PAUL STEPHEN CORN
USGS NORTHERN ROCKY MOUNTAIN SCIENCE CENTER
ALDO LEOPOLD WILDERNESS RESEARCH INSTITUTE
790 EAST BECKWITH AVENUE, MISSOULA, MONTANA 59801
(406) 542-4191, FAX (406) 542-4196
Email: scorn@usgs.gov
CURRENT POSITION
Research Zoologist (GS-15). Northern Rocky Mountain Science Center, USGS, and Aldo Leopold
Wilderness Research Institute. My duties involve applied and basic research into the natural history,
ecology, and conservation biology of nongame and Federally-listed amphibians and reptiles in the
western United States, particularly in relation to protected lands and wilderness. I participated in
the creation of the USGS Amphibian Research and Monitoring Initiative and serve as Principal
Investigator for the Rocky Mountain Region. I was based at the Midcontinent Ecological Science
Center (Fish and Wildlife Service, National Biological Service, USGS) in Fort Collins, Colorado, until I
moved to the Leopold Institute in August, 1996.
PRIOR EXPERIENCE
May 1988 to March 1991: Zoologist (GS-11). Ecology and Systematics Section, National Ecology
Research Center, U.S. Fish and Wildlife Service, Fort Collins, Colorado.
October 1988 to April 1989: Wildlife Biologist (GS-11). Pacific Northwest Research Station, Forestry
Sciences Laboratory, U.S. Forest Service, Olympia, Washington.
May 1983 to May 1988: Wildlife Biologist (GS-9). Ecology and Systematics Section, National Ecology
Research Center, U. S. Fish and Wildlife Service, Fort Collins, Colorado.
April 1982 to May 1983: Self-Employed Zoologist. Obtained contract to catalog amphibians and
reptiles in the Fish and Wildlife Service Biological Survey collection
October 1979 to September 1980: Zoologist (GS-7). U.S. Fish and Wildlife Service, Denver Wildlife
Research Center, Fort Collins, Colorado
September 1974 to May 1976; August 1978 to May 1981: Graduate Teaching Assistant. Department of Zoology and Entomology, Colorado State University
EDUCATION
Ph.D. in Zoology, 1982. Colorado State University, Fort Collins. Dissertation: Selection Pressures
Affecting a Dorsal Color Polymorphism in Rana pipiens. (David Pettus, Major Professor)
M.S. in Zoology, 1977. Colorado State University.
B.S. with Distinction in Biology, 1974. University of Illinois, Urbana-Champaign.
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CURRICULUM VITAE
COMPETITIVE GRANTS
2012 USGS National Park Monitoring Project: Status and Trend of Amphibians on a Transect on the
Continental Divide: Integrating an NPS Vital Sign with the USGS Amphibian Research and
Monitoring Initiative across Networks. Co-PI with Erin Muths, Fort Collins Science Center and
Kristen Legg, NPS Greater Yellowstone Inventory and Monitoring Network. $79.5K
2012 USGS Data Rescue Program: Capture-recapture and disease data from 1960s: an important
addition to boreal chorus frog biology. Co-PI with Erin Muths (award to Ft Collins Science
Center). 2 years, $21K
2010 USGS Park Oriented Biological Support: Assessing the Threat of Climate Change to Headwater
Amphibians in Glacier National Park. Co-PI with Winsor Lowe, University of Montana, 2 years,
$71K
2008 USGS Park Oriented Biological Support: Identifying Factors Driving the Divergent Effects of
Disease on Amphibians in Two National Park Ecosystems In the Rocky Mountains. Co-PI with
Peter Murphy and Sophie St.-Hilaire, Idaho State University. 2 years, $75K; USGS Amphibian
Research and Monitoring Initiative: Sublethal Effects of Wildfire and Logging on Amphibians:
Synergistic Effects on Vigor, Stress, and Disease. Co-PI with Blake Hossack and Winsor Lowe,
University of Montana, 3 years. $74K.
