Habitats of the Petite Camargue A natural heritage that needs to be preserved The Camargue, and specially the Petite Camargue (west from the Petit Rhône in the départements (counties) of Bouches-du-Rhône, Gard and Hérault), are famous for the diversity of their landscapes and of their natural habitats that are quite always consequences of traditional human activities. This diversity is the result of topographical variations and of the specific coastal dynamics in the Rhône delta: Gulf of Lions beaches, vast sedge meadows in the Scamandre, Crey and Charnier ponds, humid meadows in the Vistre’s low-valley, dunes in l’Espiguette, lagoons and their emblematic pink flamingos. A large number of rare plant and animal species are present in these globally protected habitats; that is why they are protected by preservation policies. In this booklet most of these habitats are presented, as well as the most specific species. Because of the natural diversity of this territory, the State, the Département and the Syndicat Mixte pour la Protection et la Gestion de la Camargue Gardoise have proposed to integrate it into the Natura 2000 network (“habitats” directive), which aim is to protect and to develop major European natural sites. 1 Habitats of the Petite Camargue Cartographie : Scamandre Center Agricultural zone Beaches and dunes Sea club rush beds, behind dunes Pine forests Lagoons Salt steppes Sansouires vegetation) (low salt-resistant Open and seasonal marches Humid meadows Sedge meadows Ponds River bank vegetation Exploration of natural habitat Property of the département Salins d’Aigues-Mortes Seaquarium Musée de la Mer Observatoire de l’Espiguette Mer Méditerrannée 2 Salt marches of Aigues-Mortes Seaquarium – Sea museum Observation post of l’Espiguette Mediterranean Sea Habitats of the Petite Camargue A natural heritage that needs to be preserved 3 The Mediterranean Sea In the Petite Camargue, the coast has an extension of about 25 km (15.5 miles), in the départements of Bouches-du-Rhône, Gard and Hérault. It is part of a greater zone: the Gulf of Lions. The coast of the département of Gard is delimited west by the Vidourle river (“grau” du Ponant) which is known for being particularly capricious, and east by the Rhône Vif river which is a former arm of the Rhône river. The shore slopes down very gently with similar characteristics under the sea . The sandy-muddy bottom is not very interesting for divers in spite of some submerged rocky outcrops located between the port of le Grau du Roi and Sète. But on this coast, which is hard to be explored, the fauna is particularly abundant due to nutritive elements of alluviums brought by the Petit Rhône and the Vidourle rivers. For the preservation of a large number of fish and invertebrates, lagoons are protected places where these species can reproduce or begin to grow in security: mullets, eels, seabasses, seabreans, crabs and many other coastal species. Formerly abundant in Aigues-Mortes Gulf, phanerogame beds (posidonia or eelgrass) have been destroyed during the 1960’s because of the construction of the towns of Port-Camargue and la Grande-Motte. Today few of them only remain in residual places between le Grau du Roi and Palavas. The slowness of their growth turns difficult the evaluation of their regeneration. The ecological importance of these phanerogame beds is considerable; they constitute a priority natural habitat of Community interest (“habitats” directive). 4 The Seabrean (sparus aurata) is one of the most common fish of the coast of the Gulf of Lions. They breed in the sea in November-December. During winter, the life of their larvae is first pelagic; then, once they become little fish (in the spring) they reach protected littoral lagoons in which they stay until they go back to the sea when weather turns colder. They are not sensible to variations of salinity and live in high depths. They eat molluscs and shellfishes that they can crush easily thanks to their strong jaw. Dunes and coastal dunes White Dune The beaches in the Gulf of Lions are very large; and l’Espiguette is certainly one of the most beautiful dune complex of the northern Mediterranean Sea. This environment is a very important natural habitat for plant or animal species that live in these very dry salty and hostile surroundings. In l’Espiguette the dune complex is a very active and diverse one with mobile dunes (non-stabilised by vegetation) (Saharan type – barkhane dunes), white dunes with European beachgrass (or marram grass, Ammophila arenaria) or dunes with Phoenician juniper (Juniperus phoenicea) located near the Baronnets massif. These natural constructions are weakened by the rise of the sea level. Consolidation or reconstruction programmes are implemented, using for example barriers made of wood stakes that hold sand (“ganivelles”). They are present in all the Camargue and Gulf of Lions coast. The Sea Daffodil (Pancratium maritimum) flowers in July and August in the white dunes of the Espiguette beach