CURRICULUM VITAE
Peter Morrison Vitousek
Academic Background
B.A. cum laude in Political Science, Amherst College, 1971
Ph.D. in Biology, Dartmouth College, 1975
Assistant Professor of Zoology and Biology, Indiana University, 1975–1979
Associate Professor of Botany and Biology, University of North Carolina—Chapel
Hill, 1980–1983
Associate Professor and Professor of Biological Sciences, Stanford University,
1984–
Clifford G. Morrison Professor of Population and Resource Studies, Stanford
University, 1993–
Honors and Awards
Pew Scholar in Conservation and Environment, 1990
W.S. Cooper Award, Ecological Society of America, 1992; R.H. MacArthur Award,
Ecological Society of America, 1993; Time/CNN America’s best ecologist,
2002; Princeton Environmental Prize, 2002; Marsh Ecology Book Award,
British Ecological Society, 2005; Tansley Lecturer, British Ecological Society,
2005; Award for Scientific Reviewing, National Academy of Sciences, 2006.
Fellow, National Academy of Sciences, 1992; American Academy of Arts and
Sciences, 1993; American Association for the Advancement of Science, 1994;
California Academy of Sciences, 1994
D.Sc., Amherst College, 2003
Teaching
Ecology (lecture course). Indiana, North Carolina, and Stanford.
Environmental Biology (non-majors course, lecture). North Carolina and Stanford.

Ecosystems of the Tropics. Stanford.
Biology and Global Change. Stanford.
Terrestrial Biogeochemistry. Stanford.
Island Ecology. Stanford.
Publications
1975 Reiners, W.A., R.H. Marks, and P.M. Vitousek. Heavy metals in sub-alpine and alpine soils of New Hampshire. Oikos 26: 264-275.
Vitousek, P.M., and W.A. Reiners. Ecosystem succession and nutrient retention: a hypothesis. BioScience 25: 376-381. ISI Citation Classic, 1991.
1976 Vitousek, P.M., and W.A. Reiners. Ecosystem development and the biological control of streamwater chemistry. p. 665-680. In Nriagu, J. (ed.),
Environmental Biogeochemistry . Ann Arbor Science, Michigan.
1977 Vitousek, P.M. The regulation of element concentrations in mountain streams in the northeastern United States. Ecological Monographs 47: 65-87.
1979 Gorham, E., P.M. Vitousek, and W.A. Reiners. The regulation of element budgets over the course of terrestrial ecosystem succession . Annual Review of
Ecology and Systematics 10: 53-84.
Vitousek, P.M., J.R. Gosz, C.C. Grier, J.M. Melillo, W.A. Reiners, and R.L.
Todd. Nitrate losses from disturbed ecosystems. Science 204: 469-474.
Vitousek, P.M. and J.M. Melillo. Nitrate losses from disturbed ecosystems: patterns and mechanisms. Forest Science 25: 605-619.
1980 Blank, J.L., R.K. Olson, and P.M. Vitousek. Nutrient uptake by a diverse spring ephermeral community. Oecologia 47: 96-98.
Foster, M.M., P.M. Vitousek, and P.A. Randolph. The effect of Ambrosia artemisiifolia on nutrient cycling in a first-year old field. American Midland
Naturalist 103: 106-113.
Rosswall, T.H. and P.M. Vitousek. Research priorities and future cooperation in
West African nitrogen cycling research. In Rosswall, T.H. and R. Soderlund
(eds .), Nitrogen Cycling in West African Ecosystems . SCOPE/UNEP
International Nitrogen Unit, Stockholm, Sweden. pp. 421-428.

Vitousek, P.M. Nitrogen losses from disturbed ecosystems--ecological considerations. In Rosswall, T.H. and R. Soderlund (eds.), Nitrogen Cycling in
West African Ecosystems . SCOPE/UNEP International Nitrogen Unit,
Stockholm, Sweden. pp. 39-53.
1981 Matson, P.A. and P.M. Vitousek. Nitrification potentials following clearcutting in the Hoosier National Forest, Indiana. Forest Science 27: 781-791.
Robertson, G.P. and P.M. Vitousek. Nitrification potentials in primary and secondary succession. Ecology 62: 376-386.
Vitousek, P.M. Clearcutting and the nitrogen cycle. Ecological Bulletin
(Stockholm) 33: 631-642.
Vitousek, P.M., W.A. Reiners, J.M. Melillo, C.C. Grier, and J.R. Gosz. Nitrogen cycling and loss following forest perturbation: the components of response. In
Barrett, G.W. and R. Rosenberg (eds.), Stress Effects on Natural Ecosystems .
Wiley and Sons, London. pp. 115-127.
Vitousek, P.M. and P.S. White. Process studies in succession. In Shugart, H.H.,
D.B. Botkin, and D. West (eds.), Forest Succession: Concept and Application .
Springer-Verlag, New York. pp. 267-276.
1982 Vitousek, P.M. Nutrient cycling and nutrient use efficiency. American Naturalist
119: 553-572.
Vitousek, P.M., J.R. Gosz, C.C. Grier, J.M. Melillo, and W.A. Reiners. A comparative analysis of potential nitrification and nitrate mobility in forest ecosystems. Ecological Monographs 52: 155-177.
1983 Bolin, B., P.J. Crutzen, P.M. Vitousek, R.G. Woodmansee, E.D. Goldberg, and
R.B. Cook. Interactions of biogeochemical cycles. p. 8-40. In Bolin, B. and
R.B. Cook (eds.), The Biogeochemical Cycles and Their Interactions . John
Wiley and Sons, Chichester.
Vitousek, P.M. The effects of deforestation on air, soil, and water. p. 223-245.
Bolin, B. and R.B. Cook (eds.), The Biogeochemical Cycles and Their
Interactions . John Wiley and Sons, Chichester.
In
Vitousek, P.M. Nitrogen turnover in a ragweed-dominated first-year old-field in southern Indiana. American Midland Naturalist 110: 46-53.

Vitousek, P.M. Mechanisms of ion leaching in natural and managed ecosystems. p. 129-144. In Mooney, H.A. and M. Godron (eds.), Disturbance and
Ecosystems: Components of Response . Springer-Verlag, New York.
Vitousek, P.M., H.L. Allen, and P.A. Matson. Impacts of management practices on soil nitrogen status. p. 25-39. In Maintaining Forest Site Productivity,
Proc. Society of American Foresters, Appalachian Chapter.
Vitousek, P.M., K. Van Cleve, N. Balakrishnan, and D. Mueller-Dombois. Soil development and nitrogen turnover on recent volcanic substrates in Hawaii.
Biotropica 15: 268-274.
1984 Mueller-Dombois, D., P.M. Vitousek, and K.W., Bridges. Canopy dieback and ecosystem processes in Pacific forests. Hawaii Botanical Science Paper 44,
100 p. University of Hawaii.
Saterson, K.A., and P.M. Vitousek. Fine root biomass and nutrient cycling in
Aristida stricta in a North Carolina coastal plain savanna. Canadian Journal of Botany 62: 823-829.
Vitousek, P.M. Litterfall, nutrient cycling, and nutrient limitation in tropical forests. Ecology 65 : 285-298.
Vitousek, P.M. Anion fluxes in three Indiana forests. Oecologia 61: 105-108.
Vitousek, P.M. Terrestrial nitrogen cycles: some unanswered questions. p. 209-
218. In Moore, B, and M. Dastoor, The Interactions of Global Biogeochemical
Cycles . Jet Propulsion Lab Publication 84-21.
Vitousek, P.M. A general theory of forest nutrient dynamics. p. 121-135. In
Agren, GI (ed.), State and Change of Forest Ecosystems--Indicators in current research. Swedish Univ. Agric. Sci. Dept. Ecology and Environmental
Research Report 13.
Vitousek, P.M., and P.A. Matson. Mechanisms of nitrogen retention in forest ecosystems: a field experiment. Science 225: 51-52.
1985 Pye, J.M., and P.M. Vitousek. Soil and nutrient removals by erosion and windrowing at a Southeastern U.S. Piedmont site. Forest Ecology and
Management 11: 145-155.
Vitousek, P.M. Plant community turnover and ecosystem nutrient flux. p. 325-
333. In Pickett, S.T.A. and P.S. White (eds.), The Ecology of Natural
Disturbance and Patch Dynamics . Academic Press, New York.