2006 USGS Park Oriented Biological Support: Assessing the Distribution and Effects of Chytrid
Fungus on Amphibians in Grand Teton National Park. Co-PI with Sophie St.-Hilaire and Peter
Murphy, Idaho State University. 2 years, $61K
2005 USGS Amphibian Research and Monitoring Initiative: Developing a Statewide Base Level
Inventory in Montana. Co-PI with Bryce Maxell, Montana Heritage Program. 3 years, $74K; Joint
Fire Sciences Program: Effects of Prescribed and Wildland Fire on Aquatic Ecosystems in
Western Forests. Co-PI with David Pilliod, Cal Poly San Luis Obispo and Bruce Bury (extension of
original project). 2 years, $75K
2004 National Park Service Rocky Mountain Cooperative Ecosystem Studies Unit: Establishing the
Distribution of Chytrid Fungus in Grand Teton National Park and Evaluating the Effects on Boreal
Toads, Co-PI with Sue Wolff, Grand Teton National Park. 1 year, $10K
2003 USGS Amphibian Research and Monitoring Initiative: Amphibian Monitoring on the Great
Divide Transect: Greater Yellowstone Ecosystem, Co-PI with Charles Peterson, Idaho State
University. 1 year, $60K; Effects of Fire Suppression and Exclusion on Boreal Toad (Bufo boreas)
Populations, Co-PI with Blake Hossack, 2 years, $53K
2002 Joint Fire Sciences Program: Effects of Prescribed and Wildland Fire on Aquatic Ecosystems in
Western Forests. Co-PI with David Pilliod, US Forest Service and Bruce Bury, USGS. 3 years,
$372K; USGS Amphibian Research and Monitoring Initiative: Amphibian Monitoring on a North
to South Transect in the Rocky Mountains: Greater Yellowstone Ecosystem. Co-PI with Charles
Peterson, Idaho State University. 1 year, $49K
2001 US Forest Service Region 1/4 Adaptive Management and Monitoring Program: Effects of
Wildland Fire and Prescribed Fire on Amphibians and Aquatic Habitats in the Northern Rocky
Mountains. Co-PI with David Pilliod, US Forest Service. 3 years, $150K
1998 USGS Midcontinent Ecological Science Center: Examining an Intact "Metapopulation" of Boreal
Toads (Bufo boreas): Use of Habitat and Large Scale Movements. Co-PI with Erin Muths. 5 years,
$115K
1996 National Biological Service Species at Risk: Geographic Variation in Resistance of Boreal Toads
to Ultraviolet Radiation. 2 years, $53K
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HONORS AND AWARDS
Science Excellence Award, U.S. Geological Survey, 2004
Star Award, U.S. Geological Survey, 1998, 2000, 2001, 2002, 2005, 2011, 2012
Superior Accomplishment Award, National Biological Service; 1995
Special Achievement Awards, U.S. Fish and Wildlife Service; 1987, 1988, 1990–1992
Best poster, 1991 meeting of the Colorado Chapter of The Wildlife Society.
Best student paper presented at the 1979 meeting of the Southwestern and Rocky Mountain
Division of the American Association for the Advancement of Science
Illinois County Scholarship, 1970–1974
Edmund J. James Scholar, University of Illinois, 1970–1974
PROFESSIONAL SOCIETIES
Ecological Society of America
Society for Conservation Biology
Society for Northwestern Vertebrate
Zoology
Society for the Study of Amphibians and Reptiles
Species Survival Commission, IUCN
The Wildlife Society
OTHER DUTIES AND SERVICE
Academic Institutions
Affiliate faculty: College of Forestry, Department of Ecosystem and Conservation Sciences,
University of Montana, 1996–present; Department of Biological Sciences, Idaho State University,
Pocatello, 1992–present; Department of Biology, Colorado State University, Fort Collins, 1995–
1996. Committee member for the following graduate students:
Rebecca McCaffery, PhD, University of Montana, 2010. Population dynamics of the Columbia spotted frog
(Rana luteiventris): inference from long-term demography.
Bryce A. Maxell, PhD, University of Montana, 2009. State-wide assessment of status, predicted distribution,
and landscape-level habitat suitability of amphibians and reptiles in Montana.
Kathleen Griffin, PhD, University of Montana, 2007. Spatial population dynamics of western painted turtles
in a wetland ecosystem in northwestern Montana.
C. Gregory Guscio, MS, University of Montana, 2007. Responses of western toads (Bufo boreas) to changes
in terrestrial habitat resulting from wildfires.
Stephen J. Amish, MS, University of Montana, 2006. Ecosystem engineering beaver and the population of
Columbia spotted frogs in western Montana.
W. Christopher Funk, PhD, University of Montana, 2004. Patterns and consequences of dispersal in
Columbia spotted frogs (Rana luteiventris).
Aimee C. Wyrick, MS, University of Montana, 2004. Demography of the Columbia spotted frog (Rana
luteiventris) in the presence or absence of fish in the Absaroka-Beartooth Wilderness, Montana.
Paul E. Bartelt, PhD, Idaho State University 2000. A biophysical analysis of habitat selection in western toads
(Bufo boreas) in southeastern Idaho.
Mike Machura, MS Candidate University of Montana, 2007–2009 (did not finish)
Sarah Barnum, PhD Candidate, Colorado State University, 1995 (did not finish)
University of Montana, Senior honors thesis committee member for J. Tomasz Giermakowski, 1998–1999;
John Syslo, 2005–2006
Guest lecturer, Department of Biology, Colorado State University (occasional lectures in General
Biology, Vertebrate Zoology, Ecology, and Herpetology); Front Range Community College, Fort
Collins. Lecture on amphibian and reptile conservation. City of Fort Collins, Master Naturalist
Class
Graduate Teaching Assistant, Colorado State University. 1974–1976, 1978–1981.
Taught laboratory sections in General Biology, Cell Biology, Ornithology, Herpetology,
Vertebrate Zoology. Developed and taught a 1-credit Herpetology course.