during the tourist season. Besides picking, the principal cause of extinction of this beautiful bulbous plant is the destruction of dune complex by men (urbanisation, camping sites, car parks) and by storms that erode more and more frequently dunes (already weakened by the diminution of the quantity of sand brought by the Rhône river because of hydroelectric barrages and the fitting-out of silt sites on beaches). The Spanish sand racer (Psammodrommus hispanicus) is a little striated lizard living in and behind dunes in neither very busy nor trampled places. The Spanish sand racer, very quick, can be seen only briefly before he hides under dune plants (Goldilocks (or Eternal Flowers), marram grass, Sea side Chamomile). Unlike many dune animal species that prefer night (snails, insects, batrachians), this protected lizard (which is still quite common in the Petite Camargue) has no problem with the burning sand of the midsummer heat. 5 Sea club rush bed, behind dunes Dune systems are complex, changing according to the distance from the sea, the topography and the salinity of ground water. Dunes and their surroundings are often under the influence of fresh water (rain water lying above salty ground water) allowing the development of lush vegetation. In herbaceous places behind dunes can be observed namely Shoenus nigricans and Erianthus ravennae (Ravenna grass or Plume grass) making barely penetrable beds. In these very particular habitats, exists a very diverse flora with several species of orchids, for example. These beds are often damaged by the development of tourist infrastructures along the beachfronts (car parks and traffic for example), even if it remains in a good state of preservation in l’Espiguette and le Grand Travers. The Bug Orchis (Orchis fragrans) is an orchid specific to low herbaceous open areas located behind dunes. It flowers in May and June and develops in humid sandy meadows behind the dune complex in Boucanet, Grand Travers and l’Espiguettes. In France this species is endangered by the destruction of its habitat by the development of urban and tourist infrastructures. Nevertheless it is a protected species. It is interesting to notice that most of the endangered orchids stations in the Petite Camargue are only located on areas owned by the Conservatoire du Littoral. 6 Summer Lady’s Tresses (Spiranthes aestivalis). This little orchid (maximum 10 cm – 40 inches) makes white tiny spiral flowers in June. It grows in humid depressions (flooded in winter) along with bug orchis In France it is a sharply declining protected species. There are only few stations of it in the Petite Camargue where it is endangered by tourist infrastructures, namely in the surroundings of Port-Camargue. Expansive pasturage (or summer mowing) is a good way to manage these habitats that naturally tend to be more and more closed, preventing orchids from developing. Dunes wooded with Umbrella pines Boucanet pine forest The geomorphologic evolution of coastal dunes leads to a progressive afforestation by bushes like Phillyrea angustifolia or Phoenician juniper (Juniperus pinaster), or by trees like Umbrella pines (Pinus pinea), Maritime pine (Pinus pinaster) or Poplars (Populus sp). These dune coastal sometimes artificial pinewoods (afforestation during the 1960’s and 1970’s) are a characteristic of the Petite Camargue, between les Saintes-Mariesde-la-Mer and Carnon. Boucanet and Grand Travers pinewoods are properties of the Conservatoire du Littoral and can be visited by the public. Pinewoods of the fossil dune complex in Montcalm tend to evolve into broad-leaved trees forests (namely: poplars and oaks). This natural phenomenon is partly the consequence of the withdrawal of the sea (and of the influence of storms and salty sea spray). Sea grape (Ephedra dystachia). This strange plant, close to conifers, forms bushes plastered down over the sand on the grey impeded dunes. Looking like a horsetail it is named sea grape because of its red edible fruits. It makes yellow flowers during the spring. In the Petite Camargue this plant only exists in isolated stations (principally in the salt marshes of Aigues-Mortes); it is very sensitive to trampling and over pasturage. Jewelled lepida (Lacerta lepida). This splendid green-yellowish lizard with lateral blue ocellus is a bit astonishing because of its height (usually 30 cm - 12 inches , maximum 60 cm - 24 inches). This lizard is the tallest one in Europe and it feeds principally on insects and sometimes on fruits and on little vertebrates. He likes open and sunny places in natural or agricultural environment (interdunar depressions, vines,…). Harmless, protected, the Jewelled lapida tends to turn less common in closing habitats (without pasturage, for example). 