General Biology 101 Chapter Outlines Starr & Taggert 10th edition Chapter 47: Community Interactions The introduction to the chapter hi-lights the concept of resource partitioning of pigeons found in New Guinea. Twelve species of pigeons each specializing on a certain food source co-exist because they have very finely defined resource preferences. Section 1 Which factors shape community structure? Habitat – A place characterized by physical and chemical features and by the array of species within it. Community – the populations of all species in the habitat that associate with one another. Factors that affect community structure. 1) Interactions among climate and topography 2) Kinds and amounts of food and other resources 3) Adaptations for survival 4) Interactions e.g. predation, competition etc. 5) Overall pattern of population size. Niche – the role/profession of a given organism in a community, sum of all activities and relationships to secure resources. Fundamental niche – the “ideal” situation without competition. Realized niche – the “actual” or “constrained” situation depending upon the prevailing conditions. Different types of species interations: 1) 2) 3) 4) 5) Commensalism* Mutualism* Competition Predation Parasitism* *Examples of symbiosis, which means “living together.” Table 47.1 demonstrates the positive and negative effects of these interactions. Section 47.2 Mutualism Mutualism – interactions between individuals of different species where each benefits by the interaction. Example: Pollination of flowers by their pollinators. Both benefit and neither is harmed. Obligatory mutualism – individuals of one speices cannot grow nad reproduce without the other species presence. “Intimate dependency on the other.” Examples: 1) Yucca and its yucca moth. 2) Mycorrhizae and roots of young plants 3) Endosymbiotic origin of eukaryotes (See Sec 20.4 for further details). Section 47.3 Competition – occurs whenever there are limited resources. Intraspecific competition – occurs between individuals of the same species. Interspecific competition – occurs between individuals of different species. This form tends to not be so intense as intraspecific competition. Types of competition interaction Exploitative – when a competitor has some selective advantage at using limiting resources. Interference – when access to resources is limited because of actions of one member involved in the competition. E.g. rufous hummingbirds outcompete broadtail hummingbirds in the Rocky Mountains because they tend to be more aggressive and have greater likelihood of procuring resources (of nectar). Competitive Exclusion – Theory that two or more species requiring identical resources cannot co-exist indefinitely. Demonstrated by Gause with an experiment of Paramecium (single celled organisms) in a flask of food (bacteria – their prey item). Where populations of two different populations coexist in nature, competitive interactions suppress the growth rate of the both of them. Demonstrated by Hairston’s studies with salamanders. Resource Partitioning – the subdividing of some category of similar resources that lets competing species coexist. > Species might co-exist in the same habitat even if their niches overlap. They are better able to do this if they share the resource in different ways or different times e.g. annual plants/weeds growing in a field. Section 47.4 predator-Prey Interactions Predators – consumers feeding on living organisms, includes carnivores and omnivores. Coevolution – the joint evolution of two or more species that exert selection pressure on each other as an outcome of close ecological interation. E.g. Crab and its prey the snails. Models of Predator–prey interactions: Type I: Each predator consumes a constant number of prey individuals over time regardless of the prey’s abundance. Graph is shaped like: / Type II: the consumption of prey increases with an increase in prey density, but not as fast as the increasing prey density. Graph is shaped like an asymptote. Type III: Predator response is slowest when prey density is lowest, similar to a logarhythmic curve. Factors that can affect predator/prey models/responses: 1) 2) 3) 4) Predator and prey reproductive rates. Hiding places for prey i.e. shelter Presence of alternative prey species Shifts in environmental conditions Stability depends on carrying capacity and population densities. Classic example: Canada lynx and prey of snowshoe hare. Their populations oscillate in synch with one another. What was found by Charles Krebs is that availability and abundance of plants also plays a significant role/factor i.e. a 3-fold model necessary to explain the observations was needed. Section 47.5 Evolutionary Arms Race Camouflage – some prey gain protection from predators by the adaptation of concealment e.g. living stone plants in the desert. Warning Coloration – conspicuous patterns on potential prey that warn the would be predators that they taste bad or are poisonous e.g. Wasp/bee colors, poison dart frogs, skunks. Mimicry – when prey species resemble dangerous or bad tasting species. Can be done with form, behavior or coloration e.g. beetles that resemble wasps but are in fact edible. Moment of Truth Defenses – when animals are cornered they may act defensively by snarling, growling, showing their bare teeth, spewing odors etc. to deter the would be predator. Predator Responses to Prey – in the evolutionary arms race predators counter prey defenses in a variety of ways with special adaptations such as: 1) Stealth (quickness and being sneaky) 2) Camouflage e.g. Scorpion fish laying on the sea floor. 3) Ways of avoiding repellents e.g. mice that behaviorally avoid spraying beetles by disabling their spray ability. Section 47.6 Evolution of Parasitism Parasites – live in (inhabit) or live in other organisms, benefit by gaining nutrients. Hosts – those organisms that carry parasites. It may or may not die from the interaction. Parastitic infections can lower the birth rate of the host, raise the death rate, and influence intraspecific and interspecific competition of the hosts. Kinds of Parasites Ectoparasites – live on a host’s body surface. Endoparasites – live within the body of the host. Microparasites – microscopic in size and fast reproducers. Macroparasites – include parasitic worms e.g. flatworms, mites, ticks, fleas. Holoparasitic plants – nonphotosynthetic plants that withdraw nutrient and water from young roots. Hemiparasitic plants – retain some ability to photosynthesize, but also gain nutrients from another plant that it resides upon e.g. mistletoe. Social parasites – alter the social behavior of another species to complete their life cycle e.g. cuckoos and north American cowbirds. Parasitoids – organisms that are part parasitic and part predatory e.g. larvae of insects that live inside and consume all of the soft tissues of the host. E.g. parasitoid wasps that lay their eggs on larvae of sawflies. Parasites as Biological Control Agents: Some parasites have been “employed” to fight pest species (of insects) but this is difficult to control and not always successful.