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CURRICULUM VITAE
Professional Societies
Editor, Herpetological Conservation 1994–2007, Northwestern Naturalist 1992–1997,
Herpetological Review (Life History Notes) 1991–1992.
Associate Editor, Journal of Herpetology 2010–present.
Board Member, Northwest Chapter, Partners in Amphibian and Reptile Conservation 2009–
present
Judge, Society for the Study of Amphibians and Reptiles 2006 Annual Meeting Siebert Award
(student paper competition); Herpetologists’ League 1997 Annual Meeting student paper
competition.
Co-Chair, Rocky Mountain Working Group, Declining Amphibian Populations Task Force, IUCN;
participated in the September 2005 Amphibian Conservation Summit, sponsored by IUCN and
Wildlife Conservation International, to develop an Amphibian Conservation Action Plan for the
new IUCN-SSC Amphibian Specialist Group.
Federal and State Agencies
USGS—Acting Director, Northern Rocky Mountain Science Center, May 2005
Bureau of Land Management—Technical Proposal Evaluation Committee
US Environmental Protection Agency—panel review of fellowship applications
Colorado Division of Wildlife—Boreal toad recovery team
U.S. Fish and Wildlife Service—Wyoming toad population habitat viability analysis
U.S. Forest Service—Black Hills National Forest, forest plan revisions; Committee to develop
ecosystem management on southwestern national forests; Scientific Analysis Team for latesuccession and old-growth forests of the Pacific Northwest, amphibian viability panel; Workshop
to develop wilderness air quality values; Workshop to evaluate old-growth and riparian
management on the Tahoe National Forest
Interagency—Boreal toad management strategy team; Interior Columbia Basin Ecosystem
Management Project; Forest Ecosystem Management Assessment Team, amphibian viability
panel
PERSONAL
Born 1 August 1952, Mattoon, Illinois
Married (Janelle Gingerich Corn), 1 child (Nicholas Darwin Corn)
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REFEREED OR MAJOR PUBLICATIONS
1980s
1. Fogleman JC, Corn PS, Pettus D. 1980. The genetic basis of a dorsal color polymorphism in Rana
pipiens. Journal of Heredity 71:439–440.
2. Corn PS. 1981. Field evidence for a relationship between color and developmental rate in the
northern leopard frog (Rana pipiens). Herpetologica 37:155–160.
3. Corn PS, Fogleman JC. 1984. Extinction of montane populations of leopard frogs (Rana pipiens) in
Colorado. Journal of Herpetology 18:147–152.
4. Corn PS. 1986. Genetic and developmental studies of albino chorus frogs (Pseudacris triseriata).
Journal of Heredity 77:164–168.
5. Corn PS, Bury RB. 1986. Habitat use and terrestrial activity by red tree voles (Arborimus longicaudus)
in Oregon. Journal of Mammalogy 67:404–406.
6. Corn PS, Bury RB. 1986. Morphological variation and zoogeography of racers (Coluber constrictor) in
the central Rocky Mountain region. Herpetologica 42:254–260.
7. Bury RB, Corn PS. 1987. Evaluation of pitfall trapping in northwestern forests: arrays of traps with
drift fences. Journal of Wildlife Management 51:112–119.
8. Bury RB, Corn PS. 1988. Douglas-fir forests in the Oregon and Washington Cascades: abundance of
terrestrial herpetofauna related to stand age and moisture. In: Szaro RC, Severson KE, Patton DR,
editors. Management of amphibians, reptiles, and small mammals in North America. Ft Collins,
CO: USDA Forest Service Rocky Mountain Forest and Range Experiment Station. General Technical
Report RM-166. p 11–22.
9. Bury RB, Corn PS. 1988. Responses of aquatic and streamside amphibians to timber harvest: a
review. In: Raedeke KJ, editor. Streamside management: riparian wildlife and forestry interactions.
Seattle: University of Washington Institute of Forest Resources. Contribution 59. p 165–181.
10. Corn PS, Bury RB. 1988. Distribution of the voles Arborimus longicaudus and Phenacomys intermedius in the central Oregon Cascades. Journal of Mammalogy 69:427–429.
11. Corn PS, Bury RB, Spies TA. 1988. Douglas-fir forests in the Oregon and Washington Cascades: is the
abundance of small mammals related to stand age and moisture? In: Szaro RC, Severson KE, Patton
DR, editors. Management of amphibians, reptiles, and small mammals in North America. Ft Collins,
CO: USDA Forest Service Rocky Mountain Forest and Range Experiment Station. General Technical
Report RM-166. p 340352.
12. Corn PS, Bury RB. 1989. Logging in western Oregon: responses of headwater habitats and stream
amphibians. Forest Ecology and Management 29:39–57.
13. Corn PS, Livo LJ. 1989. Leopard frog and wood frog reproduction in Colorado and Wyoming.
Northwestern Naturalist 70:1–9.
14. Corn PS, Stolzenburg W, Bury RB. 1989. Acid precipitation studies in Colorado and Wyoming: interim
report of surveys of montane amphibians and water chemistry. Washington: USDI Fish and Wildlife
Service. Biological Report 80(40.26). 56 p.