Vitousek, P.M. Plant and animal invasions: can they alter ecosystem processes. p. 169-176. In Halvorson, H.O., D. Pramer, and M. Rogul (eds.), Engineered organisms in the environment: Scientific issues . Am. Soc. Microbiology,
Washington, D.C.
Vitousek, P.M., and P.A. Matson. Causes of delayed nitrate production in two
Indiana forests. Forest Science 31: 122-131.
Vitousek, P.M., and P.A. Matson. Intensive harvesting and site preparation decrease soil nitrogen availability in young plantations. Southern Journal of
Applied Forestry 9: 120-125.
Vitousek, P.M., and P.A. Matson. Disturbance, nitrogen availability, and nitrogen losses in an intensively managed loblolly pine plantation. Ecology 66: 1360-
1376.
1986 Birk, E.M., and P.M. Vitousek. Nitrogen availability and nitrogen use efficiency in loblolly pine stands on the Savannah River Plant, South Carolina. Ecology
67: 69-79.
Chapin, F.S. III, P.M. Vitousek, and K. Van Cleve. The nature of nutrient limitation in plant communities. American Naturalist 127 : 48-58.
Heal, O.W., and P.M. Vitousek. Environmental controls over ecosystem processes. p. 155-159. In Van Cleve, K., F.S. Chapin, P.W. Flanagan, L.A.
Viereck, and C.T. Dyrness (eds.), Forest Ecosystems in the Alaskan Taiga: A
Synthesis of Structure and Function . Springer-Verlag, New York.
Vitousek, P.M. Biological invasions of ecosystem properties: can species make a difference? p. 163-176. In Mooney, H.A., and J. Drake (eds .), Biological
Invasions of North America and Hawaii . Springer-Verlag.
Vitousek, P.M., and S.W. Andariese. Microbial transformations of labelled nitrogen in a clearcut pine plantation. Oecologia 68: 601-605.
Vitousek, P.M., and J.S. Denslow. Nitrogen and phosphorus availability in treefall gaps in a lowland tropical rainforest. Journal of Ecology 74: 1167-
1178.
Vitousek, P.M., P.R. Ehrlich, A.H. Ehrlich, and P.A. Matson. Human appropriation of the products of photosynthesis. Bioscience 36: 368-373.
Vitousek, P.M., and R.L. Sanford, Jr. Nutrient cycling in moist tropical forest.
Annual Review of Ecology and Systematics 17: 137-167.

Williams, W.E., K. Garbutt, F.A. Bazzaz, and P.M. Vitousek. The effect of elevated CO
2
on plants. IV. Two deciduous-forest tree communities.
Oecologia 69: 454-459.
1987 Denslow, J.S., P.M. Vitousek, and J.C. Schultz. Bioassays of nutrient limitation in a tropical rainforest soil. Oecologia 74: 370-376.
Matson, P.A., and P.M. Vitousek. Cross-system comparison of soil nitrogen transformations and nitrous oxide fluxes in tropical forests. Global
Biogeochemical Cycles 1: 163-170.
Matson, P.A., P.M. Vitousek, J.J. Ewel, M.J. Mazzarino, and G.P. Robertson.
Nitrogen transformations following tropical forest felling and burning on a volcanic soil. Ecology 68: 491-502.
Mooney, H.A., P.M. Vitousek, and P.A. Matson. Exchange of materials between terrestrial ecosystems and the atmosphere. Science 238: 926-932.
Robertson, G.P., P.M. Vitousek, P.A. Matson, and J.M. Tiedje. Denitrification in a clearcut loblolly pine location in the Southeastern U.S. Plant and Soil 97:
119-129.
Turner, D.R., and P.M. Vitousek. Nodule biomass of the nitrogen-fixing alien
Myrica faya Ait. in Hawaii Volcanoes National Park. Pacific Science 41: 186-
190.
Vitousek, P.M. and J.S. Denslow. Differences in soil phosphorus among soils of the La Selva Biological Station, Costa Rica. Biotropica 19: 167-170.
Vitousek, P.M., L.L. Loope, and C.P. Stone. Introduced species in Hawai'i:
Biological effects and opportunities for ecological research. Trends in Ecology and Evolution 2: 224-227.
Vitousek, P.M. and L.R. Walker. Colonization, succession, and resource availability; ecosystem-level interactions. p. 207-223. In Gray, A., M.
Crawley, and P.J. Edwards (eds.), Colonization, Succession, and Stability .
Blackwell Scientific, Oxford.
Vitousek, P.M., L.R. Walker, L.D. Whiteaker, D. Mueller-Dombois, and P.A.
Matson. Biological invasion by Myrica faya alters ecosystem development in
Hawaii. Science 238: 802-804.
Walbridge, M.R., and P.M. Vitousek. Processes affecting the measurement of phosphorus mineralization by 32 PO
4
isotope dilution in acid organic soils.
Biology and Biochemistry 19: 709-717.
Soil

