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CURRICULUM VITAE
PERSONAL:
Name: Sergio M. Pellis
Born: January 16, 1953, Trieste, Italy. Australian citizen,
Canadian citizen
Marital Status: Married
Present position: Professor
Department of Neuroscience,
University of Lethbridge,
Lethbridge, Alberta, Canada T1K 3M4
Telephone: office: (403) 329-2078, lab: (403) 380 1811
E-mail: pellis@uleth.ca
FAX: (403) 329 2775
EDUCATIONAL HISTORY:
1971-1974. Monash University, Melbourne, Victoria, Australia, B.Sc. (Majors in Zoology
and Genetics)
1975. Monash University, Diploma of Education
1976. Monash University, B.Sc. (Honours) (Zoology)
1977-1980. Monash University, Ph. D. (Zoology/Ethology).
Title of Ph. D. Thesis: A Developmental and Comparative Study in the Understanding of
Play Behaviour (on play in Australian Magpies (Gymnorhina tibicen) and Oriental SmallClawed Otters (Amblonyx cinerea))
Supervisor: Dr. John E. Nelson
Examiners: Drs. Ilan Golani and John F. Eisenberg
SPECIALIZED TRAINING & PROFESSIONAL HISTORY:
2011-present. Associate Chair (Curriculum), Department of Neuroscience, The
University of Lethbridge, Lethbridge, Alberta, Canada
2007-present Board of Governors’ Research Chair
2007-2008 Acting Director of Canadian Centre for Behavioural Neuroscience
2005-2006. Associate Chair, Department of Neuroscience, The University of Lethbridge,
Lethbridge, Alberta, Canada
2002-2005. Chair, Department of Psychology and Neuroscience, The University of
Lethbridge, Lethbridge, Alberta, Canada
2005-present. Professor, Department of Neuroscience, The University of Lethbridge,
Lethbridge, Alberta, Canada
1997-2005. Professor, Department of Psychology and Neuroscience, The University of
Lethbridge, Lethbridge, Alberta, Canada
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1993-1997. Associate Professor, Department of Psychology, The University of
Lethbridge, Lethbridge, Alberta, Canada
1990-1993.
Assistant Professor, Department of Psychology, The University of
Lethbridge, Lethbridge, Alberta, Canada
November 1990. Invited participant in a workshop on "The Neural Bases of Aggressive
Behavior: Appraisals of Our Current Insights and Methods", sponsored by The Harry
Frank Guggenheim Foundation. This included selected short courses on modern
Neuroscience techniques for behavioral research. Seville, Spain
July 1988. Invited participant in a workshop on "The Ethoexperimental Analysis of
Behavior", sponsored by NATO (Advanced Study Institute). Pisa, Italy
1985-1990. Assistant Research Scientist, Department of Psychology, University of
Florida, Gainesville, Florida, USA
1982-1985. Post-doctoral Research Associate with Dr. Philip Teitelbaum, Department of
Psychology, University of Illinois at Urbana- Champaign, Illinois, USA
October 1984 -July 1985. Advanced training in the use of Eshkol-Wachman Movement
Notation (EWMN) for the analysis of animal movement taken at Tel Aviv University,
Ramat Aviv, Israel, under the supervision of Prof. N. Eshkol (the originator of the system)
February 1983. Course in high-speed motion picture photography taken at Sarasota,
Florida, from Visual Data Systems
August 1982. Course in Eshkol-Wachman Movement Notation taken at Tel Aviv
University, Ramat Aviv, Israel, under the supervision of Prof. N. Eskhol
1980-1982. Post-doctoral research in Ethology, Department of Zoology, Monash
University, Melbourne, Australia (with Dr. John Nelson)
TEACHING:
Graduate level
University of Florida
1986-1989. John Pierce, Ph.D. (Comparative Psychology) (Committee)
1987-1990. Stephen Taylor, Ph.D. (Comparative Psychology) (Committee)
1988-1990. Jennifer Porter, M. Sc. (Physiological Psychology) (Committee)
University of Lethbridge
Post-Doctoral Research Fellows
2000-2002. Susan Lingle (self-funded, supplemented with my NSERC award)
2001-2004. Joanna Komorowska (supported by my CSN award)
2002-2005. Susan Lingle (supported by a Post-doctoral Fellowship from the Alberta
Ingenuity Fund)
1998-2000. Karen Dean (supported by my NSERC grant)
N.B. Graduate degree programs only began at UL in 1994 (M. Sc., 1994, Ph. D, 2001).
The Honours program began in 1998.
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Graduate Student Supervisor
1994-1996. Evelyn Field, M.Sc. (Neuroscience)*
1994-1996. Lori Smith, M.Sc. (Neuroscience)
1997-1997. Linda Ganz , M. Sc. (Management) (Substitute Supervisor)
1997-1997. Su-Lin Fantella, M. Sc. (Neuroscience) (withdrew due to illness)
1998-2000. Andrew Iwaniuk, M. Sc. (Neuroscience)
1999-2002. Afra Foroud, M. Sc. (Neuroscience).
2003-2005. Christine Reinhart, M. Sc. (Neuroscience)**
2001-2006. Evelyn Field, Ph. D (Neuroscience)*
2003-2006. Cheryl Aleris, M. Sc. (Neuroscience)
2004-2006. Walter Espinoza M. Sc. (Neuroscience)
2005-2008. Christine Reinhart, Ph. D (Evolution & Behaviour)
2006-2008. Heather Bell, M. Sc. (Neuroscience)
2007-2007. Erin Hutton, MA (Applied Behavioral Analysis, St. Cloud State University)
2008-present. Heather Bell, Ph. D (Evolution & Behaviour)
2009-present. Brett Himmler, M. Sc. (Neuroscience)
2011-present. Kerri Norman, M. Sc. (Psychology)
Graduate Supervisory Committees
1994-1996. Brenda Coles, M. Sc. (Neuroscience)
1997-1999. JoAnne Golden, M. Sc. (Biological Sciences)
1997-2000. Lynne Paterson, M. Sc. (Biochemistry)
1998-2000. Katrina White, M. Sc. (Biological Sciences)
1999-2001. Ryan Sleik, M. Sc. (Kinesiology)
2000-2001. Sho Shirakashi, M. Sc. (Biological Sciences)
2000-2002. Melody Polych, M. Sc. (Neuroscience)
2001-2002. Nicole McKenzie, M. Sc. (Kinesiology)
2001-2002. Nissa Steen, M.Sc. (Psychology) (withdrew)
2001-2003. Katheryn Holgate M. Sc. (Biological Sciences)
2001-2003. Erica Hastings, M. Sc. (Neuroscience)
2002-2004. Dustin Hines, M.Sc. (Neuroscience)
2002-2004. Trevor McGill, M. Sc. (Neuroscience)
2002-2006. Jonathon Doan, Ph. D (Neuroscience)
2003-2005. Jennifer Burke, M. Sc. (Biological Sciences)
2003-2005. Stephanie Cooper, M. Sc. (Neuroscience)
2004-2008. Trevor McGill, Ph. D (Neuroscience)
2004-2007. Robin Goldsbury, M. Sc. (Neuroscience)
2005-2006. Sarah Wheatley, M. Sc. (Psychology) (Withdrew)
2005-2009. Shannon Digwood, Ph. D (Evolution & Behaviour)
2007-2008. Robin Goldsbury, Ph. D (Neuroscience) (Withdrew)
2008-present. Doug Vanderlaan Ph. D (Evolution & Behaviour)
2008-present. Nicola Forshaw, Ph. D (Evolution & Behaviour)
2008-2010. Petra McDougall, M. Sc. (Psychology)
2008-present. John Hindley, Ph. D (Evolution & Behaviour)
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2008-2011. Matt Merrifield, M. Sc. (Biological Sciences)
2008-2010. Megan Metzler, M. Sc. (Neuroscience)
2009-2010. Kurt Smith, M. Sc. (Kinesiology)
2009-2010. Andrea Rutledge, M. Sc. (Biological Sciences) (Withdrew)
2009-present. Sonja Richards, M. Sc. (Neuroscience)
2009-2011. Linda Lait, M. Sc. (Biological Sciences)
2009-present. April Takahashi, M. Sc. (Psychology)
2011-present. Deanna Forrester, Ph. D. (Evolution & Behaviour)
2011-present. Alice Martin, Ph. D. (Evolution & Behaviour)
*Won the Gold Medal for best graduate thesis of the year
**Won the Medal of Merit for the best M. Sc thesis of the year
Honours theses
1999/2000.
1999/2000.
2000/2001.
2000/2001.
2001/2002.
2002/2003.
2004/2005.
Tamara Pasztor
Takeshi Shimizu
Marvin Hofer
Erica Hastings
Kendra Massie
Christine Reinhart
James Sanders
Undergraduate level
1990-present. (University of Lethbridge) I have had over 29 undergraduate summer
research students working in my laboratory, mostly funded (see p. 7).
1990-present. (University of Lethbridge) Have taught several courses in Psychology and
Neuroscience, ranging from first to fourth year. These include: Introductory Psychology
(1000), Brain and Behaviour (2600), Introduction to Animal Behavior (2700),
Introduction to Statistics in Psychological Research (2400/3010), Multidisciplinary
Approaches to Play (3035), Human Sexuality (3630), Drugs and Behaviour (3650), Sex
and Human Ethology (3730), Motivation (3850), Evolution of Brain & Behaviour (3705),
Movement Analysis of Motor Disorders (4630), Selected Topics in Behavioural
Neuroscience (4600), Evolution and Development of Play Behaviour (4000), Competition,
Cooperation and the Social Brain (4000), and The Psychology and Biology of Emotions
(4000), Making a Playful Brain (4850), Understanding Behaviour (4850). In addition, I
have supervised 111 Independent Studies at the 4990 level, 51 at the 3990 level, and 25 at
the 2990 level. I have also supervised 56 Applied Studies students at the 3000 and 4000
levels.
1988-present. (Monash, Illinois, Florida and Lethbridge Universities) Conducted training
courses for both individuals and groups on the use of the Eshkol-Wachman Movement
Notation (EWMN). These courses have been for undergraduate and graduate students, as
well as postdoctoral and faculty researchers.
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1975-1981. (Monash University) Instructor/Demonstrator for laboratory classes in
General Zoology, Comparative Anatomy and Physiology, Ecology, Evolution, Ethology
and EWMN.