15. Jones LLC, Corn PS. 1989. Third specimen of a metamorphosed Cope's giant salamander
(Dicamptodon copei). Northwestern Naturalist 70:37–38.
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CURRICULUM VITAE
1990s
16. Corn PS, Bury RB. 1990. Sampling methods for terrestrial amphibians and reptiles. Portland, OR:
USDA Forest Service Pacific Northwest Research Station. General Technical Report PNW-GTR-256.
34 p.
17. Jones LLC, Bury RB, Corn PS. 1990. Field observation of the development of a clutch of Pacific giant
salamander (Dicamptodon tenebrosus) eggs. Northwestern Naturalist 71:93–94.
18. Bury RB, Corn PS. 1991. Sampling methods for amphibians in streams in the Pacific Northwest.
Portland, OR: USDA Forest Service Pacific Northwest Research Station. General Technical Report
PNW-GTR-275. 29 p.
19. Bury RB, Corn PS, Aubry KB. 1991. Regional comparisons of terrestrial amphibian communities in
Oregon and Washington. In: Ruggiero LF, Aubry KB, Carey AB, Huff MH, technical coordinators.
Wildlife and vegetation of unmanaged Douglas-fir forests. Portland, OR: USDA Forest Service
Pacific Northwest Research Station. General Technical Report PNW-GTR-285. p 341–350.
20. Bury RB, Corn PS, Aubry KB, Gilbert FF, Jones LLC. 1991. Aquatic amphibian communities in Oregon
and Washington. In: Ruggiero LF, Aubry KB, Carey AB, Huff MH, technical coordinators. Wildlife and
vegetation of unmanaged Douglas-fir forests. Portland, OR: USDA Forest Service Pacific Northwest
Research Station. General Technical Report PNW-GTR-285. p 353–362.
21. Corn PS, Bury RB. 1991. Small mammal communities in the Oregon Coast Range. In: Ruggiero LF, Aubry
KB, Carey AB, Huff MH, technical coordinators. Wildlife and vegetation of unmanaged Douglas-fir
forests. Portland, OR: USDA Forest Service Pacific Northwest Research Station. General Technical
Report PNW-GTR-285. p 241–254.
22. Corn PS, Bury RB. 1991. Terrestrial amphibian communities in the Oregon Coast Range. In: Ruggiero LF,
Aubry KB, Carey AB, Huff MH, technical coordinators. Wildlife and vegetation of unmanaged
Douglas-fir forests. Portland, OR: USDA Forest Service Pacific Northwest Research Station. General
Technical Report PNW-GTR-285. p 305–317.
23. Corn PS, Vertucci FA. 1992. Descriptive risk assessment of the effects of acidic deposition on Rocky
Mountain amphibians. Journal of Herpetology 26: 361–369.
24. Bury RB, Esque TC, Corn PS. 1994. Conservation of desert tortoises (Gopherus agassizii): genetics
and protection of isolated populations. In Beaman KR, editor. Proceedings of 1987–1991 symposia.
Palm Springs, CA: The Desert Tortoise Council. p 59–66.
25. Corn PS. 1994. Displacement of desert tortoises: overview of a study at the Apex Heavy Industrial
Use Zone, Clark County, Nevada. In: Beaman KR, editor. Proceedings of 1987–1991 symposia. Palm
Springs, CA: The Desert Tortoise Council. p 295–303.
26. Corn PS. 1994. Recent trends in desert tortoise populations in the Mojave Desert. In: Bury RB,
Germano DJ, editors. Biology of North American tortoises. Washington: USDI National Biological
Survey. Fish and Wildlife Research 13. p 85–93.
27. Corn PS. 1994. Straight-line drift fences and pitfall traps. In: Heyer WR, Donnelly MA, McDiarmid
RW, Hayek LC, Foster MS, editors. Measuring and monitoring biological diversity: standard
methods for amphibians. Washington: Smithsonian Institution Press. p 109–117.
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28. Corn PS. 1994. What we know and don't know about amphibian declines in the West. In: Covington
WW, DeBano LF, technical coordinators. Sustainable ecological systems: implementing an ecological
approach to land management. Proceedings of a symposium 12–15 July 1993; Flagstaff, AZ. Fort
Collins, CO: USDA Forest Service Rocky Mountain Forest and Range Experiment Station. General
Technical Report RM-247. p. 59–67.
29. Kaufmann MR, Graham RT, Boyce DA Jr, Moir WH, Perry L, Reynolds RT, Bassett RL, Mehlhop P,
Edminster CB, Block WM, Corn PS. 1994. An ecological basis for ecosystem management. Fort
Collins, CO: USDA Forest Service Rocky Mountain Forest and Range Research Station. General
Technical Report RM-246. 22 p.
30. Bury RB, Corn PS. 1995. Have desert tortoises undergone a long-term decline in abundance? Wildlife
Society Bulletin 23:41–47.