7 Lagoon Lagoon and pink flamingos In Languedoc and in the Camargue, lagoons are salty ponds (characterised by their high level of salt, sometimes higher than the one of sea water); most of them have communication with the sea thanks to channels called “grau” or canals. Generally lagoons are not very deep and some of them even dry up in summer; then a low salt-resistant vegetation called “sansouire” grows in it. The diversity of lagoons turns possible the development of an important piscicultural and avifaunistic biodiversity. Deep lagoons are nurseries for several sea fish of the Gulf of Lions (seabreans, mullets, eels, etc.). A lot of waders and limicoles (pink flamingos, plovers, shanks, avocets etc.) feed in not very deep lagoons. Most of lagoons are damaged by an excessive division (roads, canals, dikes) and by an increasing level of organic matter in water, causing in summer the “malaïgues” phenomenon (diminution of oxygen rate in water). Common shelduck (Tadorna tadorna). This big and many-coloured duck is characteristic of lagoons where it feeds in pair on shellfishes, molluscs and other aquatic invertebrates. It has a red beak and there is a bulbous tubercle on the beak of males. It nests on the ground (sometimes in former rabbit’s holes!), especially in coastal pine forests near lagoons. It is a protected sedentary species, quite common in the Camargue and in lagoons of Languedoc. 8 Mediterranean gull (Larus melanocephalus). Several laridae (terns, gulls) frequent lagoons in order to feed or to nest. Most common of them are Yellow-legged gulls (Larus cachinnans) and Blackheaded gulls (Larus minutus); but other rare species (Mediterranean gull: Larus melanocephalus, Sandwich tern: Sternus sandvicensis) frequent the protected lagoons of the Salins du Midi in Aigues-Mortes. For them the Camargue is sometimes the single nesting zone in France. The Mediterranean gull can be observed in the evening on the beaches in le Grau du Roi. Salt steppe Salt steppe and statice The Salt Steppe is an open environment with scarce vegetation and sandy ground. Salt steppes are only located near to the coast; often they have a direct or indirect communication way with nearby lagoons (when indirect via brackish ground water). In the winter floods are frequent; this environment is not damaged by sea floods occurring during strong storms. A very specific flora grows in this salty habitat, a natural habitat of Community interest (“Habitats” directive). This very specific flora is composed by several species of statices (Limonium sp) and glassworts (Salicornia sp). Salt steppes are endangered by voluntary alterations of their hydrological scheme, namely by prolonged supply of fresh water. The largest of them are located near to the Chaumadou seasonal pond in the Espiguette plain. Caspia statice (Limonium bellidifolium). Six summer-flowering statice species are present in the Camargue where they are called “fleurs de gardians” (“herdsmen flowers”). The commonest one is Limonium narbonense, used to make bunches of dried flowers. The other species grow quite always on the coast and are less common. Three of them are a characteristic of salt steppes: Caspia statice (limonium bellidifolium) whose leaves dry when flowering, Limonium virgatum and Limonium girardianum which is a protected species in France. Kentish plover (Charadrius alexandrinus). The kentish plover is a minuscule limicole living in the summer on banks of shallow coastal lagoons of the Camargue. It feeds on aquatic invertebrates living in mud and reproduces in sandy environment with little vegetation like salt steppes or even… car parks located near to the beaches of l’Espiguette, for example! In spite of being recently protected it is endangered by the development of coastal tourist infrastructures and by tourism in general (trampling of nests, horse-riding, dogs…). 9 Sansouire Sansouires are often located near lagoons and are closely linked to them. Little differences of some inches in the topography allow halophilic species to prosper in sansouire: annual or bushy glassworts (Salicornia sp, Anthrocnemun sp), seablites (Sueda sp), or Sea Purslane (Halimione portulacoides). Annual sansouires are not very dense; they are flooded from the autumn to the spring, while bushy sansouires are only flooded from time to time. This environment is characterised during the autumn and the winter by wonderful red-orange colours in the sunset light… Sansouires are not very productive for herds of bulls and horses. Nevertheless they are indispensable for extensive ranching, a traditional activity of wetlands of the Camargue. Natterjack toad (Bufo calamita). Very few batrachians live in salty environment because they prefer fresh water; the natterjack toad is one of the few frequenting puddles, ditches or seasonal rain pools in coastal sandy habitats. This little toad is easily recognisable by its vertebral yellow line. It breeds in water very soon in the spring and then hides in humid sand and vegetation in order to avoid summer drought. Like all the batrachians it is a protected species. 