1976-1978. (LaTrobe University, Melbourne, Australia) Instructor/Demonstrator for
laboratory classes in Introductory Zoology
UNIVERSITY COMMITEES:
University of Lethbridge
2011-present. Associate Chair (Curriculum), Department of Neuroscience
2010-2011. Search Committee member for Modern Languages
2009-present. NSERC General Grant/Scholarship Selection Committee
2009-present. Department of Neuroscience STP Committee
2009-2010. Department of Anthropology Search Committee
2009-2010. Department of Anthropology Chair Selection Committee
2008-2010. Ph. D Program Committee
2008-2010. Ph. D. Review Committee
2008-2009. AHFMR Travel Awards Committee
2008-2009. Search Committee for the Academic Writing Programme
2008-present. Michener Post-Doctoral Fellowship in Terrestrial Ecology Awards
Committee
2007-2008. Task Force for Graduate Studies in the Sciences
2007-2008. Chair, STP Committee for the Liberal Education Program
2007-2008. Acting Director, CCBN
2007-2008. Animal Welfare Committee
2005-2006. Chair of the Liberal Education STP Committee
2005-2006. Department of Neuroscience STP Committee
2005-2006. Associate Chair, Department of Neuroscience
2004-2006. School of Graduate Studies Council (new format)
2002-2005. Chair, Department of Psychology & Neuroscience
2002-2003. Roving Alternate for Economics Search Committee
2001-2003. School of Graduate Studies Council
2001-2004. Senate Honorary Degrees Committee
2000-2001. CCBN Planning and Building Committee
2000-2002. Faculty of Arts and Science STP Committee
2000-2001. Department of English Search Committee
1998-1999. Department of Philosophy Search Committee
1998-1999. Faculty of Management STP Committee
1993-2001. NSERC General Grant/Scholarship Selection Committee (Chairperson for
1993-1994, 1996-1997)
1997. Appointment Committee for Associate Vice President (Academic)
1997 and 2001. Grade Appeal Committee
1996-1997. Department of Physical Education Search Committee
1995-1997. 1998-2000. Department of Psychology Search Committee
1995-1996. Department of Sociology, Chair Selection Committee
1995-1997. M.A/M. Sc. Program Committee
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1995-1997. AHFMR Awards Committee
1991-1997. Animal Welfare Committee (Chairperson 1994-1997)
1994-1997. Graduate Studies Committee
1993-1997 and 2003. Department of Psychology, Chair Selection Committee
1993-1994 and 1996-1997. University of Lethbridge Research Committee
1994-1995 and 1997-2003. Department of Psychology, STP Committee
1994-1995. Department of Philosophy, STP Committee
1992-1993. Department of Economics, Chair Selection Committee
1992-1993. Department of Psychology, Curriculum Committee
1992-1993. Advisory Committee for Technical Services
INVOLVEMENT IN THE ACADEMIC COMMUNITYAT LARGE:
2011-present. Academic Editor of PLoS ONE
2010-present. Awards Chair-elect for the American Psychological Association Division 6
(Behavioral Neuroscience and Comparative Psychology)
2008-present. Council member for the International Society of Comparative Psychology
2005-2006. Chair of the ad hoc J. P. Scott Award Committee for the 2006 International
Society for Research on Aggression meeting
2005-present. Member of the Hotchkiss Brain Institute (University of Calgary)
2005-2010. Council member for the International Society for Research on Aggression
2005-present. Consulting Editor, Journal of Comparative Psychology
2004-2009. Associate Editor, Aggressive Behavior
2004-2006. Ingenuity Graduate Studentship Review Committee (AIF)
2000-present. On the editorial board of International Journal of Comparative Psychology
1997-2002. Summer Studentship Applications Advisory Committee (AHFMR)
1994-2002. Council member for the International Society for Research on Aggression
1993-2004. On the editorial board of Aggressive Behavior
Continuing ad hoc reviewer for US and Canadian funding agencies NIH; NSF;
NSERC; CIHR; AHFMR; Binational Science Foundation (Israel-USA); Netherlands
Organization for Scientific Research (Evolution & Behaviour Research Program); Harry
Frank Guggenheim Foundation for Research on Aggression; Thomas F. and Kate Miller
Jeffress Memorial Trust (Bank of America) and for international professional journals
Acta Zoologica; Aggressive Behavior; American Journal of Play; American Journal of
Primatology; Animal Behaviour; Behaviour; Behavioural Brain Research; Behavioral
Neuroscience; Behavioural Processes; Biology of Reproduction; BMC (Ecology);
Biological Psychiatry; Brain Research; Brain, Behavior & Evolution; Central European
Journal of Biology; Developmental Cognitive Neuroscience; Developmental
Psychobiology; Developmental Psychology; Ethology; European Journal of
Neuroscience; Evolution & Human Behavior; Experimental Brain Research; Genes,
Brain & Behavior; Hormones & Behavior; Human Nature; International Journal of
Comparative Psychology; International Journal of Primatology; Journal of Comparative
Psychology; Journal of Ethology; Journal of Motor Behavior; Journal of Neuroscience
Methods; Journal of Zoology; New Zealand Journal of Zoology; Neuroscience &
Biobehavioral Reviews; Pharmacology, Biochemistry & Behavior; Physiology &
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Behavior; PloS ONE; Primates; The Quarterly Review of Biology; Social Neuroscience.
In addition, I am routinely asked to review book chapters and books for colleagues and for
editors.
PROFESSIONAL SOCIETIES:
2001-present. International Society for Comparative Psychology (ISCP)
1985-present. International Society for Research on Aggression (ISRA)
1986-present. Animal Behavior Society (ABS)
1987-present. Society for Neuroscience
1987-present. International Brain Research Organization (IBRO)
1990-present. Canadian Society for Brain, Behaviour, and Cognitive Science
1993-2005. International Society for Human Ethology
1984-1999. Sigma Xi
1986-1997. New York Academy of Sciences
GRANTS AWARDED:
1984-1987. Harry Frank Guggenheim Foundation Research Grant ($US25000 /yr)
1986. Division of Sponsored Research, University of Florida (Co-PI with P. Teitelbaum)
($US7500)
1990. Natural Science & Engineering Research Council Small Grant from the University
of Lethbridge Research Services ($3000)
1991. University of Lethbridge Research Grant ($3200).
1991. Natural Science and Engineering Research Council Equipment Grant ($ 65000)
1991-1993. Natural Science & Engineering Research Council Operating Grant ($25000
/yr)
1992. University of Lethbridge Research Grant ($4000)
1992. Additional Natural Science & Engineering Research Council New Faculty award
from the University of Lethbridge Research Services ($1200)
1994. AHFMR Visiting Lecturer award (Dorothy Paul, University of Victoria, BC)
($1132)
1994-1997. Natural Science & Engineering Research Council Operating Grant
($23000/yr)
1994. AHFMR Visiting Lecturer award (John Nelson, Monash University, Melbourne,
Australia) ($1500)
1994. National Ataxia Foundation (Co-PI with D. P. O’Brien) ($US6000)
1995-1997. National Institutes of Health (Co-PI with D. P. O’Brien) ($US75000/yr)
1995. University of Lethbridge Research Grant ($3704)
1995. AHFMR Visiting Lecturer award (Philip Teitelbaum, University of Florida) ($1459)
1996. AHFMR Visiting Lecturer award (Martin Kavaliers, University of Western Ontario)
($1091)
1996. University of Lethbridge Grant in Lieu of Salary ($5953)
1996. AHFMR Visiting Lecturer award (Robert J. Blanchard, University of Hawaii)
($1455)
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1997. AHFMR Visiting Lecturer award (Joel A. Vilensky, Indiana University Medical
School) ($1469)
1997. University of Lethbridge Research Grant ($4000)
1997. Natural Science & Engineering Research Council Equipment Grant ($30330)
1997. University of Lethbridge Grant in Lieu of Salary ($5806)
1998. AHFMR Travel Grant ($1000)
1998-2003. Natural Science & Engineering Research Council Operating Grant
($27760 /yr)
1999. University of Lethbridge Research Grant ($4300)
2000. Alberta Learning Research Excellence Envelope (Co-PI with Lesley Brown)
($10000)
2000-2001. National Centre of Excellence (Canadian Stroke Foundation) ($22500/yr)
2001-2003. National Centre of Excellence (Canadian Stroke Foundation) ($40000/yr)
2001. Research Infrastructure Support and Grant from the Dean’s office (with A. Hurly,
R. Cartar & C. Goater) ($22,402)
2001. AHFMR Travel Grant ($1,000)
2001. Natural Science & Engineering Research Council Equipment Grant ($59998)
combined with a University of Lethbridge supplement ($17921) (Total: $77919)
2001-2003. Medical Services Incorporated (C0-PI with Lesley Brown) ($85000/year).
2002. AHFMR Visiting Lecturer award (Gordon M. Burghardt, University of Tennessee)
($1006)
2003-2008. Natural Science & Engineering Research Council Operating Grant
($36000/year)
2003. AHFMR Visiting Lecturer award (David Eilam, Tel Aviv University) ($1229)
2006-2013. Norlein Foundation (supports perinatal research), support for a graduate
student ($20000/year)
2008-2013. Natural Science & Engineering Research Council Discovery Grant
($43136/year)
2008. AHFMR Visiting Lecturer award (Jeffrey R. Alberts, University of Indiana)
($1068)
2008. AHFMR Visiting Lecturer award (Michael A. Huffman, Kyoto University) ($1500)
2011-213. Working group to explore “Play as a Window into Cognitive Evolution and the
Rules of Sociality" funded by the National Institute for Mathematical and Biological
Synthesis (NIMBioS) – I am one of 4 PIs with NIMBioS, supporting a total of 15
researchers from around the world to converge at Knoxville, TN, 3-4 times over the two
year tenure of the grant to explore mathematical models applicable for the study of play
and its evolution (approximately $150,000)
FUNDED SUMMER RESEARCH STUDENTS (1991-PRESENT):
Chinook (5)
NSERC (6)
AHFMR (8)
Other (6)
LISTINGS:
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1989. Who's Who in Australasia and the Far East, First Edition, Cambridge, U.K
1989. The International Directory of Distinguished Leadership, Second Edition, American
Biographical Institute, Inc., Raleigh, North Carolina
1991. Who's Who in Australasia and the Far East, Second Edition, Cambridge, U.K
1992. The International Directory of Distinguished Leadership, Third Edition, American
Biographical Institute, Inc., Raleigh, North Carolina
PUBLICATIONS:
REFEREED JOURNALS
1. Pellis, S. M. (1981). Exploration and play in the behavioural development of the
Australian magpie Gymnorhina tibicen. Bird Behaviour, 3: 37-49.
2. Pellis, S. M. (1981). A description of social play by the Australian magpie
Gymnorhina tibicen based on Eshkol-Wachman notation. Bird Behaviour, 3: 61-79.
3. Pellis, S. M. (1982). An analysis of courtship and mating in the Cape Barren goose
Cereopsis novaehollandiae Latham based on Eshkol-Wachman Movement Notation. Bird
Behaviour, 4: 30-41.
4. Pellis, S. M., & Pellis, V. C. (1982). Do post-hatching factors limit clutch size in the
Cape Barren goose (Cereopsis novaehollandiae Latham)? Australian Wildlife Research, 9:
145-149.
5. Pellis, S. M. (1983). Development of head and foot coordination in the Australian
Magpie Gymnorhina tibicen, and the function of play. Bird Behaviour, 4: 57-62.
6. Pellis, S. M. (1983). The frequency and pattern of play behaviour. Mammalia, 47: 272274.
7. Pellis, S. M., & Pellis, V. C. (1983). Locomotor-rotational movements in the ontogeny
and play of the laboratory rat Rattus norvegicus. Developmental Psychobiology, 16: 269286.
8. Pellis, S. M. (1984). Two aspects of play-fighting in a captive group of Oriental smallclawed otters Amblonyx cinerea. Zeitschrift für Tierpsychologie, 65: 77-83.
9. Pellis, S. M., & Nelson, J. E. (1984). Some aspects of predatory behaviour of the quoll
Dasyurus viverrinus. Australian Mammalogy, 7: 5-15.
10. DeVietti, T. L., Pellis, S. M., Pellis, V. C., & Teitelbaum, P. (1985). Previous
experience disrupts atropine-induced stereotyped "trapping". Behavioral Neuroscience,
99: 1128-1141.
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11. Pellis, S. M. (1985). What is "fixed" in a Fixed Action Pattern? A problem of
methodology. Bird Behaviour, 6: 10-15.
12. Pellis, S. M., Chen, Y.-C., & Teitelbaum, P. (1985). Fractionation of the cataleptic
bracing response in rats. Physiology & Behavior, 34: 816-823.
13. Pellis, S. M., Chen, Y.-C., Chesire, R. M., & Teitelbaum, P. (1985). Head
displacement and bracing in haloperidol-treated rats compared to rats with lateral
hypothalamic damage. Physiology & Behavior, 35: 799-804.
14. Chen, Y.-C., Pellis, S. M., Sirkin, D. W., Potegal, M., & Teitelbaum, P. (1986).
Bandage-backfall: Labyrinthine and non-labyrinthine components. Physiology &
Behavior, 37: 805-814.
15. Pellis, S. M., De La Cruz, F., Pellis, V. C., & Teitelbaum, P. (1986). Morphine
subtracts subcomponents of haloperidol-isolated postural support reflexes revealing
gradients of their integration. Behavioral Neuroscience, 100: 631-646.
16. Pellis, S. M., Pellis, V. C., O'Brien, D. P., De La Cruz, F., & Teitelbaum, P. (1987).
Pharmacological subtraction of the sensory controls over grasping in rats. Physiology &
Behavior, 39: 127-133.
17. De La Cruz, F., Pellis, S. M., & Pellis, V. C. (1987). Sex differences in the effects of
haloperidol, morphine and their combination on colonal temperature in rats. Experimental
Neurology, 96: 376-380.
18. Pellis, S. M., & Officer, R. C. E. (1987). An analysis of some predatory behaviour
patterns in four species of carnivorous marsupials (Dasyuridae), with comparative notes
on the eutherian cat Felis catus. Ethology, 75: 177-196.
19. Pellis, S. M., & Pellis, V. C. (1987). Play-fighting differs from serious fighting in
both target of attack and tactics of fighting in the laboratory rat Rattus norvegicus.
Aggressive Behavior, 13: 227-242.
20. Pellis, S. M., Pellis, V. C., & Teitelbaum, P. (1987). "Axial apraxia" in
labyrinthectomized lateral hypothalamic-damaged rats. Neuroscience Letters, 82: 217220.
21. Pellis, S. M., Pellis, V. C., Chesire, R. M., Rowland, N. E., & Teitelbaum, P. (1987).
Abnormal gait sequence in the locomotion released by atropine in catecholamine deficient
akinetic rats. Proceedings of the National Academy of Science (U.S.A.), 84: 8750-8753.
22. Pellis, S. M. (1987). Problems of definition and of identifying appropriate levels of
analysis. Politics & Life Sciences, 6: 49-51. Commentary on: J. C. Davies (1987).
Aggression: Some definition and some physiology. Politics & Life Sciences, 6: 27-57.
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23. Pellis, S. M., & Pellis, V. C. (1988). Play-fighting in the Syrian golden hamster
Mesocricetus auratus Waterhouse, and its relationship to serious fighting during postweaning development. Developmental Psychobiology, 21: 323-337.