31. Bury RB, Corn PS, Dodd CK Jr, McDiarmid RW, Scott NJ Jr. 1995. Amphibians. In LaRoe ET, Farris GS,
Puckett CE, Doran PD, Mac MJ, editors. Our living resources. A report to the nation on the
distribution, abundance, and health of U.S. plants, animals, and ecosystems. Washington: USDI
National Biological Service. p 124–126.
32. Corn PS, Peterson CR. 1996. Prairie legacies—amphibians and reptiles. In: Samson FB, Knopf FL,
editors. Prairie Conservation: Preserving North America's Most Endangered Ecosystem. Covelo,
CA: Island Press. p 125–134.
33. Vertucci FA, Corn PS. 1996. Evaluation of episodic acidification and amphibian declines in the Rocky
Mountains. Ecological Applications 6:449–457.
34. Corn PS, Jennings ML, Muths E. 1997. Survey and assessment of amphibian populations in Rocky
Mountain National Park. Northwestern Naturalist 78:34–55.
35. Muths E, Corn PS. 1997. Basking by boreal toads (Bufo boreas boreas) during the breeding season.
Journal of Herpetology 31:426–428.
36. Corn PS. 1998. Effects of ultraviolet radiation on boreal toads in Colorado. Ecological Applications
8:18–26.
2000–2005
37. Corn PS. 2000. Amphibian declines: review of some current hypotheses. In: Sparling DW, Bishop CA,
Linder G, editors. Ecotoxicology of amphibians and reptiles. Pensacola FL: Society of
Environmental Toxicology and Chemistry. p 663–696.
38. Muths E, Corn PS, Stanley TR. 2000. Use of oxytetracycline in batch-marking post-metamorphic
boreal toads. Herpetological Review 31:28–32.
39. Corn PS, Muths E, Iko WM. 2000. A comparison in Colorado of three methods to monitor breeding
amphibians. Northwestern Naturalist 81:22–30.
40. Corn PS, Knapp RA. 2000. Fish Stocking in protected areas: summary of a workshop. In: Cole DN,
McCool SF, Borrie WT, O’Loughlin J, compilers. Wilderness science in a time of change conference—
Volume 5: Wilderness ecosystems, threats, and management. Ogden, UT: U.S. Department of
Agriculture, Forest Service, Rocky Mountain Research Station. Proceedings RMRS-P-0-VOL-5. p
301–303.
41. Muths E, Corn PS. 2000. Boreal toads. In: Reading RP, Miller BJ, editors. Endangered animals: a
reference guide to conflicting issues. Westport, CT: Greenwood Press. p 60–65.
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CURRICULUM VITAE
42. Anderson DR, Burnham KP, Lubow BC, Thomas L, Corn PS, Medica PA, Marlow RW. 2001. Field trials
of line transect methods applied to estimation of desert tortoise abundance. Journal of Wildlife
Management 65:583–597.
43. Knapp RA, Corn PS, Schindler DE. 2001. The introduction of nonnative fish into wilderness lakes:
good intentions, conflicting mandates, and unintended consequences. Ecosystems 4:275–278.
44. Muths E, Johnson TL, Corn PS. 2001. Experimental repatriation of boreal toad (Bufo boreas) eggs,
metamorphs, and adults in Rocky Mountain National Park. Southwestern Naturalist 46:106–113.
45. Corn PS, Muths E. 2002. Variable breeding phenology affects the exposure of amphibian embryos to
ultraviolet radiation. Ecology 83:2958–2963.
46. Boone MD, Corn PS, Donnelly MA, Little EE, Niewiarowski PH. 2003. Physical stressors. In: Linder G,
Krest SK, Sparling DW, editors. Amphibian decline: an integrated analysis of multiple stressor
effects. Pensacola, FL: Society of Environmental Toxicology and Chemistry. p 129–151.
47. Corn PS. 2003. Amphibian breeding and climate change: the importance of snow in the mountains.
Conservation Biology 17:622–625.
48. Corn PS. 2003. Deteriorating status of western amphibians: can we generalize about causes? In:
Linder G, Krest SK, Sparling DW, editors. Amphibian decline: an integrated analysis of multiple
stressor effects. Pensacola, FL: Society of Environmental Toxicology and Chemistry. p 249–255.
49. Corn PS. 2003. Endangered toads in the Rockies. In: Taylor L, Martin K, Hik D, Ryall A, editors. Ecological
and earth sciences in mountain areas. Banff, AB: The Banff Centre. p 43–51.
50. Corn PS, Bury RB, Hyde EJ. 2003. Conservation of North American stream amphibians. In: Semlitsch
R, editor. Amphibian conservation. Washington, DC: Smithsonian Institution Press. p 24–36.
51. Muths E, Campbell DH, Corn PS. 2003. Hatching success in salamanders and chorus frogs at two sites
in Colorado, USA: effects of acidic deposition and climate. Amphibia-Reptilia 24:27–36.