10 Eurasian Oystercatcher (Haematopus ostralegus). This elegant limicole is easily recognisable by its large scarlet beak. It frequents coastal lagoons fishing essentially worms and shellfishes. It lives in salty environment and most of the time it nests in sansouires along with pied avocets (Recurvirostra avosetta) and common redshanks (Tringa totanus), close to shallow lagoons. These species can be easily observed in the spring from the birds observatory of the Maison des Vins in l’Espiguette (road between l’Espiguette and le Grau du Roi). Open marshes Hydrologic characteristics of the Camargue marshes are these: an important variation in water level, more or less long flood periods (cycles of one or several years) and drought periods when salt raises up from the ground. Regarding open marshes, flood periods are more frequent. In this type of marshes an annual floating or immersed vegetation can develop; for example muskgrass (Chara spp) that constitutes feeding zones for ducks. Moreover there are also areas with low density of semi-aquatic plants like rushes, an ideal habitat for aquatic micro-fauna. Traditionally, marshes are used for pasturage during the dry season so that they remain open and are very interesting for hunting. Lesser water-plantain (Baldelia ranunculoïdes). This little water-plantain makes little and fragile mauve flowers during the spring. Its repartition is not well known but it seems to be quite scarce in the Camargue even if its not a protected species. It grows in open marshes and is endangered by the closing of its habitat. For this annual species the keeping of pasturage and summer drying is essential; it is well adapted to particular and hard life conditions. It is endangered by the quite permanent flooding of marshes due to hunting management. European pond turtle (Emys orbicularis) is an aquatic black yellow spotted turtle which height reaching 20 to 25 cm (8 to 10 inches). It lives in fresh or brackish stagnant water preferably in canals with scarce vegetation on their banks. It particularly likes sunbathes. It is a very shy species than can be hardly observed in muddy water. At the beginning of summer it leaves water in order to lay its eggs in the banks of marshes, in meadows or in agricultural fallow lands. The Camargue is a place of refuge for this vulnerable and protected species that seems to suffer from pollution of aquatic habitats and non-adapted fishing techniques. 11 Seasonal marshes Seasonal marsh and alkali bulrush On the contrary of open marshes, seasonal marshes are more often dry than flooded. That is why there is more salt raising up from the ground in seasonal marshes, strongly influencing the development of the vegetation. Plants normally growing in sansouires can be observed in it, like scattered glassworts or other little and very specialised plants adapted to the extreme summer drought. This habitat is quite scarce in the Petite Camargue and it is a priority habitat of Community interest (“Habitats directive”). The maintain of natural hydrologic variations (seasonal and annual cycles) is indispensable for its preservation. Cressa cretica. Little bush-like hardy perennial making little pink flowers in the summer. It is very well adapted to seasonal floods and to salt. It grows in grounds with a scattered and short vegetation. By keeping habitats open, extensive pasturage in seasonal marshes allows the development of an highly diversified flora. There is only one station of Cressa cretica known in the Petite Camargue; the species is listed as “vulnerable” in France and is protected in Languedoc-Roussillon and in the Camargue. 12 European bee-eater (Merops apiaster). This beautiful coloured bird (yellow, orange, red, turquoise) evokes Tropics. It is also called “African-hunter” as well as an other coloured bird: the european roller (Coracias garrulus). It reaches our region quite late (end of April or beginning of May) and nests in holes it digs in sandy banks. It essentially feeds on dragonflies, wasps or bees it hunts when flying over open marshes, sansouires or vineyards. Humid meadows Humid meadows and Orchis palustris Natural Mediterranean humid meadows are places of refuge for a large number of common wild plants in continental France, like for example Poet's Narcissus (Narcissus poeticus). These habitats can develop thanks to seasonal floods due to rivers overflowing or to raising up ground-water. They are also preserved thanks to the maintain of traditional activities like extensive pasturage and reaping. The most beautiful meadows can be seen in the Vistre river’s low valley and close to the Costières; they are locally named “Prés du Cailar” (meadows of the village of Le Cailar) and “Launes de Gallician”. They are endangered by cultivation, by the excessive embankment of rivers and by the intensification of drainage. The amelioration of the ecological functioning of their catchment basins is the condition for their long-term preservation. Marsh orchis (Orchis laxiflora). The marsh orchis likes light and grows in wetlands like humid meadows or open marshes. It can be easily recognised by its height and the