Chapter Objectives: Chapters 23 and 24 Species and Populations 1. 2. 3. 4. 5. 6. 7. 8. 9. 10. 11. 12. 13. 14. 15. 16. 17. 18. 19. 20. 21. 22. 23. 24. 25. Explain what is meant by the modern synthesis Explain how microevolutionary change can affect a gene pool state the Hardy-Weinberg theorem Write the general Hardy-Weinberg equation and use it to calculate allele and genotype frequencies Explain the consequences of Hardy-Weinberg equilibrium Demonstrate that a disequilibrium population requires only one generation of random mating to establish Hardy-Weinberg equilibrium Describe the usefulness of the Hardy-Weinberg equilibrium model to population genetics List the conditions a population must meet in order to maintain HardyWeinberg equilibrium Explain how genetic drift, gene flow, mutation, nonrandom mating, and natural selection can cause microevolution Explain the role of population size in genetic drift Distinguish between the bottleneck effect and the founder effect Explain why mutation has little quantitative effect on a large population Describe how inbreeding and assortive mating affect a population's allele frequencies and genotype frequencies List factors that produce geographic variation among closely related populations Explain why even though mutation can be a source of genetic variability, it contributes a negligible amount to genetic variation in a population Explain how genetic variation may be preserved in a natural population Describe the neutral theory of molecular evolution and explain how changes in gene frequency may be nonadaptive Explain what is meant by selfish DNA Explain the concept of relative fitness and its role in adaptive evolution Explain why the rate of decline for a deleterious allele depends upon whether the allele is dominant or recessive to the more successful allele Describe what selection acts on and what factors contribute to the overall fitness of a genotype Give examples of how an organism's phenotype may be influenced by the environment Distinguish among stabilizing selection, directional selection, and diversifying selection Define sexual dimorphism and explain how it can influence evolutionary change Give at leas 4 reasons why natural selection cannot breed perfect organisms ********************* 26. Distinguish between anagenesis and cladogenesis 27. Define morphospecies and explain how this concept can be useful to biologists 28. Define biological species (E. Mayr) 29. Describe some limitations of the biological species concept 30. Explain how gene flow between closely related species can be prevented 31. Distinguish between prezygotic and postzygotic isolating mechanisms 32. Describe 5 prezygotic isolating mechanisms and give an example of each 33. Explain why many hybrids are sterile 34. Explain how hybrid breakdown maintains separate species even if gene flow occurs 35. Distinguish between allopatric and sympatric speciation 36. Explain the allopatric speciation model and describe the role of intraspecific variation and geographical isolation 37. Explain why peripheral isolates are susceptible if geographic barriers arise 38. Describe the adaptive radiation model and use it to describe how it might be possible to have many sympatric closely related species even if geographic isolation is necessary for them to evolve 39. Define sympatric speciation and explain how polyploidy can cause reproductive isolation 40. Distinguish between autopolyploidy and allopolyploidy 41. List some points of agreement and disagreement between the 2 schools of thought about the tempo of speciation (gradualism vs. punctuated equilibrium) 42. Describes the origins of evolutionary novelty Chapter Terms: Chapter 23 Terms population genetics founder effect hybrid vigor modern synthesis gene flow frequency-dependent selection population mutation species inbreeding neutral variation Darwinian fitness gene pool assortative mating relative fitness genetic structure natural selection stabilizing selection Hardy-Weinberg theorem polymorphism directional selection geographical variation Hardy-Weinberg equilibrium diversifying selection cline sexual dimorphism Hardy-Weinberg equation microevolution balanced polymorphism sexual selection heterozygote advantage bottleneck effect Chapter 24 Terms macroevoluton morphological species concept autopolyploid speciation recognition species concept allopolyploid anagenesis cohesion species concept hybrid zone phyletic evolution ecological species concept punctuated equilibrium cladogenesis evolutionary species concept exaptation branching evolution allopatric speciation paedomorphosis species sympatric speciation allometric growth prezygotic barriers adaptive radiation heterochrony postzygotic barriers polyploidy homeosis Chapter Outline Framework A. Population Genetics 1. The modern evolutionary synthesis integrated Darwinian selection and Mendelian inheritance 2. The genetic structure of a population is defined by its allele and genotype frequencies 3. The Hardy-Weinberg theorem describes a nonevolving population B. Causes of Microevolution 1. Microevolution is a generation-to-generation change in a population's allele or genotype frequencies 2. The 5 causes of microevolution are genetic drift, gene flow, mutation, nonrandom mating, and natural selection C. Genetic Variation, the Substrate for Natural Selection 1. Genetic variation occurs within and between populations 2. Mutation and sexual recombination generate genetic variation 3. Diploidy and balanced polymorphism preserve variation D. Natural selection as the Mechanism of Adaptive Evolution 1. Evolutionary fitness is the relative contribution an individual makes to the gene pool of the next generation 2. The effect of selection on a varying characteristic can be stabilizing, directional, or diversifying 3. Sexual selection may lead to pronounced 2y differences between the sexes 4. Natural selection cannot fashion perfect organisms E. What is a Species? 1. The biological species concept emphasizes reproductive isolation 2. Prezygotic and postzygotic barriers isolate the gene pools of biological species 3. The biological species concept does not work in all situations F. Modes of Speciation 1. Geographical isolation can lead to the origin of species: allopatric speciation 2. A new species can originate in the geographical midst of the parent species: sympatric speciation 3. Genetic change in populations can account for speciation 4. The punctuated equilibrium model has stimulated research on the tempo of speciation G. The Origin of Evolutionary Novelty 1. Most evolutionary novelties are modified versions of older structures 2. Genes that control development play a major role in evolutionary novelty 3. An evolutionary trend does not mean that evolution is goal-oriented