24. Pellis, S. M., O'Brien, D. P., Pellis, V. C., Teitelbaum, P., Wolgin, D. L., & Kennedy,
S. (1988). Escalation of feline predation along a gradient from avoidance through "play"
to killing. Behavioral Neuroscience, 102: 760-777.
25. Pellis, S. M. (1988). Agonistic versus amicable targets of attack and defense:
Consequences for the origin, function and descriptive classification of play-fighting.
Aggressive Behavior, 14: 85-104.
26. Pellis, S. M., & Pellis, V. C. (1988). Identification of the possible origin of the body
target which differentiates play-fighting from serious fighting in Syrian golden hamsters
Mesocricetus auratus. Aggressive Behavior, 14: 437-449.
27. Pellis, S. M., & Pellis, V. C. (1989). Targets of attack and defense in the play fighting
by the Djungarian hamster Phodopus campbelli: Links to fighting and sex. Aggressive
Behavior, 15: 217-234.
28. Pellis, S. M., Pellis, V. C., Morrissey, T. K., & Teitelbaum, P. (1989). Visual
modulation of vestibular-triggered air-righting in the rat. Behavioural Brain Research, 3:
23-26.
29. Pellis, S. M., Pellis, V. C., & Dewsbury, D. A. (1989). Different levels of complexity
in the playfighting by muroid rodents appear to result from different levels of intensity of
attack and defense. Aggressive Behavior, 15: 297-310.
30. Morrissey, T. K., Pellis, S. M., Pellis, V. C., & Teitelbaum. P. (1989). Seemingly
paradoxical jumping is triggered by postural instability in cataleptic haloperidol-treated
rats. Behavioural Brain Research, 35: 195-207.
31. Pellis, S. M., Pellis, V. C., Chen, Y.-C., Barzci, S., & Teitelbaum, P. (1989).
Recovery from axial apraxia in the lateral hypothalamic labyrinthectomized rat reveals
three elements of contact-righting: Cephalic dominance, axial rotation, and distal limb
action. Behavioural Brain Research, 35: 241-251.
32. Pellis, S. M., & Pellis, V. C. (1990). Differential rates of attack, defense and
counterattack during the developmental decrease in play fighting by male and female rats.
Developmental Psychobiology, 23: 215-231.
33. Pellis, S. M., Teitelbaum, P., & Meyer, M. E. (1990). Labyrinthine involvement in the
dorsal immobility response of adult rats. Behavioural Brain Research, 39: 197-204.
34. Porter, J. D., Pellis, S. M., & Meyer, M. E. (1990). An open-field analysis of
bilaterally labyrinthectomized rats. Physiology & Behavior, 48: 27-30.
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35. Whishaw, I. Q., & Pellis, S. M. (1990). The structure of skilled forelimb reaching in
the rat: A proximally driven stereotyped movement with a single rotatory component.
Behavioural Brain Research, 41: 49-59.
36. Whishaw, I. Q., Pellis, S. M., Gorny, B. P., & Pellis, V. C. (1991). The impairments
in reaching and the movements of compensation in rats with motor cortex lesions: A
videorecording and movement notation analysis. Behavioural Brain Research, 42: 77-91.
37. Pellis, S. M., & Pellis, V. C. (1991). Role reversal changes during the ontogeny of
play fighting in male rats: Attack versus defense. Aggressive Behavior, 17: 179-189.
38. Pellis, V. C., Pellis, S. M., & Teitelbaum, P. (1991). A descriptive analysis of the
post-natal development of contact-righting in rats (Rattus norvegicus). Developmental
Psychobiology, 24: 237-263.
39. Pellis, S. M., & Pellis, V. C. (1991). Attack and defense during play fighting appear
to be motivationally independent behaviors in muroid rodents. The Psychological Record,
41: 175-184.
40. Ben-David, M., Pellis, S. M., & Pellis, V. C. (1991). Feeding habits and predatory
behaviour of marbled polecats (Vormella peregusna syriaca): I. Killing methods with
relation to prey size and prey behaviour. Behaviour, 118: 127-143.
41. Pierce, J. D., Jr., Pellis, V. C., Dewsbury, D. A., & Pellis, S. M.* (1991). Targets and
tactics of agonistic and precopulatory behavior in montane and prairie voles: Their
relationship to juvenile play fighting. Aggressive Behavior, 17: 337-349.
42. Pellis, S. M., Pellis, V. C., Manning, C. J., & Dewsbury, D. A. (1991). The paucity of
social play in juvenile Mus musculus: What is missing from the behavioural repertoire?
Animal Behaviour, 42: 686-687.
43. Pellis, S. M. (1991). How motivationally distinct is play? A preliminary case study.
Animal Behaviour, 42: 851-853.
44. Pellis, S. M., Pellis, V. C., & Teitelbaum, P. (1991). Air-righting without the cervical
righting reflex in adult rats. Behavioural Brain Research, 45: 185-188.
45. Pellis, S. M., Pellis, V. C., & Teitelbaum, P. (1991). Labyrinthine and other
supraspinal controls over head-and-body ventroflexion. Behavioural Brain Research, 46:
99-102.
46. Pellis, S. M., Whishaw, I. Q., & Pellis, V. C. (1991). Visual modulation of
vestibularly-triggered air-righting in rats involves the superior colliculus. Behavioural
Brain Research, 46: 151-156.
13
47. Teitelbaum, P., & Pellis, S. M. (1992). Towards a synthetic physiological
psychology. Psychological Science, 3: 4-20.
48. Whishaw, I. Q., Pellis, S. M., & Gorny, B. P. (1992). Skilled reaching in rats and
humans: Evidence for parallel development or homology. Behavioural Brain Research,
47: 59-70.
49. Pellis, S. M., Pellis, V. C., & Nelson, J. E. (1992). The development of righting
reflexes in the pouch young of the marsupial Dasyurus hallucatus. Developmental
Psychobiology, 25: 105-125.
50. Whishaw, I. Q., Dringenberg, H. C., & Pellis, S. M. (1992). Forelimb use in free
feeding by rats: Motor cortex aids limb and digit positioning. Behavioural Brain
Research, 48: 113-125.
51. Pellis, S. M., & Pellis, V. C. (1992). An analysis of the targets and tactics of
conspecific attack and predatory attack in northern grasshopper mice Onychomys
leucogaster. Aggressive Behavior, 18: 301-316.
52. Pellis, S. M., Pellis, V. C., & Whishaw, I. Q. (1992). The role of the cortex in play
fighting by rats: Developmental and evolutionary implications. Brain, Behavior &
Evolution, 39: 270-284.
53. Pellis, S. M., Pellis, V. C., Manning, C. J., & Dewsbury, D. A. (1992). Supine
defense in the intraspecific fighting of male house mice Mus domesticus. Aggressive
Behavior, 18: 373-379.
54. Pellis, S. M., Vanderlely, R., & Nelson, J. E. (1992). The roles of vision and vibrissae
in the predatory behaviour of northern quolls Dasyurus hallucatus (Marsupialia:
Dasyuridae). Australian Mammalogy, 15: 55-60.
55. Whishaw, I. Q., Pellis, S. M., & Gorny, B. P. (1992). Medial frontal cortex lesions
impair the aiming component of rat reaching. Behavioural Brain Research, 50: 93-104.
56. Pellis, S. M., & McKenna, M. M. (1992). Intrinsic and extrinsic influences on play
fighting in rats: Effects of dominance, partner's playfulness, temperament and neonatal
exposure to testosterone propionate. Behavioural Brain Research, 50: 135-145.
57. Pellis, S. M., Pellis, V. C., Pierce, J. D., Jr., & Dewsbury, D A. (1992). Disentangling
the contribution of the attacker from that of the defender in the differences in the
intraspecific fighting of two species of voles. Aggressive Behavior, 18: 425-435.
58. Pellis, S. M., Pellis, V. C., & Kolb, B. (1992). Neonatal testosterone augmentation
increases juvenile play fighting but does not influence adult dominance relationships in
male rats. Aggressive Behavior, 18: 437-447.
14
59. Pellis, S. M., & Pellis, V. C. (1992). Juvenilized play fighting in subordinate male
rats. Aggressive Behavior, 18: 449-457.
60. Whishaw, I. Q., Pellis, S. M., & Pellis, V. C. (1992). A behavioral study of the
contributions of cells and fibers of passage in the red nucleus of the rat to postural
righting, skilled movements, and learning. Behavioural Brain Research, 52: 29-44.
61. Pellis, S. M. (1992). The yin and yang of behavioral analysis. Behavioral Brain
Sciences, 15: 286.
62. Pellis, S. M., & Pellis, V. C. (1993). The influence of dominance on the development
of play fighting in pairs of male Syrian golden hamsters (Mesocricetus auratus).
Aggressive Behavior, 19: 293-302.
63. Pellis, S. M., Pellis, V. C., & McKenna, M. M. (1993). Some subordinates are more
equal than others: Play fighting amongst adult subordinate male rats. Aggressive Behavior,
19: 385-393.
64. Whishaw, I. Q., Pellis, S. M., Gorny, B., Kolb, B., & Tetzlaff, W. (1993). Proximal
and distal impairments in rat forelimb use in reaching following pyramidal tract lesions.
Behavioural Brain Research, 56: 59-76.
65. Pellis, S. M., Castañeda, E., McKenna, M. M., Tran-Nguyen, L. T. L., & Whishaw, I.
Q. (1993). The role of the striatum in organizing sequences of play fighting in neonatally
dopamine-depleted rats. Neuroscience Letters, 158: 13-15.
66. Cordover, A. J., Pellis, S. M., & Teitelbaum, P. (1993). Haloperidol exaggerates
proprioceptive-tactile support reflexes and diminishes vestibular control over them.
Behavioural Brain Research, 56: 197-201.
67. Pellis, S. M. (1993). Sex and the evolution of play fighting: A review and a model
based on the behavior of muroid rodents. The Journal of Play Theory & Research, 1: 5677.
68. Pellis, S. M., Pellis, V. C., & McKenna, M. M. (1994). A feminine dimension in the
play fighting of rats (Rattus norvegicus) and its defeminization neonatally by androgens.
Journal of Comparative Psychology, 108: 68-73.
69. Whishaw, I. Q., Gorny, B., Tran-Nguyen, L. T. L., Castañeda, E., Miklyaeva, E. I., &
Pellis, S. M. (1994). Doing two things at once: Impairments in movement and posture
underlie the adult skilled reaching deficit of neonatally dopamine-depleted rats.
Behavioural Brain Research, 61: 65-77.
70. Field, E. F., & Pellis, S. M.* (1994). Differential effects of amphetamine on the attack
and defense components of play fighting in rats. Physiology & Behavior, 56: 325-330.
15
71. Pellis, S. M., & Pellis, V. C. (1994). The development of righting when falling from a
bipedal standing posture: Evidence for the dissociation of dynamic and static righting
reflexes in rats. Physiology & Behavior, 56: 659-663.
72. Pellis, S. M., & McKenna, M. M. (1995). What do rats find rewarding in play
fighting? An analysis using drug-induced non-playful partners. Behavioural Brain
Research, 68: 65
73. Pellis, S. M., & Pellis, V. C. (1996). On knowing it’s only play: The role of play
signals in play fighting. Aggression & Violent Behavior, 1: 249-268.
74. Pellis, S. M., Pellis, V. C., & Whishaw, I. Q. (1996). Visual modulation of air
righting by rats involves calculation of time-to-impact but does not require the detection of
the looming stimulus of the approaching ground. Behavioural Brain Research, 74: 207211.
75. Pellis, S. M., MacDonald, N., & Michener, G. R. (1996). The lateral display as a
combat tactic in Richardson’s ground squirrel Spermophilus richardsonii. Aggressive
Behavior, 22: 119-134
76. Pellis, S. M., McKenna, M. M., Field, E. F., Pellis, V. C., Prusky, G. T., & Whishaw,
I. Q. (1996). The uses of vision by rats in play fighting and other close quarter social
interactions. Physiology & Behavior, 59: 905-913.
77. Smith, L. K., Field, E. F., Forgie, M. L., & Pellis, S. M.* (1996). Dominance and
age-related changes in the play fighting of intact and post-weaning castrated males (Rattus
norvegicus ). Aggressive Behavior, 22: 215-226.
78. Ivanco, T. L., Pellis, S. M., & Whishaw, I. Q. (1996). Skilled movements in prey
catching and in reaching by rats (Rattus norvegicus) and opossums (Monodelphis
domestica): Relations to anatomical differences in motor systems. Behavioural Brain
Research, 79: 163-182.
79. Field, E. F., Whishaw, I. Q., & Pellis, S. M.* (1996). An analysis of sex differences in
the movement patterns used during the food wrenching and dodging paradigm. Journal of
Comparative Psychology, 110: 298-306.
80. Martens, D. J., Whishaw, I. Q., Miklyaeva, E. I., & Pellis, S. M.* (1996). Spatiotemporal impairments in limb and body movements during righting in an
hemiparkinsonian rat analogue: Relevance to axial apraxia in humans. Brain Research,
733: 253-262.
