontidae); Solaropsis gibboni, Nenia
(Andinia) barcrofti, n.sp. from Monteredondo, collected by Medem; N.karsteniana and N.
magistra from near Bogota are similar to N.barcrofti]
Pilsbry, H.A. 1958. A Colombian species of the Genus Rhodea. The Nautilus, 71(3):83-84, pl.9.
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ANSP 122975 Melania corvina Morelet
Counts: dry: 1, alc: Original number:
Locality: Columbia
Depth: Elevation: Station number:
Collector(s):
Date Collected:
Collection:
Date Cataloged: 28 Jun 1916
ANSP 63598 Helix vexans Dohrn
Counts: dry: 1, alc: Original number:
Locality: New Granada
Depth: Elevation: Station number:
Collector(s):
Date Collected:
Collection: Morelet Collection
Date Cataloged: 18 Jul 1893