52. Muths E, Corn PS, Pessier AP, Green DE. 2003. Evidence for disease-related amphibian decline in
Colorado. Biological Conservation 110:357–365.
53. Pilliod DS, Bury RB, Hyde EJ, Pearl, CA, Corn PS. 2003. Fire and amphibians in North America. Forest
Ecology and Management 178:163–181.
54. Campbell DH, Muths E, Turk JT, Corn PS. 2004. Sensitivity to acidification of subalpine ponds and
lakes in northwestern Colorado. Hydrological Processes 18:2817–2834.
55. Corn PS, Muths E. 2004. Variable breeding phenology affects the exposure of amphibian embryos to
ultraviolet radiation: reply. Ecology 85:1759–1763.
56. Adams MJ, Hossack, BR, Knapp RA, Corn PS, Diamond SA, Trenham PC, Fagre D. 2005. Distribution
patterns of lentic-breeding amphibians in relation to ultraviolet radiation exposure in western North
America. Ecosystems 8:488–500.
57. Brooks PD, O’Reilly CM, Diamond SA, Campbell DH, Knapp RA, Bradford, DM, Corn PS, Hossack BR,
Tonnessen KA. 2005. Spatial and temporal variability in the amount and source of dissolved organic
carbon: implications for UV exposure in amphibian habitats. Ecosystems 8:478–487.
58. Carey C, Corn PS, Jones MS, Livo LJ, Muths E, Loeffler CW. 2005. Factors limiting the recovery of boreal
toads (Bufo b. boreas). In: Lannoo M, editor. Amphibian declines: the conservation status of United
States species. Berkeley: University of California Press. p 222–236.
59. Corn PS. 2005. Climate change and amphibians. Animal Biodiversity and Conservation 28:59–67.
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60. Corn PS, Adams MJ, Battaglin WA, Gallant AL, James DL, Knutson M, Langtimm CA, Sauer JR. 2005.
Amphibian Research and Monitoring Initiative: concepts and implementation. Reston, VA: U.S.
Geological Survey. Scientific Investigations Report 2005-5015. 23 p.
61. Corn PS, Hossack BR, Muths E, Patla DA, Peterson CR, Gallant AL. 2005. Status of amphibians on the
Continental Divide: surveys on a transect from Montana to Colorado, USA. Alytes 22:85–94.
62. Corn PS, Muths E, Adams MJ, Dodd CK Jr. 2005. The U. S. Geological Survey’s Amphibian Research
and Monitoring Initiative. Alytes 22:65–71.
63. Diamond SA, Trenham PC, Adams MJ, Hossack, BR, Knapp RA, Stark SL, Bradford, DM, Corn PS,
Czarnowski K, Brooks, PD, Fagre D, Breen B, Detenbeck NE, Tonnessen KA. 2005. Estimated ultraviolet
radiation doses in wetlands in six national parks. Ecosystems 8:462–477.
64. Funk WC, Blouin MS, Corn PS, Maxell BA, Pilliod DS, Amish S, Allendorf FW. 2005. Population
structure of Columbia spotted frogs (Rana luteiventris) is strongly affected by the landscape.
Molecular Ecology 14:483–496.
65. Funk WC, Greene AE, Corn PS, Allendorf FW. 2005. High dispersal in a frog species suggests that it is
vulnerable to habitat fragmentation. Biology Letters 1:13–16.
66. Hossack BR, Pilliod DS, Corn PS. 2005. Lack of significant changes in the herpetofauna of Theodore
Roosevelt National Park, North Dakota, since the 1920s. American Midland Naturalist 154:423–432.
67. Muths E, Jung RE, Bailey L, Adams MJ, Corn PS, Dodd CK Jr, Fellers GM, Sadinski WJ, Schwalbe CR,
Walls SC, Fisher RN, Gallant AL, Battaglin WA , Green DE. 2005. The U.S. Department of Interior’s
Amphibian Research and Monitoring Initiative: a successful start to a national program. Applied
Herpetology 2:355–371.
68. Odum RA, Corn PS. 2005. Bufo baxteri Porter, 1968. Wyoming toad. In: Lannoo M, editor.
Amphibian declines: the conservation status of United States species. Berkeley: University of
California Press. p 390–392.
69. Scherer RD, Muths E, Noon BR, Corn PS. 2005. An evaluation of weather and disease as causes of decline
in two populations of boreal toads. Ecological Applications 15:2150–2160.
2006–2010
70. Hossack BR, Corn PS, Fagre DB. 2006. Divergent patterns of abundance and age-class structure of
headwater stream tadpoles in burned and unburned watersheds. Canadian Journal of Zoology
84:1482–1488.
71. Hossack BR, Diamond SA, Corn PS. 2006. Distribution of boreal toad populations in relation to
estimated UV-B dose in Glacier National Park, Montana, USA. Canadian Journal of Zoology 84:98–
107.