81. Pellis, S. M., & Pellis, V. C. (1997). Targets, tactics and the open mouth face during
play fighting in three species of primates. Aggressive Behavior, 23: 41-57.
16
82. Whishaw, I. Q., Coles, B. L., Pellis, S. M., & Miklyaeva, E. I. (1997). Impairments
and compensation in mouth and limb use in food handling after unilateral dopamine (DA)
depletions in a rat analogue of human Parkinson’s disease. Behavioural Brain Research,
84:167-177.
83. Field, E. F., Whishaw, I. Q., & Pellis, S. M.* (1997). The organization of sex-typical
patterns of defense during food protection in the rat: The role of the opponent’s sex.
Aggressive Behavior, 23: 197-214.
84. Field, E. F., Whishaw, I. Q., & Pellis, S. M.* (1997). A kinematic analysis of sextypical movement patterns used during evasive dodging to protect a food item: The role of
gonadal androgens. Behavioral Neuroscience, 111: 808-815.
85. Pellis, S. M., & Pellis, V. C. (1997). The pre-juvenile onset of play fighting in rats
(Rattus norvegicus). Developmental Psychobiology, 31:193-205.
86. Pellis, S. M., Field, E. F., Smith, L. K., & Pellis, V. C. (1997). Multiple differences in
the play fighting of male and female rats. Implications for the causes and functions of
play. Neuroscience & Biobehavioral Reviews, 21:105-120.
87. Pellis, S. M. (1997). Targets and tactics: The analysis of moment-to-moment decision
making in animal combat. Aggressive Behavior, 23: 107-129.
88. Pellis, S. M., & Pellis, V. C. (1998). The play fighting of rats in comparative
perspective: A schema for neurobehavioral analyses. Neuroscience & Biobehavioral
Reviews, 23: 87-101.
89. Whishaw, I. Q., Woodward, N. C., Miklyaeva, E., & Pellis, S. M. (1998). Analysis of
limb use by control rats and unilateral DA-depleted rats in the Montoya staircase test:
Movements, impairments and compensatory strategies. Behavioural Brain Research,
89:167-177.
90. Smith, L. K., Forgie, M., & Pellis, S. M.* (1998). The post-pubertal change in the
playful defense of male rats depends upon neonatal exposure to gonadal hormones.
Physiology & Behavior, 63: 151-155.
91. Smith, L. K., Forgie, M. L., & Pellis, S. M.* (1998). Mechanisms underlying the
absence of the pubertal shift in the playful defense of female rats. Developmental
Psychobiology, 33: 147-156.
92. Whishaw, I. Q., Sarna, J., & Pellis, S. M. (1998). Evidence for rodent-common and
species-typical limb and digit use in eating derived from a comparative analysis of ten
rodent species. Behavioural Brain Research, 96: 79-91.
93. Iwaniuk, A. N., Nelson, J. E., Ivanco, T. L., Pellis, S. M., & Whishaw, I Q. (1998).
Reaching, grasping and manipulation of food objects by two tree kangaroo species,
17
Dendrolagus lumholtzi and Dendrolagus matschiei. Australian Journal of Zoology, 46:
235-248.
94. Smith, L. K., Fantella, S.-L., & Pellis, S. M.* (1999). Playful defensive responses in
adult male rats depend upon the status of the unfamiliar opponent. Aggressive Behavior,
25: 141-152.
95. Pellis, S. M., & Pasztor, T. J. (1999). The developmental onset of a rudimentary form
of play fighting in mice, Mus musculus. Developmental Psychobiology, 34: 175-182.
96. Iwaniuk, A. N., Pellis, S. M., & Whishaw, I. Q. (1999). The role of the corticospinal
tract in the evolution of digital dexterity: A phylogenetic analysis of the Heffner and
Masterton data set. Behavioural Brain Research, 101: 173-187.
97. Pellis, S. M., & Iwaniuk, A. N. (1999). The roles of phylogeny and sociality in the
evolution of social play in muroid rodents. Animal Behaviour, 58: 361-373.
98. Pellis, S. M., Field, E. F., & Whishaw, I. Q. (1999). The development of a sexdifferentiated defensive motor-pattern in rats: A possible role for juvenile experience.
Developmental Psychobiology, 35: 156-164.
99. Pellis, S. M., & Iwaniuk, A. N. (1999). The problem of adult play: A comparative
analysis of play and courtship in primates. Ethology, 105: 783-806.
100. Iwaniuk, A. N., Pellis, S. M., & Whishaw, I. Q. (1999). Brain size is not correlated
with forelimb dexterity in fissiped carnivores: A comparative test of the principle of
proper mass. Brain, Behavior and Evolution, 54: 167-180.
101. Iwaniuk, A. N., Pellis, S. M., & Whishaw, I. Q. (1999). The relationship between
forelimb morphology and behaviour in North American carnivores (Carnivora). Canadian
Journal of Zoology, 77: 1064-1074.
102. Pellis, S. M., & Iwaniuk, A. N. (2000). Comparative analyses of the role of postnatal
development on the expression of play fighting. Developmental Psychobiology, 36: 136147.
103. Field, E. F., Whishaw, I. Q., & Pellis, S. M.* (2000). Sex differences in catalepsy:
Evidence for hormone-dependent postural mechanisms in haloperidol-treated rats.
Behavioural Brain Research, 109: 207-212.
104. Pellis, S. M., Pasztor, T. J., Pellis, V. C., & Dewsbury, D. A. (2000). The
organization of play fighting in the grasshopper mouse (Onychomys leucogaster): Mixing
predatory and sociosexual targets and tactics. Aggressive Behavior, 26: 319-334.
18
105. Iwaniuk, A. N., Pellis, S. M., & Whishaw, I. Q. (2000). The relative importance of
body size, phylogeny, locomotion and diet on the evolution of forepaw dexterity in
terrestrial carnivores. Canadian Journal of Zoology, 78: 1110-1125.
106. Thierry, B., Iwaniuk, A. N., & Pellis, S. M.* (2000). The influence of phylogeny on
the social behaviour of macaques (Primates: Cercopithecidae, genus Macaca). Ethology,
106: 713-728.
107. Pellis, S. M., & Iwaniuk, A. N. (2000). Adult-adult play in primates: Comparative
analyses of its origin, distribution and evolution. Ethology, 106: 1083-1104.
108. Iwaniuk, A. N., Nelson, J. E., & Pellis, S. M.* (2001). Do big-brained animals play
more? Comparative analyses of play and relative brain size in mammals. Journal of
Comparative Psychology, 115: 29-41.
109. Iwaniuk, A. N., Pellis, S. M., & Whishaw, I. Q. (2001). Are long digits correlated
with high forepaw dexterity? A comparative test in terrestrial carnivorans (Carnivora).
Canadian Journal of Zoology, 79: 900-906.
110. Pasztor, T. J., Smith, L. K., MacDonald, N. L., Michener, G. R., & Pellis, S. M.*
(2001). Sexual and aggressive play fighting of sibling Richardson’s ground squirrels.
Aggressive Behavior, 27: 323-337.
111. Whishaw, I. Q., Metz, G. A., Kolb, B., & Pellis, S. M. (2001). Accelerated nervous
system development contributes to behavioral efficiency in the laboratory mouse: A
behavioral review and theoretical proposal. Developmental Psychobiology, 39: 151-170.
112. Pellis, S. M. (2002). Sex-differences in play fighting revisited: Traditional and nontraditional mechanisms for sexual differentiation in rats. Archives of Sexual Behavior, 31:
11-20.
113. Lingle, S., & Pellis, S. M. (2002). Fight or flight? Antipredator behavior and the
escalation of coyote encounters with deer. Oecologica, 131: 154-164.
114. Whishaw, I. Q., Suchowersky, O., Davis, L., Sarna, J., Metz, G. A., & Pellis, S. M.
(2002). A qualitative analysis of reaching-to-grasp movements in human Parkinson’s
disease (PD) reveals impairments in coordination and rotational movements of pronation
and supination: a comparison to deficits in animal models of PD. Behavioural Brain
Research, 133: 165-176.
115. Pellis, S. M., & Iwaniuk, A. N. (2002). Brain system size and adult-adult play in
primates: A comparative analysis of the roles of the non-visual neocortex and the
amygdala. Behavioural Brain Research, 134: 31-39.
19
116. Wallace, D. G., Hines, D.J., Pellis, S. M., & Whishaw, I. Q. (2002). Vestibular
information is required for dead reckoning in the rat. Journal of Neuroscience, 22: 1000910017.
117. Foroud, A., & Pellis, S. M.* (2002). The development of ‘anchoring’ in the play
fighting of rats: Evidence for an adaptive age-reversal in the juvenile phase. International
Journal of Comparative Psychology, 15: 11-20.
118. Pellis, S. M. (2002). When is a trait an adaptation? Behavioral Brain Sciences, 25:
524.
119. Foroud, A., & Pellis, S. M.* (2003). The development of ‘roughness’ in the play
fighting of rats: A Laban Movement Analysis perspective. Developmental Psychobiology,
42: 35-43.
120. Foroud, A., Whishaw, I. Q., & Pellis, S. M.* (2004). Experience and cortical control
over the pubertal transition to rougher play fighting in rats. Behavioural Brain Research,
149: 69-76.
121. Pellis, S. M., & Iwaniuk, A. N. (2004). Evolving a playful brain: A levels of control
approach. International Journal of Comparative Psychology, 17: 90-116.
122. Kolb, B., Pellis, S. M., & Robinson, T. E. (2004). Plasticity and functions of the
orbital frontal cortex. Brain & Cognition, 55: 104-115.
123. Reinhart, C. J., McIntyre, D. C., & Pellis, S. M.* (2004). The development of play
fighting in kindling-prone (FAST) and kindling–resistant (SLOW) rats: How does the
retention of phenotypic juvenility affect the complexity of play? Developmental
Psychobiology, 45: 83-92.
124. Komorowska, J., & Pellis, S. M.* (2004). Regulatory mechanisms underlying
novelty-induced grooming in the laboratory rat. Behavioural Processes, 67: 287-293.
125. Field, E. F., Whishaw, I. Q., & Pellis, S. M.* (2004). Evidence for the necessity of
pubertal ovarian hormones for the organization of female-typical patterns of dodging to
protect a food item. Behavioral Neuroscience, 118: 1293-1304.
126. Field, E. F., Watson, N. V., Whishaw, I. Q., & Pellis, S. M.* (2005). A masculinized
skeletomusculature is not required for male-typical patterns of food-protective
movements. Hormones & Behavior, 47: 49-55.
127. Cooper, S., Brown, L., Doan, J., Pellis, S. M., & Whishaw, I. Q. (2005). Reducing
stability of support structure for a target does not alter reach kinematics among younger
adults. Perceptual and Motor Skills, 100: 831-838.
20
128. Melvin, K. G., Doan, J., Pellis, S. M., Brown, L., Whishaw, I. Q., & Suchowersky,
O. (2005). Pallidal deep brain stimulation and L-dopa do not improve qualitative aspects
of skilled reaching in Parkinson's disease. Behavioural Brain Research, 160: 188-194.
129. Field, E. F., Martens, D. J., Watson, N. V., & Pellis, S. M.* (2005). Sex differences
in righting from supine to prone: A masculinized skeletomusculature is not required.
Journal of Comparative Psychology, 119: 238-245.
130. Lingle, S., Pellis, S. M., & Wilson, W. F. (2005). Interspecific variation in
antipredator behaviour leads to differential vulnerability of mule deer and white-tailed
deer fawns early in life. Journal of Animal Ecology, 74: 1140-1149.
131. Doan, J., Whishaw, I. Q., Pellis, S. M., Suchowersky, O., & Brown, L. (2006). Motor
deficits in parkinsonian reaching: Dopa-sensitivity influenced by real world taskconstraint. Journal of Motor Behavior, 38: 45-59.
132. Field, E. F., Watson, N. V., Whishaw, I. Q., & Pellis, S. M.* (2006). Play-fighting in
androgen-insensitive tfm rats: Evidence that androgen receptors are critical for the
development of adult playful defense but not playful attack. Developmental
Psychobiology, 48: 111-120.
133. Pellis, S. M., Hastings, E., Shimizu, T., Kamitakahara, H., Komorowska, J., Forgie
M. L., & Kolb, B. (2006). The effects of orbital frontal cortex damage on the modulation
of defensive responses by rats in playful and non-playful social contexts. Behavioral
Neuroscience, 120: 72-84.
134. Reinhart, C. J., Metz, G., McIntyre, D. C., & Pellis, S. M.* (2006). Play fighting
between kindling-prone (FAST) and kindling-resistant (SLOW) rats. Journal of
Comparative Psychology, 120: 19-30.
