Chapter 16 Population Genetics and Speciation Section 1 Vocabulary Pretest 1. Population Genetics 2. Microevolution 3. Gene Pool 4. Allele Frequency 5. Phenotype Frequency Population Genetics _________________________ is the study of evolution from a genetic point of view (it is the study of microevolution) _______________________—a change in the collective genetic material of a population ___________________–members of the same species that can interbreed. It is the smallest unit in which evolution occurs. Variation Populations show ________________________________________. Many quantitative traits (height and weight etc.) follow a ______________ _________________________. Causes of Variation ______________________________—amount of food, quality of food, etc. __________________________ ______________________—random changes in genes ______________________—reshuffling of genes ______________________ of gametes Gene Pool __________________ —total genetic information available in a population Allele and Phenotypic Frequency ____________________________—expressed as a percent: it is determined by ________________________________________________ _____________________________________ of all types in the population _____________________________—expressed as a percent: it is the number of individual with a particular _________________ divided by the total number of individuals in the population. Hardy-Weinberg Genetic Equilibrium Developed by ____________________German physician) and ________________________________(British mathematician) States that genetic frequencies in a population tend to ________________ ____________________from generation to generation ________________ ___________________________________________ It is based on a “hypothetical population” that is ____________________. Conditions of Hardy-Weinberg Equilibrium In Genetic Equilibrium: ____________________________ occur Population size remains __________________ The population is _______________________ Individuals ____________________________ Selection _____________________________ It is ______________________ that all five of the conditions in the Hardy-Weinberg Model will happen in the real world. Therefore, Genetic Equilibrium is _________________________________. It is a theoretical state that allows us to consider what forces could disrupt such balance (equilibrium). Section 2 Vocabulary Pretest 1. Immigration 2. Emigration 3. Gene Flow 4. Genetic Drift 5. 6. 7. 8. Sexual Selection Stabilizing Selection Disruptive Selection Directional Selection Disruption of Genetic Equilibrium Disruptions to the Hardy-Weinberg equilibrium can ___________________ _________________________. The five requirements for genetic equilibrium can be disrupted by the following outside forces: __________________________ __________________________ __________________________ __________________________ __________________________ Requirement #1: No Net Mutations Occur Mutations _______________________ at very low rates under normal conditions. Exposure to __________________ (mutation-causing agents, i.e. radiation and chemicals) can increase mutations rates. Mutations produce new alleles for a trait They can be __________________________________________ Helpful mutations are a vital part of evolution. Requirement #2: Population Size Remains Constant Individuals enter and leave populations constantly. Their “genes” move with them. This is called __________________________. Factors influencing gene flow include: ___________________—movement of individuals into a population __________________—movement of individuals out of a population ____________________________________ can also influence the movement of individuals into new populations ___________________________________ also remove or add genes from individuals to a population. Requirement #3 Population is Infinitely Large In nature, population sizes are restricted rather than infinitely large. _______________________ can occur in small populations of organisms ________________________—the random change in allele frequency in a population Significant changes can happen in small populations if even a single organism either fails to reproduce or reproduces too much. If the frequency of an ________________________ in a population, then (assuming you started with two alleles), there is only one left. All individuals will be ______________________ for that trait---__________ _________________________________ This __________________________ Ex: Northern Elephant Seal Bottleneck Effect Genetic Drift can lead to a _____________________________in which variations are reduced overtime. Requirement #4: Random Matings Organisms ____________________________________ in nature. Mate selection is influence by: _______________________________—choose mates nearby: can result in kinship mating _______________________________—choose mates with similar traits: reduces variation ______________________________—choose mates based on favorable traits Requirement #5: Selection Does Not Occur ___________________________—organisms with favorable traits are more likely to survive and reproduce, passing on their favorable genes to the next generation. It is an ongoing process in nature and an important disruption to equilibrium. _____________________ of Natural Selection: __________________________: individuals with the average form of a trait have the highest fitness. Ex: Lizard body size __________________________: individuals with either extreme variation of a trait have the highest fitness. Ex: Shell Color of Limpets _____________________________: individuals with one extreme of a trait have the highest fitness Ex: Nose and tongue lengths of anteaters Section 3 Vocabulary Pretest 1. Speciation 2. Geographic Isolation 3. Allopatric Speciation 4. Reproductive Isolation 5. Sympatric Speciation 6. Gradualism 7. Punctuated Equilibrium Speciation ______________________—formation of a new species _______________: a single kind of organism whose members are ________________________________________ to produce fully fertile offspring. Two types of speciation: _____________________________________________ Allopatric Speciation ____________________________: species arise as a result of geographic isolation. (Allopatric = different homelands) ___________________________—physical separation of members of a population Gene flow between the new subpopulations stops and the two begin to diverge Eventually, the become incompatible for mating, creating new species. Debate exists as to whether or not allopatric species are different enough to be considered new species. Sympatric Speciation _____________________________ —occurs when two subpopulations become reproductively isolated within the same geographic area. ___________________________—the inability of members of the same species to mate Can be caused by disruptive selection Two types: ____________________________________________ ___________________________: (different mating seasons, different mating calls, etc.) ______________________________: offspring do not fully develop, die, or are infertile. Rates of Speciation Two Theories: __________________ —slow change over millions of years ____________________________—short bursts of rapid change Evidence exists that suggests that both have taken place over time. The Hardy-Weinberg Equation Hardy and Weinberg went on to develop an equation that can be used to discover the probable genotype frequencies in a population and to track their changes from one generation to another. The equation is: ________________________________ p= frequency of the dominant allele q = frequency of the recessive allele Significance of the Hardy-Weinberg Equation ___________________________ allow geneticists to predict the probability of offspring genotypes for particular traits based on the known genotypes of their two parents The Hardy-Weinberg equation essentially allowed geneticists to do the _____________ ___________________________________. Before Hardy and Weinberg, it was thought that dominant alleles must, over time, wipe out recessive alleles _________________________________ According to this wrong idea, dominant alleles always increase in frequency from generation to generation. Hardy and Weinberg demonstrated that _______________________________________ ________________________________________________________________________