72. Karraker NE, Pilliod DS, Adams MJ, Bull EL, Corn PS, Diller LV, Dupuis LA, Hayes MP, Hossack BR,
Hodgson GR, Hyde EJ, Lohman K, Norman BR, Ollivier LM, Pearl CA, Peterson CR. 2006. Taxonomic
and geographic variation in oviposition by tailed frogs (Ascaphus spp.). Northwestern Naturalist
87:87–97.
73. Muths E, Scherer RD, Corn PS, Lambert BR. 2006. Estimation of temporary emigration in male toads.
Ecology 87:1048–1056.
74. Corn PS. 2007. Amphibians and disease: implications for conservation in the Greater Yellowstone
Ecosystem. Yellowstone Science 15(2):11–16.
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CURRICULUM VITAE
75. Field KJ, Tracy CR, Medica PA, Marlow RW, Corn PS. 2007. Return to the wild: translocation as a tool
in conservation of the desert tortoise (Gopherus agassizii). Biological Conservation 136:232–245.
76. Hossack BR, Corn PS. 2007. Responses of pond-breeding amphibians to wildfire: short-term patterns
in occupancy and colonization. Ecological Applications 17:1403–1410.
77. Guscio CG, Hossack BR, Eby LA, Corn PS. 2008. Post-breeding habitat use by adult boreal toads (Bufo
boreas boreas) after wildfire in Glacier National Park, USA. Herpetological Conservation and Biology
3:55–62.
78. Hossack BR, Corn PS. 2008. Wildlfire effects on water temperature and selection of breeding sites by
the boreal toad (Bufo boreas) in seasonal wetlands. Herpetological Conservation and Biology 3:46–
54.
79. Corn PS. 2009. Selection of species and sampling areas: the importance of inference. In: Dodd CK Jr,
editor. Amphibian ecology and conservation, a handbook of techniques. New York: Oxford
University Press. p 431–446.
80. Corn PS. 2009. The USGS Amphibian Research and Monitoring Initiative in the Pacific Northwest. In:
Olson DH, coordinating editor. Herpetological conservation in northwestern North America.
Northwestern Naturalist 90:66.
81. Goebel AM, Ranker TA, Corn PS, Olmstead RG. 2009. Mitochondrial DNA evolution in the Anaxyrus
boreas species group. Molecular Phylogenetics and Evolution 50:209–225.
82. Hossack BR, Eby LA, Guscio CG, Corn PS. 2009. Thermal characteristics of amphibian microhabitats in
a fire-disturbed landscape. Forest Ecology and Management 258:1414–1421.
83. Hossack BR, Muths E, Anderson CW, Kirshtein JA, Corn PS. 2009. Distribution limits of
Batrachochytrium dendrobatidis: a case study in the Rocky Mountains, USA. Journal of Wildlife
Diseases 45:1198–1202.
84. Murphy PJ, St-Hilaire S, Bruer S, Corn PS, Peterson CR. 2009. Distribution and pathogenicity of
Batrachochytrium dendrobatidis in boreal toads from the Grand Teton area of western Wyoming.
EcoHealth 6:109–120.
85. Hossack BR, Adams MJ, Grant EHC, Pearl CA, Bettaso JB, Barichivich WJ, Lowe WH, True K, Ware JL,
Corn PS. 2010. Low prevalence of chytrid fungus (Batrachochytrium dendrobatidis) in amphibians of
U.S. headwater streams. Journal of Herpetology 44:253–260.
86. Pilliod DS, Hossack BR, Bahls PF, Bull EL, Corn PS, Hokit G, Maxell BA, Munger JC, Murphy P, Wyrick
A. 2010. Non-native salmonids affect amphibian occupancy at multiple spatial scales. Diversity and
Distributions 16:959–974.
87. Pilliod DS, Muths E, Scherer RD, Bartelt PE, Corn PS, Hossack BR, Lambert BA, McCaffery R, Gaughan
C. 2010. Effects of amphibian chytrid fungus on individual survival probability in wild boreal toads.
Conservation Biology 24:1259–1267.
2011–
88. Corn PS, Muths E, Kissel AM, Scherer RD. 2011. Breeding chorus indices are weakly related to
estimated abundance of boreal chorus frogs. Copeia 2011:365–371.
P.S. CORN
11
89. Corn PS, Muths E, Pilliod DS. 2011. Long-term observations of boreal toads at an ARMI apex site. In:
Anderson C, editor. Questioning Greater Yellowstone’s Future: Climate, Land Use, and Invasive
Species. Proceedings of the 10th Biennial Scientific Conference on the Greater Yellowstone
Ecosystem. Yellowstone National Park, WY, and Laramie, WY: Yellowstone Center for Resources and
University of Wyoming William D. Ruckelshaus Institute of Environment and Natural Resources. p
101–104.
90. Murphy PJ, St-Hilaire S, Corn PS. 2011. Temperature, hydric environment, and prior pathogen
exposure alter the experimental severity of chytridiomycosis in boreal toads. Diseases of Aquatic
Organisms 95:31–42.