135. Brown, L. A., Cooper, S. A., Doan, J. B., Dickin, D. C., Whishaw, I. Q., Pellis, S. M.
& Suchowersky, O. (2006). Parkinsonian deficits in sensory integration for postural
control: temporal response to changes in visual input. Parkinsonism & Related Disorders,
12: 376-381.
136. Field, E. F., Metz, G. A., Pellis, S. M., & Whishaw, I. Q. (2006). Sexually dimorphic
postural adjustments during vertical behaviour are altered in a unilateral 6-OHDArat
model of Parkinson's disease. Behavioural Brain Research, 174: 39-48.
137. Kamitakahara, H., Monfils, M.-H., Forgie, M. L., Kolb, B., & Pellis, S. M.* (2007).
The modulation of play fighting in rats: Role of the motor cortex. Behavioral
Neuroscience, 121: 164-176.
138. Pellis, S. M., & Pellis, V. C. (2007). Rough and tumble play and the development of
the social brain. Current Directions in Psychological Science, 16: 95-98.
21
139. Lingle, S., Rendall, D., & Pellis, S. M. (2007). Altruism and recognition in the
antipredator defence of deer: I. Species and individual variation in fawn distress calls.
Animal Behaviour, 73: 897-905.
140. Lingle, S., Rendall, D., Wilson, W. F., Deyoung, R. W., & Pellis, S. M. (2007).
Altruism and recognition in the antipredator defence of deer: II. Why mule deer help nonoffspring fawns. Animal Behaviour, 73: 907-916.
141. Field, E. F., & Pellis, S. M.* (2008). The brain as the engine of sex differences in the
organization of movement in rats. Archives of Sexual Behavior, 37: 30-42.
142. Pellis, S. M., & Pellis, V. C. (2009). Play and the socially competent brain.
PlayRights, Issue 2: 4-7.
143. Pellis, S. M., Gray, D., & Cade, W. H. (2009). The judder of the cricket: The
variance underlying the invariance in behavior. International Journal of Comparative
Psychology, 22: 188-205.
144. Bell, H. C., McCaffrey, D., Forgie, M. L., Kolb, B., & Pellis, S. M.* (2009). The role
of the medial prefrontal cortex in the play fighting in rats. Behavioral Neuroscience, 123:
1158-1168.
145. Bell, H. C., Pellis, S. M.,* & Kolb, B. (2010). Juvenile peer play experience and
development of the orbitofrontal and medial prefrontal cortices. Behavioural Brain
Research, 207: 7-13.
146. Reinhart, C. J., Pellis, V. C., Thierry, B., Gauthier, C.-A., VanderLaan, D. P., Vasey,
P. L., & Pellis, S. M.* (2010). Targets and tactics of play fighting: Competitive versus
cooperative styles of play in Japanese and Tonkean macaques. International Journal of
Comparative Psychology, 23: 166-200.
147. Pellis, S. M., Pellis, V. C., & Bell, H. C. (2010). The function of play in the
development of the social brain. American Journal of Play, 2: 278-296.
148. Pellis, S. M. (2010). Conservative motor systems, behavioral modulation and neural
plasticity. Behavioural Brain Research, 214: 25-29.
149. Doan, J. B., Whishaw, I. Q., Pellis, S. M., Suchowersky, O., de Bruin, N., & Brown,
L. A. (2010). Challenging context affects standing reach kinematics among Parkinson's
disease patients. Behavioural Brain Research, 214: 135-141.
150. Pellis, S. M., & Pellis, V. C. (2011). To whom the play signal is directed: A study of
headshaking in black-handed spider monkeys (Ateles geoffroyi). Journal of Comparative
Psychology, 125: 1-10.
22
151. Muhammad, A., Hossain, S., Pellis, S. M. & Kolb, B. (2011). Tactile stimulation
during development attenuates amphetamine sensitization and structurally reorganizes
prefrontal cortex and striatum in a sex-dependent manner. Behavioral Neuroscience, 125:
161-174.
152. Pellis, S. M. (2011). Head and foot coordination in head scratching and food
manipulation by purple swamp hens (Porphyrio porphyrio): Rules for minimizing the
computational costs of combining movements from multiple parts of the body.
International Journal of Comparative Psychology, 24: 255-271.
153. Pellis, S. M., Pellis, V. C., Reinhart, R. J., & Thierry, B. (2011). The use of the
bared-teeth display during play fighting in Tonkean macaques (Macaca tonkeana):
Sometimes it is all about oneself. Journal of Comparative Psychology, in press.
154. Bell, H. C. & Pellis, S. M.* (2011). A cybernetic perspective on food protection in
rats: Simple rules can generate complex and adaptable behaviour. Animal Behaviour, in
press.
155. Pellis, S. M. & Bell, H. C. (2011). Closing the circle between perceptions and
behavior: A cybernetic view of behavior and its consequences for studying motivation and
development. Developmental Cognitive Neuroscience, in press.
156. Pellis, S. M. & Pellis, V. C. (2011). Anatomy is important, but need not be destiny:
Novel uses of the thumb in aye-ayes compared to other lemurs. Behavioural Brain
Research, in press.
BOOK CHAPTERS:
1. Pellis, S. M. (1989). Fighting: The problem of selecting appropriate behavior patterns.
In: R. J. Blanchard, P. F. Brain, D. C. Blanchard & S. Parmigiani (Eds.), Ethoexperimental
Approaches to the Study of Behavior (pp. 361-374). Dordrecht, The Netherlands: Kluwer
Academic Publishers.
2. Teitelbaum, P., Pellis, S. M., & DeVietti, T. L. (1990). Disintegration into stereotypy
induced by drugs or brain damage: A micro-descriptive behavioral analysis. In: S. J.
Cooper & C. T. Dourish (Eds.), Neurobiology of Behavioral Stereotypy (pp. 169-199).
Oxford, U.K.: Oxford University Press.
3. Teitelbaum, P., Pellis, V. C., & Pellis, S. M. (1991). Can allied reflexes promote the
recovery of a robot's behavior? In: J.-A. Meyer & S. W. Wilson (Eds.), Simulation of
Adaptive Behavior: From Animals to Animats (pp.97-104). The MIT Press/Bradford
Books.
23
4. Pellis, S. M. (1996). Righting and the modular organization of motor programs. In: K.P. Ossenkopp, M. Kavaliers & P. R. Sanberg (Eds.), Measuring Movement and
Locomotion: From Invertebrates to Humans (pp. 115-133). Landes Company, Austin, TX.
5. Field, E. F., & Pellis, S. M.* (1998). Sex differences in the organization of behavior
patterns: Endpoint measures do not tell the whole story. In: L. Ellis & L. Ebertz (Eds.),
Males, Females, and Behavior: Towards Biological Understanding (pp. 147-162).
Praeger, Westport, CT.
6. Pellis, S. M., & Pellis, V. C. (1998). The structure-function interface in the analysis of
play fighting. In: M. Bekoff & J. A. Byers (Eds.), Play Behavior: Comparative,
Evolutionary, and Ecological Aspects (pp. 115-140). Cambridge University Press,
Cambridge, U. K.
7. Pellis, S. M. (2002). Keeping in touch: Play fighting and social knowledge. In: M.
Bekoff, C. Allen & G. M. Burghardt (Eds.), The Cognitive Animal: Empirical and
Theoretical Perspectives on Animal Cognition (pp. 421-427). MIT Press, Cambridge, MA.
8. Pellis, S. M. (2005). Cross-species comparisons. In: B. Hopkins (Ed.), Cambridge
Encyclopedia of Child Development (pp. 112-114). Cambridge University Press,
Cambridge, UK.
9. Pellis, S. M., Pellis, V. C., & Foroud, A. (2005). Play fighting: Aggression, affiliation
and the development of nuanced social skills. In: R. Tremblay, W. W. Hartup & J. Archer
(Eds.), Developmental Origins of Aggression (pp. 47-62). Guilford Press, New York.
10. Pellis, S. M., & Pellis, V. C. (2005). Posture. In: I. Q. Whishaw & B. Kolb (Eds.),
The Behaviour of the Laboratory Rat. A Handbook with Tests (pp. 121-128). Oxford
University Press; Oxford, UK.
11. Pellis, S. M., & Pellis, V. C. (2005). Play and fighting. In: I. Q. Whishaw & B. Kolb
(Eds.), The Behaviour of the Laboratory Rat. A Handbook with Tests (pp. 298-306).
Oxford University Press; Oxford, UK.
12. Pellis, S. M., & Pellis, V. C. (2006). Play and the development of social engagement:
A comparative perspective. In: P. J. Marshall & N. A. Fox, (Eds.), The Development of
Social Engagement: Neurobiological Perspectives (pp. 247-274). Oxford University
Press; Oxford, UK.
13. Pellis, S. M., & Pellis, V. C. (2009). Play fighting. In: H. T. Reis & S. K. Sprecher
(Eds.), Encyclopedia of Human Relationships. Vol. 3 (P-Z) (pp. 1255-1256). Sage
Publications, Inc.; Thousand Oaks, CA.
14. Pellis, S. M., Pellis, V. C. & Reinhart, C. J. (2010). The evolution of social play. In: C.
Worthman, P. Plotsky, D. Schechter & C. Cummings (Eds.), Formative Experiences: The
24
Interaction of Caregiving, Culture, and Developmental Psychobiology (pp. 404-431).
Cambridge University Press; Cambridge, UK.
15. Pellis, S. M. & Pellis, V. C. (2010). Social play, social grooming and the regulation of
social relationships. In: A. V. Kalueff, J. I. LaPorte & C. L. Bergner (Eds.), Neurobiology
of grooming behavior (pp. 66-87). Cambridge University Press; Cambridge, UK.
16. Pellis, S. M. (2010). Behavioral development and socialization. In: G. Koob, R. F.
Thompson & M. Le Moal (Eds.), Encyclopedia of Behavioral Neuroscience (pp. 133138). Elsevier, Ltd.; Oxford, UK.
17. Pellis, S. M., & Pellis, V. C. (2010). Play. In: D. Mills (Ed.), Encyclopedia of Applied
Animal Behaviour and Welfare (pp. 477-479). CABI Publishing; Nosworthy Way, UK.
18. Pellis, S. M. & Pellis, V. C. (2011). Rough-and-tumble play – training and using the
social brain. In: A. D. Pellegrini (Ed.), Oxford Handbook of the Development of Play (pp.
245-259). Oxford University; Oxford, UK.
19. VanderLaan, D. P., Pellis, S. M. & Vasey, P. L. (2011). Mounting and social play in
juvenile male Japanese macaques. In: J.-B. Leca, M. A. Huffman, and P. L. Vasey (Eds.),
The Monkeys of Stormy Mountain: 60 years of Primatological Research on the Japanese
Macaques of Arashiyama. Cambridge University Press; Cambridge, UK, in press.
20. Pellis, S. M. & Pellis, V. C. (2011). Play fighting in Japanese macaques: A
comparative perspective. In: J.-B. Leca, M. A. Huffman, and P. L. Vasey (Eds.), The
Monkeys of Stormy Mountain: 60 years of Primatological Research on the Japanese
Macaques of Arashiyama. Cambridge University Press; Cambridge, UK, in press.
21. Pellis, S. M. & Pellis, V. C. (2011). Play-fighting during early childhood and its role
in preventing later chronic aggression. In: R. E. Tremblay (Ed.), Encyclopedia on Early
Childhood Development – Aggression. Online resource supported by the Centre of
Excellence for Early Childhood Development (CEECD) and the Strategic Knowledge
Centre on Early Childhood Development (SKC-ECD) (http://www.childencyclopedia.com/), in press
*I am the senior author in these papers. The work was supported by grants awarded to me,
and conducted by students in my laboratory under my supervision.
BOOKS AND EDITED WORKS:
1. Marino, L., & Pellis, S. M. (Eds.) (2004). The Evolution of Vertebrate Brain and
Behavior. Special issue of International Journal of Comparative Psychology, Volume 17
(1).
25
2. Ellis, L., Hershberger, S., Field, E. F., Wersinger, S., Pellis, S. M., Geary, D., Palmer,
C., Hoyenga, K., Hetsroni, A., & Karadi, K. (2008). Sex Differences: Summarizing more
than a Century of Scientific Research. Psychology Press; New York, NY.
3. Pellis, S. M., & Pellis, V. C. (2009). The Playful Brain. Venturing to the Limits of
Neuroscience. Oneworld Press; Oxford, UK.
4. Pellis, S. M., & Pellis, V. C. (2010). The Playful Brain. Venturing to the Limits of
Neuroscience. Oneworld Press; Oxford, UK. Updated paperback edition.
5. Teskey, G. C., & Pellis, S. M. (Eds.) (2010). Behavioural and Neural Plasticity. Special
issue of Behavioural Brain Research, Volume 214 (1).