1988 Andariese, S.W., and P.M. Vitousek. Soil nitrogen turnover is altered by herbicide treatment in a piedmont forest soil. Forest Ecology and Management
23: 19–25.
Livingston, J., P.M. Vitousek, and P.A. Matson. Nitrous oxide flux and nitrogen transformations across a landscape gradient in Amazonia. Journal of
Geophysical Research 93: 1593–1599.
Vitousek, P.M. Diversity and biological invasions of oceanic islands. p. 181–189.
In Wilson, E.O. (ed.), BioDiversity . National Academy Press.
Vitousek, P.M., T. Fahey, D.W. Johnson, and M.J. Swift. Element interactions in forest ecosystems: succession, allometry, and input-output budgets.
Biogeochemistry 5: 7–34.
Vitousek, P.M. and P.A. Matson. Nitrogen transformations in tropical forest soils. Soil Biology and Biochemistry 20: 361–367.
Vitousek, P.M., P.A, Matson, and D.R. Turner. Elevational and age gradients in
Hawaiian montane rainforest: foliar and soil nutrients. Oecologia 77: 565–
570.
1989 Binkley, D., and P.M. Vitousek. Soil nutrient availability. p. 75–96. In Pearcy,
R.W., J.R. Ehleringer, H.A. Mooney and P.W. Rundel (eds .), Physiological
Plant Ecology: Field Methods and Instrumentation . Chapman and Hall,
London.
Ehrlich, P.R., G.C. Daily, A.H. Ehrlich, P.A. Matson, and P.M. Vitousek. Global change and carrying capacity: implications for life on Earth. p. 19–27. In
DeFries, R.S., T.F. Malone (eds), Global Change and Our Common Future ,
National Academy Press, Washington, D.C.
Gower, S.T., and P.M. Vitousek. Effects of nutrient amendments on fine root biomass in a primary successional forest in Hawai’i. Oecologia 81: 566–568.
Luizão, F.J., P.A. Matson, G.L. Livingston, R. Luizão, and P.M. Vitousek.
Nitrous oxide flux following tropical land clearing. Global Biogeochemical
Cycles 3: 281–285.
Matson, P.A., P.M. Vitousek, and D.S. Schimel. Regional extrapolation of trace gas flux based on soils and ecosystems. p. 97–108. In Andreae, M.O., and D.S.
Schimel (eds.), Exchange of Trace Gases Between Terrestrial Ecosystems and the Atmosphere . John Wiley and Sons, Chichester.

Palm, C., P. Robertson, and P.M. Vitousek. Nitrogen availability. p. 162–168. In
Anderson, J.M., and J.S. Ingram (eds.), Tropical Soil Biology and Fertility: A
Handbook of Methods . CAB International, Wallingford, England.
Ramakrishnan, P.S., and P.M. Vitousek. Ecosystem-level consequences of biological invasions. p. 281–300. In Drake, J., F. DiCastri, R. Groves, F.
Kruger, H.A. Mooney, M. Rejmanek, and M. Williamson (eds.), Biological
Invasions: a Global Perspective . Wiley and Sons, Chichester.
Vitousek, P.M. Terrestrial Ecosystems. p. 78–83. In DeFries, R.S., and T.F.
Malone (eds.), Global Change and Our Common Future . National Academy
Press, Washington, D.C.
Vitousek, P.M. What are the relative roles of biological production, micrometeorology, and photochemistry in controlling the flux of trace gases between terrestrial ecosystems and the atmosphere. p. 249–261. In Andreae,
M.O. and D.S. Schimel (eds.), Exchange of Trace Gases Between Terrestrial
Ecosystems and the Atmosphere . John Wiley and Sons, Chichester.
Vitousek, P.M., P.A. Matson, and K. Van Cleve. Nitrogen availability and nitrification during succession: primary, secondary, and old-field seres. Plant and Soil 115: 229–239.
Vitousek, P.M., P.A. Matson, C. Volkmann, J.M. Maass, and G. Garcia. Nitrous oxide flux from seasonally-dry tropical forests: a survey. Global
Biogeochemical Cycles 3: 375–382.
Vitousek, P.M., G. Shearer, and D.H. Kohl. Foliar 15 N natural abundance in
Hawaiian rainforest: patterns and possible mechanisms. Oecologia 78: 383–
388.
Vitousek, P.M., and L.R. Walker. Biological invasion by Myrica faya in Hawai'i: plant demography, nitrogen fixation, and ecosystem effects. Ecological
Monographs 59: 247–265. W.S. Cooper Award, 1992.
1990 Denslow, J.S., J.C. Schultz, and P.M. Vitousek, and B.R. Strain. Growth responses of tropical shrubs to treefall gap environments. Ecology 71: 165–
179.
Huenneke, L.F., S.P. Hamburg, R. Koide, H.A. Mooney, and P.M. Vitousek.
Effects of soil resources on plant invasion and community structure in
Californian serpentine grassland. Ecology 71: 478–491.

Huenneke, L.F., and P.M. Vitousek. Seedling and clonal recruitment of the invasive tree Psidium cattleianum : implications for management of native
Hawaiian forests. Biological Conservation 53: 199–211.
Matson, P.A., and P.M. Vitousek. Remote sensing and trace gas flux. p. 157–
167. In Hobbs, R., and H.A. Mooney (eds.), Remote Sensing and Global
Change . Springer-Verlag.
Matson, P.A., and P.M. Vitousek. Ecosystem approach to a global nitrous oxide budget. BioScience 40: 667–672.
Matson, P.A., P.M. Vitousek, G. Livingston, and N. Swanberg. Sources of variation in N
2
O flux from forest ecosystems of central Amazonia. Journal of
Geophysical Research 95: 16,789–16,798.
Vitousek, P.M. Biological invasions and ecosystem processes: towards an integration of population biology and ecosystem studies. Oikos 57: 7–13.
Vitousek, P.M., C.B. Field, and P.A. Matson. Variation in foliar  13 C in
Hawaiian Metrosideros polymorpha : a case of internal resistance? Oecologia
84: 362–370.
Woodward, S.A., P.M. Vitousek, K.A. Matson, R.F. Hughes, K. Benvenuto, and
P.A. Matson. Avian use of the introduced nitrogen-fixer Myrica faya in
Hawaii Volcanoes National Park. Pacific Science 44: 88–93.
1991 Davidson, E.A., P.M. Vitousek, P.A. Matson, R. Riley, G. Garcia-Méndez, and
J.M. Maass. Soil emissions of nitric oxide in a seasonally dry tropical forest of
Mexico. Journal of Geophysical Research 96: 15,439–15,445.
Garcia-Méndez, G., J.M. Maass, P.A. Matson, and P.M. Vitousek. Nitrogen transformations and nitrous oxide flux in a tropical deciduous forest in Mexico.
Oecologia 88: 362–366.
Hughes, R.F., P.M. Vitousek, and J.T. Tunison. Effects of invasion by fireenhancing C
4
grasses on native shrubs in Hawaii Volcanoes National Park.
Ecology 72: 743–747.
Vitousek, P.M. An analysis of forests as a means for counteracting the build-up of carbon dioxide in the atmosphere. Journal of Environmental Quality 20:
348–354.
Vitousek, P.M., and R.W. Howarth. Nitrogen limitation on land and in the sea: how can it occur? Biogeochemistry 13: 87–115.

