Lecture Exam 4: Summer 2009 Update 7/12 Chapter 11, 12, 13 Late Paleozoic Earth History, Invertebrate and Vertebrate Paleozoic Life The test will cover selected material from textbook Chapter 11, 12 and 13 AND several pages from Chapter 9. Ch 9 (Suggested questions to reveiw: 6) Eukaryotic Cell Evolution: from prokaryotes to eukaryotes through endosymbiosis Review text Pages 179-181 Prokaryote vs Eukaryote (Fig. 9.14) Evolution of complex cell with nucleus containing genetic material and organelles Advantages of eukaryotes: specialized cells to perform specific functions allow for larger organisms sexual reproduction leads to variations which may be favorable for adaptation One of central themes of evolution is that cells became more complex over time. The evolution of eukaryotic cells as observed in the fossil record supports this. Ch 11 (Suggested questions to review: 2, 3, 4, 6, 7, 10, 11, 12, 13, 16) Figure 10.3 Cratonic sequences will be provided. Construction of Pangaea: During the early Paleozoic, Laurentia had collided with Baltica to form Laurasia This collision is marked by Caledonian orogeny and closed the northern Iapetus Ocean. Acadian orogeny – second step in building the Appalachian mountains, affected present day New England area Erosion of Acadians into present day New York State formed the Catskill Delta with coarse sediments nearer the mountain range and finer sediments away from the mountain range. The Kaskaskia sea extended to western New York State during the Devonian. The northern European counterpart to the North American Catskill Delta is the “Old Red Sandstone”, a clastic wedge forming on the other side of the mountain ranges formed during the collision of Laurasia and Baltica. (Fig. 11.1a and Fig. 11.17) During the Carboniferous, (Mississippian and Pennsylvanian periods) Gondwana moved toward Laurasia and rotated clockwise, causing deformation by folding and uplift of mountains in a NE to SW direction from the northern Appalachians to the Ouachita mountains of present day Oklahoma. The final stage of building the Appalachians (Allegheny, Ouachita) (Fig. 11.2 a) By the late Permian Gondwana had sutured to Laurasia, other continents had joined to Laurasia, and Pangaea had formed. Pangaea contained most of earth’s land masses and left a huge Panthalassa ocean. Late Paleozoic events: Devonian: Kaskaskian Cratonic sequence inundated North America (Fig. 11.4) with transitional marine-continental boundaries just west of the Appalachian highlands. Extensive reef development is observed today in western Canada and western Texas as evidence of the transgression. Carboniferous mountain building – Cordilleran – Ancestral Rockies were uplifed by faulting. Late Carboniferous, inland sea (from Absaroka transgression) caused transitional delta environments in some areas of the north-central and eastern U.S. (Fig. 11.8) Cyclothems: repeated alternations of marine and nonmarine environments, usually in areas of low relief. Represent transgressive and regressive sequences at transitional areas such as present day Gulf Coast deltas and swamps. Associated with coal formation in swamps as sea level rises, floods, buries, then drops and erodes an area; then repeats the cycle. Carboniferous – Permian: Growth of glaciation over Gondwana which spread upwards into the temperate latitudes. (See Fig. 11.2b) Natural resources of the Late Paleozoic include coal, evaporite deposits (i.e. salt), and metallic minerals formed during mountain building. Ch 12 (Suggested questions to review: 5, 6, 8, 10, 11, 12,) invertebrate Cambrian Explosion Shelly Fauna (shelled organisms) – why so abundant in the Cambrian fossils record? archeocyathid – first reef builder – emerged in Cambrian and were extince at end of Cambrian Sponges, brachiopods, corals all appear in early Paleozoic trilobite – arthropod (insect-like marine organism) – abundant in early Paleozoic – extinct by Permian planktonic – nektonic – benthic organisms (their marine environments) sessile (attach to bottom to filter feed) trophic levels – tiers of food production and consumptions within a feeding hierarchy = food web primary producers (autotrophs) – phytoplankton primary consumers – feed on producers (filter feeders) and secondary consumers (predators) feed on primary consumers decomposers – bacteria break down dead organisms not eaten by scavengers brachiopods (bivalve filter feeders) Burgess Shale – Cambrian - importance and type of organisms notochord, arthropod exoskeleton – advantages for living organism; advantages for fossil preservation mass extinction – possible causes; THREE extinctions in Paleozoic -- benefits (see Fig. 12.19 – can you read the chart?) Ch 13 (Suggested questions to review Q 10, 11, 15) chordate (has notochord, dorsal hollow nerve cord and gill slits) vertebrate (a subdivision of chordates) Fig. 13.5: major fish groups (note lines indicate range) no need to memorize – figure would be given for interpretation Devonian – “Age of Fishes” Which fish types survived to recent time? (loss of shallow habitat at end of Permian favored adaptation to deep waters) Jawless fish, jawed fish, ray-finned vs lobe finned Challenges to animals moving to land (avoid dessication, accomplish reproduction, mobility, skeletal support, respiration) Challenges for plants to move (reproduction) Crossopterygians – lob-finned fish – why important? amphibian (returns to water to reproduce and can live on land) Compare lobe-fin fish and amphibian (Fig. 13.11 – note rib cage for lungs) Tiktaalik rosaea – “missing link” – between fish and amphibian amniote egg – significant – contains hard shell, food supply -- reptiles could reproduce completely on land endothermic (warm blooded – mammals, birds) vs exothermic (cold blooded organisms – reptiles, snakes) Permian extinction: Cause is not clear but there are multiple possible contributing factors. Effects of formation of a supercontinent: Collisions of land masses eliminate continental shelf area which provide warm, shallow marine environments favorable to Paleozoic life forms. Significant loss of habitat for marine organisms. Long, high mountain ranges cause “rain shadow effect” causing a windy dry interior.