6. Bacchus, S., Pellis, S. M., Rowland, N., Stellar, J. & Szechtman, H. (Eds.) (2011). Quo
Vadis Behavioral Neuroscience: Festschrift for Philip Teitelbaum. Special issue of
Behavioural Brain Research, in press
TEACHING-RELATED PUBLICATIONS:
1. Pellis, S. M., O'Brien, D. P., Pellis, V. C., Teitelbaum, P., Wolgin, D. L., & Kennedy,
S. (1988). Escalation of feline predation along a gradient from avoidance through "play"
to killing. Reprinted in J. P. J. Pinel (Ed.), Current Research in Biopsychology (pp. 7895). Boston, MA: Allyn and Bacon, 1991.
2. Pellis, S. M. (1994). What is the point of play? In J. R. Vokey (Ed.), Psychological
Sketches (pp. 51-59). RealScience Publishing, Department of Psychology, University of
Lethbridge, Lethbridge, Alberta, Canada.
3. Pellis, S. M. (1994). Standing up straight, and staying there. In J. R. Vokey (Ed.),
Psychological Sketches (pp. 95-101). RealScience Publishing, Department of Psychology,
University of Lethbridge, Lethbridge, Alberta, Canada.
BOOK REVIEWS:
1. Pellis, S. M. (1990). When cause meets effect. Comtemporary Psychology, 35: 24-25.
Review of J. Archer (1988). The Behavioural Biology of Aggression. Cambridge, U.K.:
Cambridge University Press.
2. Pellis, S. M. (1992). Review of Fear and Defense, edited by P. F. Brain, S. Parmigiani,
R. Blanchard & D. Mainardi (1990). London: Harwood Academic Publishers. Aggressive
Behavior, 18: 159-161.
3. Pellis, S. M. (1993). Review of Aggression and Peacefulness in Humans and Other
Primates, edited by J. Silverberg & J. P. Gray (1991). Oxford: Oxford University Press.
Aggressive Behavior, 19: 241-243.
26
4. Pellis, S. M. (1994). Review of The Aggressive Female, edited by M. Haug, D. Benton,
P. F. Brain, B. Olivier & J. Mos (1992). CIP-Cegevens Koninklïjke Bibliotheek, Den
Haag, The Netherlands. Aggressive Behavior, 20: 466-469.
5. Pellis, S. M. (2010). Review of The Role of Play in Human Development, A. D.
Pellegrini (2010). Oxford and New York: Oxford University Press. The Quarterly Review
of Biology, 85: 363.
ABSTRACTS:
1. Pellis, S. M. (1980). Behavioural development and the function of play in the
Australian magpie Gymnorhina tibicen. Australasian Society for the Study of Animal
Behaviour, Launching Place, Victoria, Australia.
2. Pellis, S. M., De La Cruz, F., Pellis, V. C., & Teitelbaum, P. (1983). Fractionation of
postural support mechanisms with combined systemic administration of haloperidol and
morphine. Society for Neuroscience, Boston, MA, USA.
3. De La Cruz, F., Pellis, S. M., Pellis, V. C., Teitelbaum, P., & Satinoff, E. (1984).
Effecto de lesiones hipotalamicas sobre la temperatura de la rata. Congreso Nacional de
Ciencias Fisologicas, Mexico City, Mexico.
4. Pellis, S. M., De La Cruz, F., Pellis, V. C., Teitelbaum, P., & Satinoff, E. (1984).
Accion hipotermicas de la administracion combinade de haloperidol y morphinas en la
rata. Congreso Nacional de Ciencias Fisologicas, Mexico City, Mexico.
5. Pellis, S. M., Pellis, V. C., & Teitelbaum, P. (1986). Abnormal gait sequence in the
locomotion released by atropine in catecholamine-deficient-akinetic rats. Society for
Neuroscience, Washington, DC, USA.
6. Pellis, S. M. (1986). Play-fighting differs from serious fighting in both target of attack
and tactics of fighting, as exemplified by the laboratory rat (Rattus norvegicus ) and by the
Syrian golden hamster (Mesocricetus auratus). Invited paper, International Society for
Research on Aggression, Chicago, IL, USA.
7. Ben-David, M., Hellwing, S., Pellis, S. M., & Pellis, V. C. (1987). Feeding habits and
predatory behaviour in the marbled polecat Vormela peregusna syriaca (Mustelidae) in
Israel. German Society for Mammalian Biology, West Berlin, West Germany.
8. Pellis, S. M., Morrissey, T. K., Pellis, V. C., & Teitelbaum, P. (1987). Fractionation of
the subcomponents and transmitter controls of contact-righting in labyrinthectomized
lateral hypothalamic-damaged rats. Society for Neuroscience, New Orleans, LA, USA.
9. Pierce, J. D., Jr., Pellis, S. M.,* Pellis, V. C., & Dewsbury, D. A. (1988). Analysis of
the body target areas approached by the male and defended by the female during
27
copulatory behavior in two species of voles, Microtus ochrogaster and Microtus
montanus. American Society of Mammalogists, Clemson, SC, USA.
10. Pellis, S. M. (1988). Targets and "decision points" in the comparative analysis of the
tactics of fighting. Invited paper, International Society for Research on Aggression,
Swansea, Wales, U.K.
11. Pellis, V. C., & Pellis, S. M.* (1988). Playful versus agonistic counterattack in
muroid rodents: Useful markers for comparative and ontogenetic studies of play-fighting.
International Society for Research on Aggression, Swansea, Wales, U.K.
12. Pellis, S. M. (1988). Fighting: The problem of selecting appropriate behavior
patterns. Invited paper, NATO Advanced Study Institute "The Ethoexperimental
Analysis of Behavior", Pisa, Italy.
13. Pellis, S. M., Pellis, V. C., & Dewsbury, D. A. (1988). The evolution of play-fighting:
A comparative analysis of some muroid rodents. Joint meeting of the Australasian Society
for the Study of Animal Behaviour and the International Society for Comparative
Psychology, Sydney, Australia.
14. Pellis, V. C., Pellis, S. M., & Teitelbaum. P. (1988). Subcomponent reflexes involved
in the contact-righting of neonatal altricial mammals. Joint meeting of the Australasian
Society for the Study of Animal Behaviour and the International Society for Comparative
Psychology, Sydney, Australia.
15. Morrissey, T. K., Pellis, S. M., Pellis, V. C., & Teitelbaum, P. (1988). Dopamine
blockade releases de-encephalized jumping triggered by postural instability. Society for
Neuroscience, Toronto, ON, Canada.
16. O'Brien, D. P., Pellis, S. M., Pellis, V. C., & Teitelbaum, P. (1988). Diazepam
deactivation of haloperidol-induced catalepsy in the rat. Society for Neuroscience,
Toronto, ON, Canada.
17. Pellis, S. M., Pellis, V. C., & Lennon, R. L. (1988). Nicotine enhances playful attack
in the playfighting by juvenile rats. Society for Neuroscience, Toronto, ON, Canada.
18. Natoli, E., Pellis, V. C., Pellis, S. M., & Dewsbury, D. A. (1989). Analysis of the
body target areas approached by the male and defended by the female during copulatory
behavior in pine voles, Microtus pinetorum. Italian Society for the Study of Ethology,
Perugia, Italy, June.
19. Marshall, H. M., Pellis, S. M., Pellis, V. C., & Teitelbaum, P. (1989). Dopaminergic
drugs differentially affect attack versus defense in play fighting by juvenile rats. Society
for Neuroscience, Phoenix, AZ, USA, October.
28
20. Pellis, S. M., Pellis, V. C., & Nelson, J. E. (1989). Development of tactile righting
reflexes in the marsupial Dasyurus hallucatus. Society for Neuroscience, Phoenix,
Arizona, USA, October.
21. Salo, A. L., Pellis, S. M.,* Pellis, V. C., & Dewsbury, D. A. (1990). Differences in
play behavior of two species of voles (Microtus pennsylvanicus and M. pinetorum).
Animal Behavior Society, Binghampton, NY, USA, June.
22. Pellis, S. M. (1990). Targets and tactics: Conspecific attack versus predatory attack in
Northern grasshopper mice Onychomys leucogaster. International Society for Research on
Aggression, Banff, AB, Canada, June.
23. Pellis, S. M., & Pellis, V. C. (1990). The role of defense as an expression of
dominance in the play fighting of male rats. International Society for Research on
Aggression, Banff, AB, Canada, June.
24. Teitelbaum, P., Pellis, V. C., & Pellis, S. M. (1990). Can allied reflexes promote the
integration of a robot's behavior? Simulation of Adaptive Behavior: From Animals to
Animats, Paris, France, September.
25. Porter, J. D., Pellis, S. M., & Meyer, M. E. (1990). An open field activity analysis of
unilaterally labyrinthectomized rats. Society for Neuroscience, St. Louis, MO, USA,
October.
26. Pellis, S. M. (1991). The brain behind the playing rat. Canadian Society for Brain,
Behaviour, and Cognitive Science, Calgary, AB, June.
27. Pellis, S. M., & Pellis, V. C. (1991). Disassociation of juvenile play fighting and adult
dominance in male rats treated neonatally with testosterone propionate. Society for
Neuroscience, New Orleans, LA, USA, November.
28. Cordover, A. J., Pellis, S. M., & Teitelbaum, P. (1991). Haloperidol exaggerates
proprioceptive-tactile support reflexes and diminishes vestibular dominance over them.
Society for Neuroscience, New Orleans, LA, USA, November.
29. Whishaw, I. Q., Gorny, B. P., & Pellis, S. M. (1991). Rat skilled reaching: Movement
components and neocortical organization. Society for Neuroscience, New Orleans, LA,
November.
30. Faulkes, Z., Paul, D. H., & Pellis, S. M. (1991). Digging by the sand crab
Blepharipoda occidentalis. Society for Neuroscience, New Orleans, LA, USA, November.
31. Pellis, S. M., Pellis, V. C., & Gaudry, B.-D. (1992). Individual differences in
temperament influence the development of play fighting and dominance in male rats
(Rattus norvegicus). Animal Behavior Society, Kingston, ON, Canada, June.
29
32. Pellis, S. M., Whishaw, I. Q., McKenna, M. M., & Castañeda, E. (1992). The role of
the striatum in organizing complex behaviors as revealed by play in neonatally 6-OHDA
depleted rats. Society for Neuroscience, Anaheim, CA, USA, October.
33. Whishaw, I. Q., Tetzlaff, W., Pellis, S. M., & Gorny, B. (1992). Pyramidal tract
lesions and skilled reaching in the rat. Society for Neuroscience, Anaheim, CA, USA,
October.
34. Field, E., & Pellis, S. M.* (1993). Differential effects of amphetamine on the attack
and defense components of play fighting in rats. Society for Neuroscience, Washington,
DC, USA, November.
35. McKenna, M. M., Pellis, V. C., Prusky, G., & Pellis, S. M.* (1993). Modulating
effects of vision on juvenile play behavior in rats. Society for Neuroscience, Washington,
DC, USA, November.
36. Pellis, S. M., & Pellis, V. C. (1994). The development of righting when falling from a
bipedal standing posture: Evidence for the dissociation of dynamic and static righting
reflexes in rats. Canadian Society for Brain, Behaviour, & Cognitive Science, Vancouver,
B. C., Canada, June.
37. Lambert, T., & Pellis, S. M.* (1994). Hunger increases object manipulation and
decreases social play in rats. Canadian Society for Brain, Behaviour, & Cognitive Science,
Vancouver, B. C. Canada, June.
38. Field, E. F., Pellis, S. M.,* & Whishaw, I. Q. (1994). An analysis of sex differences in
the movement patterns used during the food wrenching and dodging paradigm. Canadian
Society for Brain, Behaviour, & Cognitive Science, Vancouver, B. C., Canada, June.
39. Pellis, S. M., MacDonald, N., & Michener, G. (1994). The lateral display as a combat
tactic in Richardson’s ground squirrels, Spermophilus richardsonii. International Society
for Research on Aggression, Delray Beach, FL, USA, July.
40. Smith, L. K., Field, E. F., Forgie, M. L., & Pellis, S. M.* (1994). Effects of postweaning castration on the structure of play fighting in male rats. International Society for
Research on Aggression, Delray Beach, FL, USA, July.
41. Pellis, S. M., & Pellis, V. C. (1994). A comparative analysis of the so-called primate
play face. Animal Behavior Society, Seattle, WA, USA, July.
42. MacDonald, N. L., Pellis, S. M.,* & Michener, G. R. (1994). Sex differences in play
behaviour of juvenile Richardson’s ground squirrels (Spermophilus richardsonii ). Animal
Behavior Society, Seattle, WA, USA, July.
43. Pellis, S. M., & Field, E. F. (1995). Difference or differences in the play fighting of
male and female rats? The sex of play revisited. Invited paper, International Behavioral
30
Development Symposium: Biological Basis of Sexual Orientation & Sex-Typical
Behavior. Minot, ND, USA, May.
44. Pellis, S. M. (1995). Targets and tactics: The analysis of decision making in animal
combat. Plenary presentation, International Ethological Conference, Honolulu, HI, USA,
August.