Vitousek, P.M., and P.A. Matson. Gradient analysis of ecosystems. p. 287–298.
In Cole, J.J., G. Lovett, and S. Findlay (eds.), Comparative Analysis of
Ecosystems: Patterns, Mechanisms, and Theories . Springer-Verlag.
Vitousek, P.M., and P.A. Matson. Commentary: Ecological perspectives on tropical deforestation and biosphere-atmosphere interaction. Climatic Change
19: 159–162.
Walker, L.R., and P.M. Vitousek. Interactions of an alien and a native tree during primary succession in Hawaii Volcanoes National Park. Ecology 72: 1449–
1455.
1992 D’Antonio, C.M., and P.M. Vitousek. Biological invasions by exotic grasses, the grass-fire cycle, and global change. Annual Review of Ecology and Systematics
23: 63–87.
Townsend, A., P.M. Vitousek, and E. Holland. Tropical soils could dominate the short-term carbon cycle feedbacks to increased global temperatures. Climatic
Change 22: 293–303.
Vitousek, P.M. Perspectives on the nitrogen cycle. p. 151–163. In Moore, B., and
D.S. Schimel (eds.), Trace Gases and the Biosphere . Office of
Interdisciplinary Earth Studies, University Corp. for Atmospheric Research,
Boulder, Colorado.
Vitousek, P.M. Global environmental change: An introduction. of Ecology and Systematics 23: 1–14.
Annual Review
Vitousek, P.M. Effects of alien plants on native ecosystems. p. 29–41. In Stone,
C.P., C.W. Smith, and J.T. Tunison (eds.), Alien Plant Invasions in Native
Ecosystems of Hawai’i: Management and Research . Cooperative National
Park Resources Study Unit, University of Hawaii.
Vitousek, P.M., S.W. Andariese, P.A. Matson, L. Morris, and R.L. Sanford.
Effects of harvest intensity, site preparation, and herbicide use on soil nitrogen transformations in a young loblolly pine plantation. Forest Ecology and
Management 49: 277–292.
Vitousek, P.M., G. Aplet, D.R. Turner, and J.J. Lockwood. The Mauna Loa environmental matrix: foliar and soil nutrients. Oecologia 89: 372–382.
Vitousek, P.M., and P.A. Matson. Tropical forests and trace gases: potential interactions between tropical biology and the atmospheric sciences. Biotropica
24: 233–239.

1993 Davidson, E.A., P.A. Matson, P.M. Vitousek, R. Riley, K. Dunkin, G. Garcia-
Mendez, and J.M. Maass. Processes regulating soil emissions of NO and N
2
O in a seasonally dry tropical forest. Ecology 74: 130–139.
Hughes, F., and P.M. Vitousek. Barriers to shrub reestablishment following fire in seasonal submontane woodland in Hawaii. Oecologia 93: 557–563.
Menaut, J.C., L. Abbadie, and P.M. Vitousek. Nutrient and organic matter dynamics in tropical ecosystems. p. 215–231. In Crutzen, P., and J.
Goldammer (eds), Fire in the Environment: The Ecological, Atmospheric, and
Climatic Importance of Vegetation Fires . John Wiley and Sons, Chichester.
Potter, C.S., J.T. Randerson, C.B. Field, P.A. Matson, P.M. Vitousek, H.A.
Mooney, and S.A. Klooster. Terrestrial ecosystem production: a process model based on global satellite and surface data. Global Biogeochemical
Cycles 7: 811–842.
Vitousek, P.M. Global dynamics and ecosystem processes: Scaling up or scaling down? p. 169–177. In Ehleringer, J.R., and C.B. Field (eds), Scaling
Physiological Processes: Leaf to Globe . Academic Press, San Diego.
Vitousek, P.M., and D.U. Hooper. Biological diversity and terrestrial ecosystem biogeochemistry. p. 3–14. In Schulze, D., and H.A. Mooney (eds), Biological
Diversity and Ecosystem Function . Springer-Verlag.
Vitousek, P.M., and P.A. Matson. Agriculture, the global nitrogen cycle, and trace gas flux. p. 193–208, In Oremland, R. (ed), Biogeochemistry of Global
Change: Radiatively Active Trace Gases . Chapman and Hall, New York.
Vitousek, P.M., L.R. Walker, L.D. Whiteaker, and P.A. Matson. Nutrient limitation to plant growth during primary succession in Hawaii Volcanoes
National Park. Biogeochemistry 23: 197–215.
1994 Aplet, G.H., and P.M. Vitousek. An age-elevation matrix analysis of Hawaiian rainforest succession. Journal of Ecology 82: 137–147.
Chorover, J., P.M. Vitousek, D.A. Everson, A.M. Esperanza, and D. Turner.
Solution chemistry profiles of mixed-conifer forests before and after fire.
Biogeochemistry 26: 115–144.
Joel, G., G. Aplet, and P.M. Vitousek. Leaf morphology along environmental gradients in Hawaiian Metrosideros polymorpha . Biotropica 26: 17–22.
Karpa, D.M., and P.M. Vitousek. Successional development of a Hawaiian montane grassland. Biotropica 26: 2–11.

Potter, C.S., P.A. Matson, and P.M. Vitousek. Evaluation of soil database attributes in a terrestrial carbon cycle model: implications for global change research. p. 281–302. In : Michener, W.K. (ed), Environmental Information
Systems Analysis . Taylor and Francis, U.K.
Raich, J.W., R.H. Riley, and P.M. Vitousek. Use of root-ingrowth cores to assess nutrient limitations in forest ecosystems. Canadian Journal of Forest
Research 24: 2135–2138.
Vitousek, P.M. Factors controlling ecosystem structure and function. p. 87–97.
In Amundson, R., J. Harden, and M. Singer (eds .), Factors of Soil Formation; a 50th Anniversary Symposium . Special Publication of the Soil Science
Society of America.
Vitousek, P.M. Potential nitrogen fixation during primary succession in Hawaii
Volcanoes National Park. Biotropica 26: 234–240.
Vitousek, P.M. Beyond global warming: ecology and global change. Ecology 75:
1861–1876. ISI Citation Classic, 2001.
Vitousek, P.M., D.R. Turner, W.J. Parton, and R.L. Sanford. Litter decomposition on the Mauna Loa environmental matrix, Hawaii: Patterns, mechanisms, and models. Ecology 75: 418–429.
1995 Crews, T.E., K. Kitayama, J. Fownes, D. Herbert, D. Mueller-Dombois, R.H.
Riley, and P.M. Vitousek. Changes in soil phosphorus and ecosystem dynamics across a long soil chronosequence in Hawai’i. Ecology 76: 1407–
1424.
DeFries, R.S., C.B. Field, I. Fung, C.O. Justice, S. Los, P.A. Matson, E.
Matthews, H.A. Mooney, C.S. Potter, K. Prentice, P.J. Sellers, J.R.G.
Townshend, C.J. Tucker, S.L. Ustin, and P.M. Vitousek. Mapping the land surface for global atmosphere-biosphere models: Toward continuous distributions of vegetation's functional properties. Journal of Geophysical
Research 100: 20,867–20,882.
Holland, E.A., A.R. Townsend, and P.M. Vitousek. Variation in temperature regulation of CO
2
fluxes and N mineralization from five Hawaiian soils:
Implications for a changing climate. Global Change Biology 1: 115–123.
Kitayama, K., D. Mueller-Dombois, and P.M. Vitousek. Primary succession of the Hawaiian montane rainforest on a chronosequence of eight lava flows.
Journal of Vegetation Science 6: 211–222.