colour of its flowers (crimson). It flowers in April and May. In France this species is decreasing because of the drying of its habitats. Its a protected species in the Camargue that can be found only in fluvial and lake areas of the Petite Camargue. Its distribution depends on the maintain of traditional extensive pastoral activities. Southern festoon (Zerynthia polyxena). This butterfly is characteristic of wetlands like rivers banks or humid meadows where it gathers pollen from asteraceae. It flies in March/April and lays its eggs under birthwort leaves (Aristolochia spp), the single plant caterpillars can feed. In France it is a protected vulnerable species endangered by the destruction of its habitat by the urbanisation and intensive farming. The Rhône Delta is certainly its biggest refuge in France. 13 Reedbeds A reedbed is a fresh or brackish water marsh characterised by a thick emergent vegetation (Reed grass (Phragmites communis)). It spontaneously appears on banks of ponds. It exists for centuries in periphery of the Scamandre, Crey and Charnier ponds. With quite 3000 ha (7,500 acres) it is the biggest western European reedbed. Reeds are mechanically cropped in winter since the 1950’s; This renewal allows the maintain of sedge meadows. The diversity of birds in this habitat is also interesting for hunting. Moreover sedge meadows are places where pond fish can breed. This habitat is hardly damaged by increasing paludal activities creating compartmentalisation and preventing renewal of water. Reeds are weakened by too short drought periods due to these activities. Purple heron (Ardea purpurea), an African migratory bird. It is littler than the grey heron (Ardea cinerea) and their colours are different. It lives in old non-cropped sedge meadows where it builds its nest using dry reed twigs. In France it is a protected species. It shares its habitat with another mimetic sedentary heron: the great bittern (Botaurus stellaris). Both of them are extremely sensitive to the structure and the age of the sedge meadows, as well as to the water level during their breeding period. Conflicts for use of water or cropping prevent these species from nesting. 14 Bearded tit (Panurus biarmicus). Living exclusively in sedge meadows this little bird can be hardly seen in spite of the grey head and the black whiskers of males. Bearded tits need old sedge meadows in order to build their nests within dry reed twigs entanglements. By eliminating all refuge zones, systematic cropping and hydraulic management are very prejudicial to it. A possible solution is to crop reeds making a mosaic of cropped and non-cropped areas. Ponds The Camargue is a recent geologic formation (10,000 years) resulting of a progressive backward movement of the Mediterranean sea and of alluviums deposits in several beds of the Rhône river, constituting sandy bars in some places. These bars are marks of successive shores that formerly delimited lagoons progressively isolated from the sea and that became shallow brackish ponds. In these ponds, aquatic beds can develop: for example pondweed beds (Potamogeton sp), locally named “gratte”. In the Petite Camargue, Scamandre, Crey and Charnier ponds are brackish ponds with 5 to 15 g/l of salt. They are used for several purposes and are a place where thousands of wintering ducks can rest. Conflicts for use of water endanger this habitat and disturb other habitats like sedge meadows. Common teal (Anas crecca). Most of the individuals of the littlest European duck can be seen in wetlands in November and December. It rests on ponds all day long and it feeds at night in shallow open marshes. This migratory species breeds in Northern Europe (namely in Finland) and in Siberia. In spite of being very popular among hunters it is less endangered by this activity than by the destruction of wetlands, like many other species. This destruction is the very cause of its decrease. Common kingfisher (Alcedo atthis). The Common kingfisher is certainly the most beautiful bird of wetlands thanks to its shimmering colours (orange, blue-green). It can be observed all year long in habitats that are rich in fish (ponds, lagoons, canals, rivers). It digs its nest in wooded soft banks. It is very sensitive to cold (frozen water) and has strongly decreased because of water pollution and transformations of banks (dyking up and rip-rap.) 15 River bank vegetation Near the Mediterranean sea this habitat has been strongly damaged. In the Camargue it has been destroyed by banks transformations (dyking up, recalibrations, creation of canals). It remains only thin borders made of white poplars, white willows and Syrian ashes that sometimes are well developed on banks of the secondary hydraulic network. The relevance of this vegetation for a correct management of rivers has been demonstrated in all recent studies. It contributes to the regulation of water flow, to the protection of banks and of nearby lands against erosion, to the improvement of water quality and to the embellishment landscapes. Finally it is a refuge place for very diversified flora and fauna. Summer snowflake (Leucojum aestivum). This bulbous plant flowers in our region very early in March. Despite of being protected in all the national territory it is strongly decreasing. Camarguese stations on the Or pond banks (Etang de l’Or) and Vitres river’s low valley are among the most important in France. It grows in fresh-water habitats like humid natural meadows, open marshes and even on banks of hydraulic networks forming river bank vegetation. It is endangered by the development of infrastructures and by wrong maintenance of banks (non-respect of its natural cycle). 