45. Pellis, S. M., Pellis, V. C., & Field, E. F. (1995). Spontaneous turning behavior in
male and female adult rats: The effects of neonatal castration and testosterone treatment
on sex typical patterns of turning. Society for Neuroscience, San Diego, CA, USA,
November.
46. Pellis, V. C., Field, E. F., & Pellis, S. M.* (1995). The influence of sex and neonatal
hormone exposure on locomotion in adult rats. Society for Neuroscience, San Diego, CA,
USA, November.
47. Field, E. F., Whishaw, I. Q., & Pellis, S. M.* (1995). The relationship between sex,
neonatal hormone exposure and movement patterns in a competitive feeding paradigm
using adult rats. Society for Neuroscience, San Diego, CA, USA, November.
48. Pellis, S. M., & Pellis, V. C. (1996). On growing young: The onset of play fighting in
rats. Spring Conference on Behaviour & Brain, Fernie, B.C., Canada, May.
49. Field, E. F., Whishaw, I. Q., & Pellis, S. M.* (1996). A kinematic analysis of sextypical movement patterns used during evasive dodging to protect a food item: The role of
gonadal hormones. Spring Conference on Behaviour & Brain, Fernie, B. C., Canada, May.
50. Coles, B., Miklyaeva, E. I., Pellis, S. M., & Whishaw, I. Q. (1996). Impairments in
spontaneous eating following unilateral dopamine depletions in rats: An analogue of
human hemiparkinson’s disease. Spring Conference on Behaviour & Brain, Fernie, B. C.,
Canada, May.
51. Pellis, S. M., & Pellis, V. C. (1996). The structure-function interface in the analysis of
play fighting. Invited paper for the symposium on “Animal Play”, Animal Behavior
Society, Flagstaff, AZ, USA, August.
52. Smith, L. K., & Pellis, S. M.* (1996). Age-related effects of gonadal hormones on
play fighting and dominance in male rats (Rattus norvegicus). Animal Behavior Society,
Flagstaff, AZ, USA, August.
53. Pellis, S. M., Pellis, V. C., & Smith, L. K. (1996). The ontogeny of playful defense in
rats. International Society for Research on Aggression, Strasbourg, France, August.
54. Smith, L. K., & Pellis, S. M.* (1996). Making your behaviour match your partner:
Play fighting between familiar and unfamiliar male and female rats (Rattus norvegicus).
International Society for Research on Aggression, Strasbourg, France, August.
31
55. Pellis, S. M., Field, E. F., & Whishaw, I. Q. (1996). Sex differences in postural
compensation to an inclined plane during haloperidol catalepsy: The role of neonatal
androgen exposure. Society for Neuroscience, Washington, DC, USA, November.
56. Field, E. F., Pellis, S. M.*, & Whishaw, I. Q. (1996). Sex differences in the
organization of lateral movements are preserved during apomorphine induced stereotypy.
Society for Neuroscience, Washington, DC, USA, November.
57. Martens, D. J., Field, E. F., Pellis, V. C., & Pellis, S. M.* (1996). The development of
rotatory movements around the longitudinal axis during contact-righting: Sex differences
in the organization of rotatory movements. Society for Neuroscience, Washington, DC,
USA, November.
58. Pellis, S. M., & Pellis, V. C. (1997). Signalling play: Play fighting in primates. Spring
Conference on Behaviour & Brain, Fernie, B.C., Canada, March.
59. Woodward, N., Whishaw, I. Q., Miklyaeva, E. I., & Pellis, S. M. (1997). What do
hemiParkinson rats do in the Montoya task? Spring Conference on Behaviour & Brain,
Fernie, B.C., Canada, March.
60. Field, E. F., Pellis, S. M.*, & Whishaw, I. Q. (1997). Sex differences in response to a
postural challenge during haloperidol induced catalepsy: The role of gonadal hormones.
Spring Conference on Behaviour & Brain, Fernie, B.C., Canada, March.
61. Sarna, J., Whishaw, I. Q., & Pellis, S. M*. (1997). Rodent-typical and speciesspecific limb use in eating: A comparative analysis of ten species. Spring Conference on
Behaviour & Brain, Fernie, B.C., Canada, March.
62. Smith, L. K., & Pellis, S. M.* (1997). The pubertal transition in the play fighting of
rats. What regulates it? Spring Conference on Behaviour & Brain, Fernie, B.C., Canada,
March.
63. Field, E. F., Whishaw, I. Q., & Pellis, S. M.* (1997). Seemingly paradoxical jumping
in cataleptic haloperidol-treated rats is sexually dimorphic: The organizational and
activational influences of gonadal steroid hormones on postural support. Joint meeting of
the Society for Behavioral Neuroendocrinology and the Conference on Reproductive
Behavior, Baltimore, MD, USA, May.
64. Pellis, S. M., Pellis, V. C., & Thierry, B. (1997). The use of the bared-teeth display as
a play signal in Tonkean macaques Macaca tonkeana. International Ethological
Conference, Vienna, Austria, August.
65. Woodward, N. C., Miklyaeva, E. L., Pellis, S. M., & Whishaw, I. Q. (1997). Analysis
of limb use by unilateral DA-depleted rats in the staircase test: Evidence for multiple
compensatory strategies. Society for Neuroscience, New Orleans, LA, USA, November.
32
66. Field, E. F., Whishaw, I. Q., & Pellis, S. M.* (1997). Lack of experience during the
juvenile phase does not affect the development of sex-typical movement patterns. Society
for Neuroscience, New Orleans, LA, USA, November.
67. Pellis, S. M. (1998). Play: It’s not just sex anymore. Plenary presentation, Spring
Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
68. Iwaniuk, A. N., Pellis, S. M., & Whishaw, I. Q. (1998). Comparative food handling in
three small carnivores. Animal Behavior Society, Carbondale, IL, USA, July.
69. Iwaniuk, A. N., Pellis, S. M., & Whishaw, I. Q. (1998). The role of the spinocortical
tract in the evolution of digital dexterity. A phylogenetic analysis of the Heffner and
Masterton data set. Society for Neuroscience, Los Angeles, CA, USA, November.
70. Pellis, S. M. (1999). Sex differences in play fighting: The missing half of the story.
Spring Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
71. Dean, K., MacNeil-Alcock, A., & Pellis, S. M.* (1999). What rat is that? It’s all in
how you crack your seeds. Spring Conference on Behaviour & Brain, Fernie, B.C.,
Canada, February.
72. Iwaniuk, A. N., Pellis, S. M., & Whishaw, I. Q. (1999). Are brainy carnivores handy?
Spring Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
73. Davis, J. L., Pellis, S. M., Whishaw, I. Q., & Suchowersky, O. (1999). Pallidal DBS
normalizes upper limb movement coordination in individuals with PD. International
Congress on Parkinson’s Disease, Vancouver, B. C., July.
74. Ballermann, M., Tompkins, G., Pellis, S. M., & Whishaw, I. Q. (1999). A new
method of testing sensory ability in the forepaw. Society for Neuroscience, Miami Beach,
Fl., USA, October.
75. Pellis, S. M. (2000). Rough play and the female rat: A non-traditional mechanism for
the regulation of a supposedly male-typical trait. Invited paper, International Behavioral
Development Symposium: Biological Basis of Sexual Orientation and Sex-Typical
Behavior. Minot, ND, USA, May.
76. Ballerman, M., McKenna, J., Pellis, S. M., & Whishaw, I. Q. (2000). The dorsal
columns: Impaired forepaw tactile discrimination vs. preserved reaching success and force
application in the rat. Society for Neuroscience, New Orleans, LA, USA, November.
77. Pellis, S. M. (2001). Is play multi-orgasmic? Spring Conference on Behaviour &
Brain, Fernie, B.C., Canada, February.
33
78. Foroud, A., & Pellis, S. M.* (2001). A pin is a pin is a pin, or is it? Spring
Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
79. Hines, D. J., Wallace, D. G., Pellis, S. M., & Whishaw, I. Q. (2001). Selective
impairment of deductive reckoning in rats with vestibular lesions. Society for
Neuroscience, San Diego, CA, USA, November.
80. Pellis, S. M. (2002). Follow the bouncing cricket. Spring Conference on Behaviour &
Brain, Fernie, B.C., Canada, February.
81. Foroud, A., & Pellis, S. M.* (2002). Experience and cortical control over the pubertal
transition to rougher play fighting in rats. Spring Conference on Behaviour & Brain,
Fernie, B.C., Canada, February.
82. Komorowska, J., & Pellis, S. M.* (2002). The escalation of aggression in the degu,
Octodon degu. Spring Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
83. Pellis, S. M., & Harvey, S. (2002). Fixed action pattern or modal action pattern: A
cricket’s perspective. Canadian Society of Zoologists, Lethbridge, AB, Canada, May.
84. Pellis, S. M. (2002). The development of aggression and affiliation: The role of play
fighting. Invited Keynote Speaker, International Society for the Study of Aggression,
Montreal, QE, Canada, July.
85. Pellis, S. M. (2002). Evolving levels of control in play fighting. Invited paper,
symposium on “Play”, International Society for the Study of Behavioral Development,
Ottawa, ON, Canada, August.
86. Doan, J., Brown, L. A., Whishaw, I. Q., Suchowersky, O., & Pellis, S. M. (2002).
Target characteristics affect reaching kinematics for parkisonian patients. Society for
Neuroscience, Orlando, FL, USA, October.
87. Pellis, S. M., Hastings, E., DeLong, M. S., Gibb, R. L., & Kolb, B. E. (2002).
Perinatal orbital frontal lesions in rats disrupt social behavior but allow sparing of
cognitive and motor behaviors. Society for Neuroscience, Orlando, FL, USA, October.
88. Pellis, S. M. (2003). Action patterns and the economy of movement. Spring
Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
89. Reinhart, C., & Pellis, S. M.* (2003). The development of social play in kindlingresistant and kindling-prone rats. Spring Conference on Behaviour & Brain, Fernie, B.C.,
Canada, February.
90. Field, E. F., Whishaw, I. Q., & Pellis, S. M.* (2003). The absence of pre- and postpubertal hormones in the development of female-typical patterns of dodging. Spring
Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
34
91. Field, E. F., Watson, N. V., Whishaw, I. Q., & Pellis, S. M.* (2003). The
development of male-typical patterns of dodging to protect a food item: Do tfm-affected
male rats exhibit male-typical patterns of movement? International Society for Behavioral
Neuroendocrinology, Cincinnati, OH, USA, June.
92. Hamilton, D. A., Silasi, G., Pellis, S. M., & Kolb, B. (2003). Experience-dependent
synaptogenesis in motor, occipital and orbitofrontal cortex of the rat. Society for
Neuroscience, New Orleans, LA, USA, November.
93. Doan, J. B., Brown, L. A., Whishaw, I. Q., Suchowersky, O., & Pellis, S. M. (2003).
Improved timing of reach components with L-DOPA in Parkinson’s disease: A kinematic
analysis. Society for Neuroscience, New Orleans, LA, USA, November.
94. Pierchrka, D. M., Kleim, J. A., Pellis, S. M., & Whishaw, I. Q. (2003). Partial
pyramidal tract lesions decrease forelimb representation in motor cortex and produce a
qualitative forelimb impairment in the rat. Society for Neuroscience, New Orleans, LA,
USA, November.
95. Pellis, S. M., & Pellis, V. C. (2004). Scratching and the evolution of behaviour
patterns. Spring Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
96. Reinhart, C., & Pellis, S. M.* (2004). What is the currency of reciprocity in play
fighting? Spring Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
97. Field, E. F., Pellis, S. M., & Whishaw, I. Q. (2004). The Whishaw reaching task: Sex
differences in posture and orientation to a single food pellet. Spring Conference on
Behaviour & Brain, Fernie, B.C., Canada, February.
98. Hamilton, D. A., Pellis, S. M., & Savage, D. D. (2004). Abnormal social behavior in
rats exposed to moderate levels of ethanol in utero. Research Society on Alcoholism,
Vancouver, B.C., Canada, June.
99. Doan, J., Cooper, S., Tiede, S., Pellis, S. M., Whishaw, I. Q., Suchowersky, O., &
Brown, L. A. (2004). Manipulating visual information alters postural stability among
Parkinson’s patients. North American Society for the Psychology of Sport & Physical
Activity, Vancouver, B. C., June.
100. Reinhart, C. J., McIntyre, D. C., Pellis, S. M., & Kolb, B. (2004). An examination of
dendritic morphology in seizure-prone (fast-kindling) and seizure-resistant (slow-kindling)
rat strains. Society for Neuroscience, San Diego, CA, October.
101. Hamilton, D. A., Silasi, G., Carroll, C. E., Pellis, S. M., & Kolb, B. (2004).
Experience differentially affects orbital and medial prefrontal cortex of the rat. Society for
Neuroscience, San Diego, CA, October.
35
102. Pellis, S. M., Field, E. F., Whishaw, I. Q., & Watson, N. V. (2004). A
developmental analysis of changes in playful attack and defense by tfm-affected male rats
and their wild-type controls. Society for Neuroscience, San Diego, CA, October.