Koch, G., P.M. Vitousek, W.T. Steffen, and B.H. Walker. Terrestrial transects for global change research. Vegetatio 121: 53–65.
Lohse, K.A., D. Nullet, and P.M. Vitousek. The effects of an extreme drought on the vegetation of a single lava flow on Mauna Loa, Hawaii. Pacific Science
49: 212–220.
Matson, P.A., and P.M. Vitousek. Biosphere-atmosphere interactions in a tropical deciduous forest ecosystem. p. 384–398. In Bullock, S.H., H.A. Mooney, and
E. Medina (eds.), Seasonally Dry Tropical Forests . Cambridge University
Press, New York.
Riley, R.H. and P.M. Vitousek. Nutrient dynamics and trace gas flux during ecosystem development in Hawaiian montane rainforest. Ecology 76: 292–
304.
Townsend, A.R., P.M. Vitousek, and S.E. Trumbore. Temperature controls over long-term soil organic matter dynamics. Ecology 76: 721–733.
Vitousek, P.M. The Hawaiian Islands as a model system for ecosystem studies.
Pacific Science 49: 2–16.
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1996 Potter, C.S., P.A. Matson, P.M. Vitousek, and E.A. Davidson. Process modelling of controls on nitrogen trace gas emissions from soils world-wide . Journal of
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Raich, J.W., A.E. Russell, T.E. Crews, H. Farrington, and P.M. Vitousek. Both nitrogen and phosphorus limit plant production on young Hawaiian lava flows.
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Vitousek, P.M., C.M. D’Antonio, L.L. Loope, and R. Westbrooks. Biological invasions as global environmental change. American Scientist 84: 468–478.
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1997 Daily, G.C., P.A. Matson, and P.M. Vitousek. Ecosystem services supplied by soil. p. 113–132. In Daily, G.C. (ed .), Nature's Services: Societal
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Malmstrom, P.M. Vitousek, G.J. Collatz, P.J. Sellers, D.A. Randall, A.S.
Denning, F. Badeck, and J. John. Carbon-13 exchange between the atmosphere and biosphere. Global Biogeochemical Cycles 11: 507–533.
Hooper, D.U., and P.M. Vitousek. Plant composition and diversity effects on ecosystem processes. Science 277: 1302–1305.
Raich, J.W., A.E. Russell, and P.M. Vitousek. Primary production and ecosystem development along an elevational gradient in Hawaii. Ecology 78: 707–721.
Russell, A.E., and P.M. Vitousek. Decomposition and potential nitrogen fixation in Dicranopteris linearis litter on Mauna Loa, Hawai’i, USA. Journal of
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Thompson, M.V., and P.M. Vitousek. Asymbiotic nitrogen fixation and decomposition during long-term soil development in Hawaiian montane rain forest. Biotropica 29: 134–144.

Torn, M.S., S.E. Trumbore, O.A. Chadwick, P.M. Vitousek, and D.M. Hendricks.
Mineral control of soil carbon storage and turnover. Nature 389: 170–173.
Townsend, A.R., P.M. Vitousek, D.J. Des Marais, and A. Tharpe. Effects of temperature and soil carbon pool structure on CO
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Vitousek, P.M. On regression and residuals: response to Knops et al. Oecologia
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Vitousek, P.M. Stopping biological invasions before they do lasting damage.
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Vitousek, P.M. The ecosystems of Mauna Loa Volcano, Hawaii. Natural History
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Vitousek, P.M., and H. Farrington. Nutrient limitation and soil development: experimental test of a biogeochemical theory. Biogeochemistry 37: 63–75
Vitousek, P.M., J.D. Aber, R.W. Howarth, G.E. Likens, P.A. Matson, D.W.
Schindler, W.H. Schlesinger, and D. Tilman. Human alteration of the global nitrogen cycle: sources and consequences. Ecological Applications 7: 737–
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Vitousek, P.M., O.A. Chadwick, T. Crews, J. Fownes, D. Hendricks, and D.
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Vitousek, P.M., C.M. D’Antonio, L.L. Loope, M. Rejmanek, and R. Westbrooks.
Introduced species: a significant component of human-caused global change .
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1998 Aplet, G.H., R.F. Hughes, and P.M. Vitousek. Ecosystem development on
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Aplet, G.H., R.L. Loh, J.T. Tunison and P.M. Vitousek. Experimental restoration of a closed Faya tree stand. Technical Report 121, Cooperative National Park
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Austin, A.T., and P.M. Vitousek. Nutrient dynamics on a precipitation gradient in
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Cordell, S., G. Goldstein, D. Mueller-Dombois, D. Webb, and P.M. Vitousek.
Physiological and morphological variation in Metrosideros polymorpha , a dominant Hawaiian tree species, along an altitudinal gradient: the role of phenotypic plasticity. Oecologia 113: 188–196.
D’Antonio, C.M., R.F. Hughes, M. Mack, D. Hitchcock, and P.M. Vitousek. The response of native species to removal of invasive exotic grasses in a seasonally-dry Hawaiian woodland. Journal of Vegetation Science 9 : 699–
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Freifelder, R., P.M. Vitousek, and C.M. D’Antonio. Microclimate change and effect on fire following forest-grass conversion in a seasonally dry tropical woodland. Biotropica 30 : 286–297.
Hooper, D.U., and P.M. Vitousek. Effects of plant composition and diversity on nutrient cycling in serpentine grassland. Ecological Monographs 68: 121–149.
Kennedy, M.J., O.A. Chadwick, P.M. Vitousek, L.A. Derry, and D.M. Hendricks.
Replacement of weathering with atmospheric sources of base cations during ecosystem development, Hawaiian Islands. Geology 26 : 1015–1018.
Loh, R.K., J.T. Tunison, L.R. Walker and P.M. Vitousek. Population growth and nitrogen-fixation of faya tree, 1986–1993. Technical Report 118, Cooperative
National Park Resources Study Unit, University of Hawaii.
Russell, A.E., J.W. Raich, and P.M. Vitousek. The ecology of the climbing fern,
Dicranopteris linearis , on windward Mauna Loa, Hawai’i. Journal of Ecology
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Tanner, E.V.J., P.M. Vitousek, and E. Cuevas. Experimental investigation of nutrient limitation of forest growth on wet tropical mountains. Ecology 79:
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Vitousek, P.M. The structure and functioning of montane tropical forests: control by climate, soils, and disturbance. Ecology 79: 1–2.
Vitousek, P.M. Foliar and litter nutrients, nutrient resorption, and decomposition in Hawaiian Metrosideros polymorpha . Ecosystems 1 : 401–407.
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1999 Chadwick, O.A., L.A. Derry, P.M. Vitousek, B.J. Huebert, and L.O. Hedin.
Changing sources of nutrients during four million years of ecosystem development. Nature 397 : 491–497.
Ewel, J.J., D.J. O’Dowd, J. Bergelson, C.C. Daehler, C.M. D’Antonio, L.D.
Gomez, D.R. Gordon, R.J. Hobbs, A. Holt, K.R. Hopper, C.E. Hughes, M.
LaHart, R.R.B. Leakey, W.G. Lee, L.L. Loope, D.H. Lorence, S.M. Louda,
A.E. Lugo, P.B. McEvoy, D.M. Richardson, and P.M. Vitousek. Deliberate introductions of species: research needs. Bioscience 49 : 619–630.
Herbert, D.A., J.H. Fownes, and P.M. Vitousek. Hurricane damage and recovery of a Hawaiian forest: effects of increased nutrient availability on ecosystem resistence and resilience. Ecology 80 : 908–920.
Huebert, B., P. Vitousek, J. Sutton, T. Elias, J. Heath, S. Coeppicus, S. Howell, and B. Blomquist. Volcano fixes nitrogen into plant-available forms.
Biogeochemistry 47 : 111–118.
Kurina, L.M., and P.M. Vitousek. Controls over the accumulation and decline of a nitrogen fixing lichen, Stereocaulon vulcani , on young Hawaiian lava flows.
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Martinelli, L.A., M.C. Piccolo, A.R. Townsend, and P.M. Vitousek. Nitrogen stable isotope composition of leaves and soil: tropical versus temperate forests.
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Matson, P.A., W.H. McDowell, A.R. Townsend, and P.M. Vitousek. The globalization of N deposition: ecosystem consequences in tropical environments. Biogeochemistry 46 : 67–83.
Vitousek, P.M. Nutrient limitation to nitrogen fixation in young volcanic sites.
Ecosystems 2 : 505–510.
Vitousek, P.M., and C.B. Field. Ecosystem constraints to symbiotic nitrogen fixers: a simple model and its implications. Biogeochemistry 46 : 179–202.
Vitousek, P.M., M.J. Kennedy, L.A. Derry, and O.A. Chadwick. Weathering versus atmospheric sources of strontium in ecosystems on young volcanic soils. Oecologia 121 : 255–259.

