Big Idea 1 Review: The process of evolution explains the diversity and unity of life. Connecting evolutionary changes in a population over time to change(s) in the environment by describing 2–3 examples peppered moth, sickle cell anemia, DDT resistance in insects, introduction of nonnative species, introduction of predator, cataclysmic event The sickle-cell allele causes mutations in hemoglobin but also confers malaria resistance It exemplifies the heterozygote advantage Describing a limited set of given conserved features and core processes shared within and between domains of life DNA as the genetic material, a shared genetic code, common metabolic pathways (photosynthesis and CR, ETC’s), number of limbs, etc as evidence of common ancestry. Explaining how a phylogenetic representation reflects ancestral differences and similarities In addition to fossils, phylogenetic history can be inferred from morphological and molecular similarities in living organisms Organisms with very similar morphologies or similar DNA sequences are likely to be more closely related than organisms with vastly different structures or sequences In constructing a phylogeny, systematists need to distinguish whether a similarity is the result of homology or analogy Homology is similarity due to shared ancestry Analogy is similarity due to convergent evolution Describing how given data support the concept of a common ancestry within and between phylogenetic domains and for all life. Analyzing data related to questions of speciation and extinction throughout Earth’s history. Analysis may include (1) identifying patterns of speciation and/or extinction, (2 determining rates of speciation and/or extinction, and (3) connecting changes in gene frequency to speciation. Justifying the selection of data that address questions related to reproductive isolation and speciation pre- and post-zygotic and allopatric and sympatric isolation Describing a model that represents evolution within a population and providing evidence to support the description evolution due to genetic variation, such as antibiotic resistance, structure, or process, such as the brain, immune system, or linkage of a given population to common ancestors, through genetic, physiological, and morphological data Evaluating scientific hypotheses about the origin of life on Earth organic “soup” model, solid surface, and predicting how a hypothesis would be revised in light of new evidence “RNA World” hypothesis, new ideas about reducing atmosphere Protobionts are aggregates of abiotically produced molecules surrounded by a membrane or membrane-like structure Experiments demonstrate that protobionts could have formed spontaneously from abiotically produced organic compounds For example, small membrane-bounded droplets called liposomes can form when lipids or other organic molecules are added to water The “RNA World” and the Dawn of Natural Selection The first genetic material was probably RNA, not DNA RNA molecules called ribozymes have been found to catalyze many different reactions, including: Self-splicing Making complementary copies of short stretches of their own sequence or other short pieces of RNA Early protobionts with self-replicating, catalytic RNA would have been more effective at using resources and would have increased in number through natural selection Describing several sources of evidence from multiple scientific disciplines that support biological evolution Fossil records, morphological features, DNA and/or protein sequences, radioactive dating, distribution, or extant and extinct species The absolute ages of fossils can be determined by radiometric dating The magnetism of rocks can provide dating information Magnetic reversals of the magnetic poles leave their record on rocks throughout the world The Permian extinction killed about 96% of marine animal species and 8 out of 27 orders of insects It may have been caused by volcanic eruptions The Cretaceous extinction doomed many marine and terrestrial organisms, notably the dinosaurs It may have been caused by a large meteor impact Mass extinctions provided life with unparalleled opportunities for adaptive radiations into newly vacated ecological niches The First Prokaryotes Prokaryotes were Earth’s sole inhabitants from 3.5 to about 2 billion years ago Photosynthesis and the Oxygen Revolution The earliest types of photosynthesis did not produce oxygen Oxygenic photosynthesis probably evolved about 3.5 billion years ago in cyanobacteria The First Eukaryotes The oldest fossils of eukaryotic cells date back 2.1 billion years Endosymbiotic Origin of Mitochondria and Plastids The theory of endosymbiosis proposes that mitochondria and plastids were formerly small prokaryotes living within larger host cells The prokaryotic ancestors of mitochondria and plastids probably gained entry to the host cell as undigested prey or internal parasites In the process of becoming more interdependent, the host and endosymbionts would have become a single organism Key evidence supporting an endosymbiotic origin of mitochondria and plastids: Similarities in inner membrane structures and functions Both have their own circular DNA The Earliest Multicellular Eukaryotes Molecular clocks date the common ancestor of multicellular eukaryotes to 1.5 billion years The oldest known fossils of eukaryotes are of relatively small algae that lived about 1.2 billion years ago The “Cambrian Explosion” Most of the major phyla of animals appear in the fossil record of the first 20 million years of the Cambrian period Two animal phyla, Cnidaria and Porifera, are somewhat older, dating from the late Proterozoic Molecular evidence suggests that many animal phyla originated and began to diverge much earlier, between 1 billion and 700 million years ago Colonization of Land by Plants, Fungi, and Animals Plants, fungi, and animals colonized land about 500 million years ago Symbiotic relationships between plants and fungi are common today and date from this time Applying mathematical models (e.g., Hardy-Weinberg formula) to convert a data set from a table of numbers reflecting a change in the genetic makeup of a population over time and explaining the cause(s) and effect(s) of this change, such as natural selection, genetic drift, changes in population size, migration, mutations, and nonrandom mating. Hardy-Weinberg Their formulas are used to establish allele frequencies at genetic equilibrium (no evolution is occurring) The following conditions must all be fulfilled The population is very, very large There is no migration of individuals No mutations Mating is completely random All members survive and reproduce successfully The formulas Allele frequency: fraction of that particular allele in the population The sum of all the allele frequencies = 1 p = frequency of the dominant allele q = frequency of the recessive allele p+q=1 To figure the frequency of each genotype use: p2 + 2pq + q2 = 1 Microevolution Change in relative allele frequency over time; if allele frequency changes evolution occurs Causes of microevolution Genetic drift: change in a small gene pool due to chance Bottleneck event: population size is drastically reduced, leaving only the alleles of the survivors in the gene pool Founder effect: the small group starting a new colony contribute only their alleles to the new population Gene flow: gain or loss of alleles through immigration or emigration Non-random mating: organisms tend to mate with neighbors although they are capable of mating with any member of their species anywhere on earth