103. Field, E. F., Metz, G. A., Pellis, S. M.,* & Whishaw, I. Q. (2004). A kinematic
analysis of sex differences in postural support during horizontal and vertical movements in
the cylinder test after a unilateral 6-OHDA lesion. Society for Neuroscience, San Diego,
CA, October.
104. Pellis, S. M. (2005). The evolution of social play. Invited paper at a special
conference on “Four dimensions of childhood: Brain. Mind, Culture, and Time”,
sponsored by the Foundation for Psycho-Cultural Research. UCLA, Los Angeles,
February.
105. Field, E. F., Pellis, S. M., & Whishaw, I. Q. (2005). Sex differences in activity in a
6-OHDA model of attention deficit hyperactivity disorder. Spring Conference on
Behaviour & Brain, Fernie, B.C., Canada, February.
106. Arelis, C., & Pellis, S. M.* (2005). The effects of stress on play. Spring Conference
on Behaviour & Brain, Fernie, B.C., Canada, February.
107. Reinhart, C., & Pellis, S. M.* (2005). Play: It’s not all in your genes. Spring
Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
108. Espinoza, W., Pellis, S. M., & Hurd, P. (2005). Sexual differentiation and adult
behaviour in a cichlid fish. Spring Conference on Behaviour & Brain, Fernie, B.C.,
Canada, February.
109. Cooper, S., Doan, J., Pellis, S. M., Whishaw, I. Q., Suchowersky, O., & Brown, L.
(2005). Current treatment interventions for Parkinson’s disease and the integration of
visual information for postural control: Subthalamic brain stimulation versus levodopa
therapy. Spring Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
110. Kurek, J., Doan, J., Whishaw, I. Q., Pellis, S. M., Suchowersky, O. & Brown, O.
(2005). Deficits in gait initiation associated with Parkinson’s disease exacerbated by
postural threat. Spring Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
111. Doan, J., Suchowersky, O., Whishaw, I. Q., Pellis, S. M., & Brown, L. (2005).
Reaching kinematics not matched by postural context by Parkinson’s disease patients.
Spring Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
112. Lingle, S., Rendall, D., & Pellis, S. M. (2005). Altruism and recognition in the
antipredator defense of deer fawns. Animal Behavior Society, Snowbird, Utah, August.
113. Field, E. F., & Pellis, S. M.* (2005). Sex differences in movement: The role of
gonadal steroids and body morphology. Invited paper, 3rd International Behavioral
36
Development Symposium: Biological Basis of Sexual Orientation & Sex-Typical
Behavior. Minot, ND, USA, August.
114. Pellis, S. M., Field, E. F., Sherren, N., Yamazaki, H., & Whishaw, I. Q. (2005). The
dissociation of hyperactivity in a neonatal dopamine depletion model of ADHD by sex,
age and task. Society for Neuroscience, Washington, DC, November (Press Book)
115. Field, E. F., Sherren, N., Pellis, S. M., & Whishaw, I. Q. (2005). The neonatal 6OHDA depletion model of ADHD: Do developmental manipulations of the dopaminergic
system affect sex-typical patterns of movement organization? Society for Neuroscience,
Washington, DC, November (Press Book)
116. Pellis, S. M. (2006). How to build a playful brain. Winter Animal Behavior
Conference, Steamboat Springs, CO, January.
117. Pellis, S. M. (2006). What does a playful brain circuit look like? Spring Conference
on Behaviour & Brain, Fernie, B.C., Canada, February.
118. Reinhart, C., Thierry, B., & Pellis, S. M.* (2006). Do all monkeys play the same?
Spring Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
119. Field, E. F., Metz, G. A., Whishaw, I. Q., & Pellis, S. M. * (2006). The choice of
sex-typical patterns of movement is dependent on an intact nigrostriatal system. Spring
Conference on Behaviour & Brain, Fernie, B.C., Canada, February.
120. Lingle, S., Rendall, D., and Pellis, S. M. (2006). Altruism and recognition in the
antipredator defence of deer fawns. Canadian Society of Zoologists, Edmonton, AB, May.
121. Lingle, S., Rendall, D., & Pellis, S. M. (2006). Behavioural discrimination and
altruism in the antipredator defence of deer. International Society of Behavioral Ecology,
Tours, France, July.
122. Lingle, S., Rendall, D., & Pellis, S. M. (2006). Why help? The evolution of altruistic
antipredator defence in deer. The 6th International Deer Biology Congress, Prague, Czech
Republic. August 2006.
123. Lingle, S., Wilson, W. F., & Pellis, S. M. (2006). When prey fight back: higher
levels of aggressive defence by mule deer than whitetail females lowers vulnerability of
mule deer fawns to coyotes early in life. The 6th International Deer Biology Congress,
Prague, Czech Republic, August.
124. Pellis, S. M., & Pellis, V. C. (2006). When does a head shake signal play? A
developmental perspective from a troop of black-handed spider monkeys (Ateles
geoffroyi). International Society for Comparative Psychology, Christchurch, New Zealand,
September.
37
125. Pellis, S. M., Field, E. F., Sherren, N., Yamazaki, H., Pittman, Q. J., & Whishaw, I.
Q. (2006). The effects of neonatal dopaminergic lesions on the development of
hyperactivity and social play in male and female rats: Implications for the study of
attention deficit hyperactivity disorder. Society for Neuroscience, Washington, DC,
November.
126. Pellis, S. M., & Pellis, V. C. (2007). Cooperation and competition: The many paths
to fairness in play fighting. Winter Animal Behavior Conference, Steamboat Springs, CO,
January.
127. Pellis, S. M. & Pellis, V. C. (2007). Play and the neural underpinnings of fairness.
Canadian Spring Conference on Brain and Behaviour, Fernie, British Columbia, February.
128. Bell, H., Kolb, B., & Pellis, S. M.* (2007). Play: A mind-altering experience.
Canadian Spring Conference on Brain and Behaviour, Fernie, British Columbia, February.
129. Reinhart, C., Thierry, B. & Pellis, S. M.* (2007). Targets and tactics of play fighting
in juvenile Tonkean macaques. Canadian Spring Conference on Brain and Behaviour,
Fernie, British Columbia, February.
130. Reinhart, C., Thierry, B., Pellis, V. C., & Pellis, S. M.* (2007). Asymmetry and
fairness in the play fighting of Tonkean macaques. International Conference for Ethology,
Halifax, Newfoundland, August.
131. Bell, H. C., Kolb, B. E., & Pellis, S. M.* (2007). It’s not child’s play: Brain
development is altered by horsing around. Society for Neuroscience, San Diego,
November (Press Book)
132. Pellis, S. M. (2008). Brain begets play and play begets brain. Winter Animal
Behavior Conference, Steamboat Springs, CO, January.
133. Pellis, S. M. (2008). Play and the development of the socially competent brain.
Invited address to the Canadian Institute for Advanced Research (CIFAR), Experiencebased Brain and Biological Development (EBBD) Program meeting, Vancouver, BC,
February.
133. Reinhart, C. J., Thierry, B., Pellis, V. C., & Pellis, S. M.* (2008). Play fighting in
Tonkean and Japanese macaques: Same game, different rules. Canadian Spring
Conference on Brain and Behaviour, Fernie, British Columbia, February.
134. Bell, H., Kolb, B., & Pellis, S. M.* (2008). Playing with the brain: Relationships
between frontal cortex and social experience. Canadian Spring Conference on Brain and
Behaviour, Fernie, British Columbia, February.
135. Pellis, S. M., & Pellis, V. C. (2008). Why do spider monkeys shake their heads?
Canadian Spring Conference on Brain and Behaviour, Fernie, British Columbia, February.
38
136. Pellis, S. M. (2008). Rough and tumble play and the development of social
competency. Southern Alberta Council for Public Affairs (SACPA), Lethbridge, AB,
March.
137. Pellis, S. M. (2008). Rough and tumble play and the development of the social brain.
McGill University Health Challenge Think Tank. Active Living & Energy Balance.
Halting Childhood Obesity: Towards Sustainable Economic, Environmental, Cultural &
Health Footprints. Montreal, QC, October.
138. Bell, H. C., Kolb, B. E., & Pellis, S. M.* (2008). The medial prefrontal cortex and
the regulation of social movements. Society for Neuroscience, Washington, DC,
November
139. Pellis, S. M. (2008). Ethology and Animal Welfare. Invited address for the Scientists
Center for Animal Welfare: Animal Behavior, Welfare & Science: IAUC Roles. San
Antonio, TX, November.
140. Bell, H. C., Kolb, B. E., & Pellis, S. M.* (2009). Sociality and the medial prefrontal
cortex. Canadian Spring Conference on Brain and Behaviour, Fernie, British Columbia,
February.
141. Bell, H. C., & Pellis, S. M.* (2009). Perceptual Control Theory – Bernard and
Darwin together at last! Graduate Students’ Association Multidisciplinary Conference.
Lethbridge, Alberta, March.
142. Pellis, S. M. (2009). Rough and tumble play and the development of a socially
competent brain. Invited public lecture for Brain Awareness Week, Yates Memorial
Centre, Lethbridge, AB, March.
143. Supina, R., Smith, L., Muhammed, A., Pellis, S., & Metz, G. (2009). Strategies in
play fighting in a rat model of Parkinson’s disease suggest altered social behaviour. 13th
Congress of the European Federation of Neurological Societies, Florence, Italy,
September.
144. Pellis, S. M. (2009). Nurturing the social brain. Invited paper, Workshop on Rodent
Social Behaviour, Champalimaud Neuroscience Programme at the Instituto Gulbenkian de
Ciencia, Lisbon, Portugal, September.
145. Bell, H. C., & Pellis, S. M.* (2009). Perceptual control theory: The undiscovered
revolution? Society for Neuroscience, Chicago, October.
146. Pellis, S. M. (2010). From behaviour to brain, and back again. Bryan Kolb
Festschrift, Lethbridge, AB, February.
39
147. Bell, H. C., & Pellis, S. M.* (2010). Robbing and dodging in rats: Ballistic behaviour
versus flexible control. Canadian Spring Conference on Brain and Behaviour, Fernie,
British Columbia, February.
148. Pellis, S. M. (2010). Rough-and-tumble play and developing a social brain. Early
Childhood Education Conference. Lethbridge, AB, March.
149. Pellis, S. M. (2010). Bench to backyard: Kids need to play. Canadian Association of
School Social Workers & Attendance Counsellors Conference. Winnipeg, MB, April.
150. Pellis, S. M., & Pellis, V. C. (2010). From foot to hand: The evolution of head
scratching in primates. International Society for Comparative Psychology, Awaji, Japan,
May.
151. Pellis, S. M., Pellis, V. C., & Bell, H. C. (2010). Combat: Targets as controlled
variables, tactics as compensatory actions. Control Systems Group International
Conference. Manchester, UK, July.
152. Bell, H. C., & Pellis, S. M.* (2010). Back off – get your own sandwich! Distance
regulation in a defensive action by rats. Control Systems Group International Conference.
Manchester, UK, July.
153. VanderLaan, D. P., Pellis, S. M., & Vasey, P. L. (2010). Mounting and social play in
juvenile Japanese macaques. International Primatology Society, Kyoto, September.
154. Pellis, S. M. (2011). Behavior as an epiphenomenon: The cybernetics of animal
action. Winter Animal Behavior Conference, Steamboat Springs, CO, January.
155. Himmler, B., Modlinska, K., Stryjek, R., Pisula, W., Kolb, B., & Pellis, S. M.*
(2011). Run Rattus Run. Canadian Spring Conference on Brain and Behavior, Fernie,
British Columbia, February.
156. Pellis, S. M. (2011). Play fighting, reciprocity, uncertainty and the making of a social
brain. Invited plenary presented at the Animal Behavior Conference, Bloomington, IN,
April.
157. Himmler, B., Stryjek, R., Modlinska, K., Pisula, W., Kolb, B., & Pellis, S. M.*
(2011). How the Wild Things Play. Alberta Graduate Conference, Calgary, Alberta, May
158. Pellis, S. M. (2011). Peer-peer play, social competence and academic performance.
Invited presentation to the Alberta Education Research Seminar: The Early Years – Brain
Research and the Implications for Practice, Edmonton, May
159. Pellis, S. M. (2011). The anatomy of behavior: Lessons learned from aye-ayes.
Invited presentation to be presented for the Philip Teitelbaum Festschrift, Washington,
DC, November
40
160. Himmler, B., Stryjek, R., Modlinska, K., Pisula, W., Kolb, B., & Pellis, S. M.*
(2011). Play gone wild: Contrasting sex differences in the social play of wild and domestic
rats. To be presented at the Society for Neuroscience, Washington, DC, November
*I am the senior author in these presentations. The work was supported by grants awarded
to me, and conducted by students in my laboratory under my supervision.
Last updated 08/03/2011