2000 Austin, A. T. and P. M. Vitousek. Precipitation, decomposition and decomposability of Metrosideros polymorpha in native forests on Hawai'i.
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Blackmore, M., and P.M. Vitousek. Cattle grazing, forest loss, and fuel loading in a Hawaiian dry forest at Pu'u Wa'aWa'a ranch, Hawai'i. Biotropica 32 : 625–
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Chapin, F.S. III, E.S. Zavaleta, V.T. Eviner, R.L. Naylor, P.M. Vitousek, S.
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Crews, T.E., H. Farrington, and P.M. Vitousek. Changes in asymbiotic, heterotrophic nitrogen fixation on leaf litter of Metrosideros polymorpha with long-term ecosystem development in Hawaii. Ecosystems 3 : 386–395.
Hättenschwiler, S., and P.M. Vitousek. The role of polyphenols in terrestrial ecosystem nutrient cycling. Trends in Ecology and Evolution 15 : 238–243.
Hobbie, S.E., and P.M. Vitousek. Nutrient regulation of decomposition in
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Hotchkiss, S.C., P.M. Vitousek, O.A. Chadwick, and J.P. Price. Climate cycles, geomorphological change, and the interpretation of soil and ecosystem development. Ecosystems 3 : 522–533.
Neff, J.C., S.E. Hobbie, and P.M. Vitousek. Nutrient and minerological control on dissolved organic C, N and P fluxes and stoichiometry in Hawaiian soils.
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Olander, L.P., and P.M. Vitousek. Regulation of soil phosphatase and chitinase activity by N and P availability. Biogeochemistry 49 : 175–190.
Raich, J.W., W.J. Parton. A.E. Russell, R.L. Sanford Jr., and P.M. Vitousek.
Analysis of factors regulating ecosystem development on Mauna Loa using the
Century model. Biogeochemistry 51 : 161–191.
Riley, R.H., and P.M. Vitousek. Hurricane effects on nitrogen trace gas emissions in Hawaiian montane rainforest. Biotropica 32 : 751–756.
Scowcroft, P.G., D.R. Turner, and P.M. Vitousek. Decomposition of polymorpha leaf litter along elevational gradients in Hawaii. Global Change
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Cordell, S., G. Goldstein, F.C Meinzer, and P.M Vitousek. Morphological and physiological adjustment to N and P fertilization in nutrient-limited
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Cordell, S., G. Goldstein, F.C Meinzer, and P.M Vitousek. Regulation of leaf lifespan and nutrient-use efficiency of Metrosideros polymorpha trees at two extremes of a long chronosequence in Hawaii. Oecologia 127 : 198–206.
Crews, T.E., L.M. Kurina, and P.M. Vitousek. Organic matter and nitrogen accumulation and nitrogen fixation during early ecosystem development in
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D’Antonio, C.M., R.F. Hughes, and P.M. Vitousek. Factors influencing dynamics of two invasive C
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Harrington, R.A., J.H. Fownes, and P.M. Vitousek. Production and resource-use efficiencies in N- and P-limited tropical forest ecosystems. Ecosystems 4 : 646–
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Kurina, L.M., and P.M. Vitousek. Nitrogen fixation rates at Stereocaulon vulcani on young Hawaiian lava flows. Biogeochemistry 55 : 179–194.
Pearson, H.L., and P.M. Vitousek. Symbiotic nitrogen fixation in regenerating stands of Acacia koa , a tropical leguminous tree. Ecological Applications 11 :
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Rastetter, E.B., P.M. Vitousek, C. Field, G.R. Shaver, D. Herbert, and G.A.
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Restrepo, C., and P.M. Vitousek. Landslides, alien species, and the diversity of a
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Treseder, K.K., and P.M. Vitousek. Effects of soil nutrient availability on investment in acquisition of N and P in Hawaiian rain forests. Ecology 82 :
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Treseder, K.K., and P.M. Vitousek. Potential ecosystem-level effects of genetic variation among populations of Metrosideros polymorpha from a soil fertility gradient in Hawaii. Oecologia 126 : 266–275.
Vitousek, P.M., and C.B. Field. Input-output balances and nitrogen limitation in terrestrial ecosystems. Pp. 217–235 in Schulze E.D., S.P. Harrison, M.
Heimann, E.A. Holland, J. Lloyd, I.C. Prentice, and D. Schimel (eds), Global
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2002
Benning, T., D. LaPoint, C. Atkinson, and P.M. Vitousek. Interactions of climate change with biological invasions and land use in the Hawaiian Islands:
Modeling the fate of endemic birds using a geographic information system.
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Pearson, H.L., and P.M. Vitousek. Nitrogen and phosphorus dynamics and symbiotic nitrogen fixation across a substrate-age gradient in Hawaii.
Ecosystems 5 : 587-596.
Vitousek, P.M. Oceanic islands as model systems for ecological studies. of Biogeography 29 : 573–582.
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2003
Goodale, C.L., J.D. Aber, and P.M. Vitousek. Changes in streamwater chemistry on Mt. Moosilauke and Mt. Washington, New Hampshire between 1973–74 and 1996–97. Ecosystems 6 : 75–86.
Hättenschwiler, S., A.E. Hagerman, and P.M. Vitousek. Polyphenols in litter from tropical montane forests across a wide range in soil fertility. Biogeochemistry
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Hedin, L.O., P.M. Vitousek, and P.A. Matson. Pathways and implications of nutrient losses during four million years of tropical forest ecosystem development. Ecology 84 : 2231–2255.
Hungate, B.A., R.J. Naiman, M. Apps, J.J. Cole, B. Moldan, K. Satake, J.
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Matzek, V.A., and P.M. Vitousek. Nitrogen fixation in bryophytes, lichens, and decaying wood along a soil age gradient in Hawaiian montane rainforest.
Biotropica 35 : 12–19.
Neff, J.C., F.S. Chapin III, and P.M. Vitousek. Leaks and short-circuits in the nitrogen cycle: Dissolved organic nitrogen and its role in terrestrial ecosystems.
Frontiers in Ecology and Environmental Science 1 : 205–211.
Restrepo, C., P. Vitousek, and P. Neville. Landslides significantly alter land cover and the distribution of biomass: an example from the Ninole Ridges of
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Vitousek, P.M. Stoichiometry and flexibility in the Hawaiian model system. p.
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A. Luers, E. Mecking, V. Monastra, and S. Porder. Erosion and the rejuvenation of weathering-derived nutrient supply in an old tropical landscape.
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2004
Allison, S.D., and P.M. Vitousek. Extracellular enzyme activities and carbon chemistry as drivers of tropical plant litter decomposition. Biotropica 36 : 285–
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Allison, S.D., and P.M. Vitousek. Decomposition and nutrient dynamics of native and exotic Hawaiian understory plants. Oecologia 141 : 612–619.
Kirch, P.V., A.S. Hartshorn, O.A. Chadwick, P.M. Vitousek, D.R. Sherrod, J.
Coil, L. Holm, and W.D. Sharp. Environment, agriculture, and settlement patterns in a marginal Polynesian landscape. Proc. Natl. Acad. Sci. USA 101 :
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Olander, L.P., and P.M. Vitousek. Biological and geochemical sinks for phosphorus in a wet tropical forest soil. Ecosystems 7 : 404–419.
Rothstein, D.E., P.M. Vitousek, and B.L Simmons. An exotic tree accelerates decomposition and nutrient turnover in a Hawaiian montane rainforest.
Ecosystems 7 : 805–814 .
Vitousek, P.M. Nutrient Cycling and Limitation: Hawai‘i as a Model System.
Princeton University Press.
Vitousek, P.M., T.L. Ladefoged, P.V. Kirch, A.S. Hartshorn, M.W. Graves, S.C.
Hotchkiss, S. Tuljapurkar, and O.A. Chadwick. Agriculture, soils, and society in precontact Hawai’i. Science 304 : 1665–1669.
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Allison, S.D., and P.M. Vitousek. Responses of extracellular enzymes to simple and complex nutrient inputs. Soil Biology and Biochemistry 37: 937-944
Asner, G.P., and P.M. Vitousek. Remote analysis of biological invasion and biogeochemical change. Proc. Natl. Acad. Sci. USA 102 : 4383–4386.
Asner, G.P., A.J. Elmore, R.F. Hughes, A.S. Warner, and P.M. Vitousek. 2005.
Ecosystem structure along bioclimatic gradients in Hawaii from imaging spectroscopy. Remote Sensing of Environment 96 :497-508.
Goodale, C.L., J.D. Aber, P.M. Vitousek, and W.H. McDowell. Long-term decreases in stream nitrate: successional causes unlikely, possible links to
DOC? Ecosystems 8 : 334-337.
Henry, H.A.L., E.E. Cleland, C.B. Field, and P.M. Vitousek. Interactive effects of elevated CO
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, N deposition and climate change on plant litter quality in a
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Henry, H.A.L, J.D. Juarez, C.B. Field & P.M. Vitousek.
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Change Biology 11 : 1808-1815.
Kirch, P.V., J. Coil, A. S. Hartshorn, M. Jeraj, P. M. Vitousek, and O. A.
Chadwick. Intensive Dryland Farming on the Leeward Slopes of Haleakala,
Maui, Hawaiian Islands: Archaeological, Archaeobotanical, and Geochemical
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Ladefoged, T., C. Stevenson, P. Vitousek, and O.A. Chadwick. Soil nutrient depletion and the collapse of Rapa Nui society. Rapa Nui Journal 19 : 100-
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Olander, L.P., and P.M. Vitousek. Short-term controls over inorganic phosphorus during soil and ecosystem development. Soil Biol. Biochem. 37 : 651–659.
Parton, W.J., J. Neff, and P.M. Vitousek. Modelling phosphorus, carbon, and nitrogen dynamics in terrestrial ecosystems. Pp. 325–327 in Turner, B.L., E.
Frosserd, and D.S. Baldwin (eds.), Organic Phosphorus in the Environment .
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Porder, S.J., P.M. Vitousek, and G. Asner. Ground-based and remotely-sensed nutrient availability across a tropical landscape. PNAS 102 : 10909-10912.
Porder, S., A. Paytan, and P.M. Vitousek. Erosion and landscape development affect plant nutrient status in the Hawaiian Islands. Oecologia 142 : 440–449.
Torn, M.S., P.M. Vitousek, and S.E. Trumbore. The influence of nutrient availability on soil organic matter turnover estimated by incubating and radiocarbon modeling. Ecosystems 8 : 352-372.
2006
Walther, G.-R., L. Hughes, P. Vitousek, and N.C. Stenseth. Consensus on climate change. TREE 20 : 648-649.
Wiegand, B.A., O.A. Chadwick, P.M. Vitousek, and J.L. Wooden. Ca cycling and isotopic fluxes in forested ecosystems in Hawaii. Geophysical Research
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Asner, G.P., R.E. Martin, K.M. Carlson, U. Rascher, and P.M. Vitousek.
Vegetation-climate interactions among native and invasive species in Hawaiian rainforest. Ecosystems 9:1106-1117.
Goldstein, J.H., G.C. Daily, J.B. Friday, P.A. Matson, R.L. Naylor, and P.M.
Vitousek. Business strategies for conservation on private lands: koa forestry as a case study. PNAS 103: 10140-10145.
Hartshorn, A.S., O. A. Chadwick, P.M. Vitousek, and P.V. Kirch. Prehistoric agricultural depletion of soil nutrients in Hawaii. PNAS 103 : 11092-11097.
Henry, H.A.L., N.R. Chiariello, P.M. Vitousek, H.A. Mooney & C.B. Field.
Interactive effects of fire, elevated CO2 and N deposition on a California annual grassland. Ecosystem s 9 : 1066-1075.