16 Orange-spotted emerald (Oxygastra curtisii). This mediumsized dragonfly with green metallic reflections is only present in flowingwater zones where it has a well delimited territory. It particularly likes bushy river bank vegetation. This species is a vulnerable and protected one and it has been observed in some places in the Petite Camargue (Vidourle and Vistre rivers). Its development depends on water quality but also on transformations of banks and of their vegetation (elimination of bushes). Wetlands in the Petite Camargue are also… Bulls and Camargue Horses ranching Reeds cropping in winter Waterbirds hunting Harvesting of salt Fishing in ponds and lagoons In the Petite Camargue the wetlands and the diversity of natural environments are preserved thanks to traditional human activities (bulls and Camargue Horses ranching, reeds cropping, salt culture, fishing, hunting…) The maintain of these traditional activities is essential for the preservation of the faunistic and the floristic species described in this booklet. However an intensification of these activities can origin a degradation of these habitats and the disappearance of species. An equilibrium (called “sustainable development”) has to be found in order to protect both ecological diversity and human activities in the Petite Camargue. That is why the Petite Camargue is part of the Ramsar network since 1996. The Ramsar Convention on Wetlands of International Importance especially as Waterfowl Habitat provides the framework for the conservation and wise use of wetlands and their resources. The “wise use” of these natural resources is the common objective of the 800 wetlands of international importance making part of the Ramsar network all over the world. 17 18 Bibliography: Flore de Camargue J. MOLINA – Parc Naturel Régional de Camargue Camargue, canards et foulques Fonctionnement et devenir d’un prestigieux quartier d’hiver A. TAMISIER / O. DEHORTER – Centre Ornithologique du Gard Oiseaux de Camargue – The Birds of Camargue Annotated check-list of the 336 bird species recorded in the Camargue (written in English and French) P. ISENMANN – Société d’Etudes Ornithologiques La Nature méditerranéenne en France Les Ecologistes de l’Euzière – Editions Delachaux et Niestlé “Faune du Littoral Gardois” brochures (Reptiles et Amphibiens, Mammifères terrestres, Mammifères marins, Mollusques marins, Poissons marins, Invertébrés marins, Oiseaux marin et lagunaires, Oiseaux des marais doux et des roselières, Oiseaux des zones sèches et agricoles). Collectif “Faune du Littoral Gardois” Routes buissonnières en Camargue Gardoise 10 road itineraries for exploration of the Camargue Gardoise Syndicat Mixte pour la Protection et la Gestion de la Camargue Gardoise Documentary films (videos) (made by the Syndicat Mixte pour la Protection et la Gestion de la Camargue Gardoise). Les voies du sagneur (12 minutes) Hommes et métiers de Camargue (23 minutes) Les petits métiers de la pêche en Camargue Gardoise (18 minutes) “Flore rare et menacée en Camargue Gardoise” (Bill) (Syndicat Mixte pour la Protection et la Gestion de la Camargue Gardoise / Association pour la Connaisance et la Conservation des Milieux naturels). Available at: Centre de Découverte du Scamandre Telephone: 04 66 73 52 05 Website: www.camarguegardoise.com 19 Addresses: Syndicat Mixte pour la Protection et la Gestion de la Camargue Gardoise Hôtel du Département – Rue Guillemette – 30044 Nîmes cedex (Telephone: 04 66 76 77 89 – Fax: 04 66 76 52 76) Centre de Découverte du Scamandre Les Iscles – Gallician – 30600 Vauvert (Telephone: 04 66 73 52 05 – Fax: 04 66 73 52 16) Voluntary Nature Reserve, exploration of natural fluvial and lake habitats, museography, visits, thematic activities for schools. Collectif “Faune du Littoral Gardois” A.C.C.M. – B.P. 6 – 30006 Nîmes cedex 04 (Telephone and Fax: 04 66 67 26 87) Observatoire des oiseaux de l’Espiguette Maison des Vins de l’Espiguette – Route de l’Espiguette 30240 Le Grau-du-Roi Observation of avifauna of lagoons, guided tours by the Centre de Découverte du Scamandre (telephone 04 66 73 52 05) Seaquarium / Musée de la Mer Rive Gauche – Avenue du Palais de la Mer 30240 Le Grau-du-Roi (Telephone: 04 66 51 57 57) Exploration of the sea, sea aquariums, muesography, visits and thematic activities for schools. 20