Lee, C., S. Tuljapurkar, and P. Vitousek. Risky business: temporal and spatial variation in pre-industrial dryland agriculture. Human Ecology 34 : 739-763.
Matson, P.A., and P.M. Vitousek. Agricultural intensification: Will land spared from farming be land spared for nature? Conservation Biology 20 : 709-710.
Porder, S., D.A. Clark, and P.M. Vitousek. Persistence of rock-derived nutrients in the wet tropical forests of La Selva, Costa Rica. Ecology 87 : 594-602.
Raich, J.W., A.E. Russell, K. Kitayama, W.J. Parton, and P.M. Vitousek.
Temperature influences carbon accumulation in moist tropical forests. Ecology
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Vitousek, P.M. Ecosystem science and human-environment interactions in the
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2007
Benner, J.W., S. Conroy, C.K. Lunch, N. Toyoda, and P.M. Vitousek. Phosphorus fertilization increases the abundance and nitrogenase activity of the cyanolichen Pseudocyphellaria crocata in Hawaiian montane forests.
Biotropica 39: 400-405.
Benner, J., and P.M. Vitousek. Development of a diverse epiphyte community in response to phosphorus fertilization. Ecology Letters .
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Chadwick, O.A., E.F. Kelly, S.C. Hotchkiss, and P.M. Vitousek. Precontact vegetation and soil nutrient status in the shadow of Kohala Volcano, Hawaii.
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Funk, J. and P.M. Vitousek. Resource use efficiency and plant invasion in lowresource systems. Nature 446 : 1078-1081.
Meyer, M., T. Ladefoged, and P.M. Vitousek. Soil phosphorus and agricultural development in the Leeward Kohala Field System. Pacific Science 61 : 347-
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Kirch PV, Chadwick OA, Tuljapurkar S, Ladefoged TN, Graves MW, Hotchkiss
S, Vitousek PM. Human ecodynamics in the Hawaiian ecosystem, 1200-200
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Porder, S., P.M. Vitousek, O.A. Chadwick, C.P. Chamberlain, and G.E. Hilley.
Uplift, erosion, and phosphorus limitation in terrestrial ecosystems.
Ecosystems 10 : 158-170.
Vitousek, P.M. Islands and global environmental change. p. 99-103, in Wai ki
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Amatangelo, K.L., and P.M. Vitousek. Fern phylogeny and nutrient stoichiometry. Oecologia 157 : 619-627
Anderson-Teixeira, K.J., P.M. Vitousek, and J.H. Brown. Positive feedbacks drive exceptional temperature dependence in ecosystems developing on the lava flows of Mauna Loa, Hawaii. PNAS 105 : 228-233
Asner, G.P., R.F. Hughes, P.M. Vitousek, D.E. Knapp, T. Kennedy-Bowdoin, J.
Boardman, R.E. Martin, M. Eastwood, and R.O. Green. Invasive plants transform the 3-D structure of rainforests. PNAS 105 : 4519-4523.
Houlton, B.Z., Y.-P. Wang, P.M. Vitousek, and C.B. Field. A unifying framework for di-nitrogen (N
2
) fixation in the land biosphere. Nature 454 :
327-330.
Kurten, E.L., C.P. Snyder, T. Iwata, and P.M. Vitousek. Morella cerifera invasion and nitrogen cycling on a lowland Hawaiian lava flow. Biological
Invasions 10 : 19-24.
2009
Amatangelo, K.A., and P.M. Vitousek. Contrasting predictors of fern versus angiosperm decomposition in a common garden. Biotropica 41 : 154-161.
Chadwick, O.A., L.A. Derry, C.R. Bern, and P.M. Vitousek. Sources of strontium to soil minerals and ecosystems across the Hawaiian Islands. Chemical
Geology doi:10.1016/jchemgeo.2009.01.009.
In press
Benner, J., P.M. Vitousek, and R. Ostertag. Nutrient cycling in tropical montane cloud forest. In Bruijnzeel, L.A., J. Juvik, and F. Scatena (eds.), Tropical
Montane Cloud Forests .
Busby, P.E., P. Vitousek, and R. Dirzo. Prevalence of tree regeneration by sprouting and seeding along a rainfall gradient in Hawai`i. Biotropica

Matzek, V., and P.M. Vitousek. N:P stoichiometry and protein:RNA ratios in vascular plants: an evaluation of the growth rate hypothesis. Ecology Letters .
Palmer, M.A., M. Graves, T.N. Ladefoged, O.A. Chadwick, T.K. Duarte, S.
Porder, and P.M. Vitousek. Sources of nutrients to windward agricultural systems in pre-contact Hawai`i. Ecological Applications .
Robertson, G.P., and P.M. Vitousek. Nitrogen in Agriculture: Balancing the Cost of an Essential Resource. Annual Review of Environment and Resources.
Vitousek, P.M., and P.A. Matson. Nutrient cycling and biogeochemistry. In:
Loreau, M. (editor), Ecology. Princeton University Press.
Vitousek, P.M., G.P. Asner, O.A. Chadwick, and S.C. Hotchkiss. Landscapelevel variation in forest structure and biogeochemistry across a substrate age gradient in Hawai`i. Ecology .
Vitousek, P.M., R.L. Naylor, T. Crews, M.B. David, L.E. Drinkwater, E. Holland,
P. Johnes, J. Katzenberger, L.A. Martinelli, P.A. Matson, G. Nziguheba, D.
Ojima, C. A. Palm, G.P. Robertson, P.A. Sanchez, A.R. Townsend, and F.-S.
Zhang. Nutrient imbalances in agricultural development. Science.
Vitousek, P.M., S. Porder, B.Z. Houlton, and O.A. Chadwick. Terrestrial phosphorus limitation: mechanisms, implications, and nitrogen-phosphorus interactions. Ecological Applications .
Submitted
Ladefoged. T.N., P.V. Kirch, S.O. Gon III, O.A. Chadwick, A.S. Hartshorn, and
P.M. Vitousek. Opportunities and constrains for intensive agriculture in the
Hawaiian Archipelago prior to European contact. Journal of Archaeological
Science.
Ladefoged, T.N., P.V. Kirch, O.A. Chadwick, S.M. Gon III, A.S. Hartshorn, and
P.M. Vitousek. Hawaiian agro-ecosystems and their spatial distribution. In
P.V. Kirch, Roots of Conflict: Soils, Agriculture, and Sociocultural
Complexity in Ancient Hawai`i. School of Advanced Research Press, Santa
Fe, N.M.
Vitousek, P.M., O.A. Chadwick, A.S. Hartshorn, and S.C. Hotchkiss. The “model system” approach and its application to understanding agriculture and intensification in Hawai`i. In P.V. Kirch, Roots of Conflict: Soils, Agriculture, and Sociocultural Complexity in Ancient Hawai`i. School of Advanced
Research Press, Santa Fe, N.M.

Vitousek, P.M., M.A. Tweiten, J. Kellner, S.C. Hotchkiss, O.A. Chadwick, and
G.P. Asner. Top-down analyses of forest structure and biogeochemistry across
Hawaiian landscapes. Pacific Science
Professional Service
Editorial Boards (various times):
American Naturalist , Annual Review of Ecology and Systematics ,
Biogeochemistry , Biotropica , Climatic Change , Conservation Ecology ,
Ecology / Ecological Monographs , Ecosystems (Current) , Global Biogeochemical
Cycles , Oecologia .
NASA Life Sciences Strategic Planning Study Committee, 1986-1988
National Science Foundation (NSF) Task Force on Biological, Behavioral, and
Social
Sciences in the 21st Century, 1990–1991
Coordinating Panel on Terrestrial Ecosystem/Climate Interactions, International
Geosphere-Biosphere Program (IGBP), 1988–1990
Steering Committee Member for IGBP Core Project Global Change in Terrestrial
Ecosystems; Activity Leader for Biogeochemistry, 1990–1993
`Alala (Hawaiian Crow) Recovery Team member, U.S. Fish and Wildlife Service,
1993–
NRC Panel “Valuation of Biodiversity,” 1995–1996
NRC Board on Global Change, 1995–1998
NSF Ecosystem Studies Panel, 2000–2003
Co-Director, First Nations Futures Institute (Consortium of Stanford, Kamehameha
Schools, and the Ngai Tahu Tribe) 2005-
Board of the National Park and Conservation Association, 2009-
(revised 1/16/09)