91. Shoo LP, Olson DH, McMenamin SK, Murray KA, Van Sluys M, Donnelly MA, Stratford D, Terhivuo J,
Merino-Viteri A, Herbert SM, Bishop PJ, Corn PS, Dovey L, Griffiths RA, Lowe K, Mahony M,
McCallum H, Shuker JD, Simpkins C, Skerratt LF, Williams SE, Hero JM. 2011. Engineering a future for
amphibians under climate change. Journal of Applied Ecology 48:487–492.
92. Gould WR, Patla DA, Daley R, Corn PS, Hossack BR, Bennetts R, Peterson CR. 2012. Estimating
occupancy in large landscapes: evaluation of amphibian monitoring in the Greater Yellowstone
Ecosystem. Wetlands 32:379–389.
93. Nussear KE, Tracy CR, Medica PA, Wilson DS, Marlow RW, Corn PS. 2012. Translocation as a
conservation tool for Agassiz's desert tortoises: survivorship, reproduction, and movements. Journal
of Wildlife Management 76:1341–1353.
94. Adams MJ, Miller DAW, Muths E, Corn PS, Grant EHC, Bailey LL, Fellers G, Fisher RN, Sadinski WJ,
Waddle H, Walls SC. 2013. Trends in amphibian occupancy in the United States. PLoS ONE 8(5):
e64347.
95. Hossack BR, Adams MJ, Pearl CA, Wilson K, Bull E, Lohr K, Patla DA, Pilliod DS, Jones JL, Wheeler K,
McKay S, Corn PS. 2013. Roles of patch characteristics, drought frequency, and restoration in longterm trends of a widespread amphibian. Conservation Biology 27:1410–1420.
96. Hossack BR, Lowe WH, Corn PS. 2013. Rapid increases and time-lagged declines in amphibian
occupancy after wildfire. Conservation Biology 27:219–228.
97. Hossack BR, Lowe WH, Honneycutt RK, Parks SA, Corn PS. 2013. Interactive effects of wildfire, forest
management, and isolation on amphibian and parasite abundance. Ecological Applications 23:479–
492.
98. Hossack BR, Lowe WH, Ware JL, Corn PS. 2013. Disease in a dynamic landscape: Host behavior and
natural disturbance reduce prevalence of amphibian chytrid. Biological Conservation 157:293–299.
99. Hossack BR, Lowe WH, Webb MAH, Talbott MJ, Kappenman KM, Corn PS. 2013. Population-level
thermal performance of a cold-water ectotherm is linked to ontogeny and local environmental
heterogeneity. Freshwater Biology 58:2215–2225.
100.McCaffery RM, Eby LA, Maxell BA, Corn PS. 2014. Breeding site heterogeneity reduces variability in
frog recruitment and population dynamics. Biological Conservation 170:169–176.
Submitted/In Prep
101.Amish S, Maxell BA, McIntire E, Eby LA, Corn PS. In prep. Ecosystem engineering: beaver, landscape
patterns of lentic habitat, and the distribution of Columbia spotted frogs in southwestern Montana.
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CURRICULUM VITAE
102.Corn PS, Battaglin WA, Clark M, Hossack BR, Muths E, Olson DH, Patla DA, Pilliod DS, Vredenburg VT.
In prep. Amphibians and snow: estimated changes in breeding phenology in the mountains of the
western United States.
NON-REFEREED REVIEWS, ABSTRACTS, NOTES, AND LETTERS
1. Corn PS. 1979. Size at metamorphosis and growth rates of juvenile frogs from montane populations
of Rana pipiens. Journal of the Colo-Wyo Academy of Sciences 11:90. [abstract]
2. Corn PS. 1980. Polymorphic reproductive behavior in male chorus frogs (Pseudacris triseriata).
Journal of the Colo-Wyo Academy of Sciences 12:6–7. [abstract]
3. Corn PS. 1980. Comment on the occurrence of Pseudacris clarki in Montana. Bulletin of the Chicago
Herpetological Society 15:77–78.
4. Corn PS. 1982. Color and pattern polymorphism in anurans. Bulletin of the Chicago Herpetological
Society 17:57–68.
5. Corn PS. 1982. Ecological genetics of albino chorus frogs (Pseudacris triseriata). Journal of the ColoWyo Academy of Sciences 14:57. [abstract]
6. Corn PS. 1982. Natural selection in leopard frog tadpoles. Bulletin of the Ecological Society of
America 63:83. [abstract]
7. Corn PS, Bury RB, Welsh HH. 1984. A selected bibliography of Wyoming amphibians and reptiles.
Smithsonian Herpetological Information Service 59:1–34.
8. Corn PS, Gingerich LJ. 1987. Geographical distribution: Phrynosoma douglassii brevirostre.
Herpetological Review 18:20.
9. [Corn PS] 1987. Rediscovery of the Wyoming toad. Endangered Species Technical Bulletin 12(11–
12):7.
10. Corn PS, Bury RB. 1988. Habitat associations of terrestrial vertebrates along a chronosequence of
Douglas-fir forests in the Coast Range of Oregon. Bulletin of the Ecological Society of America
69(suppl):107. [abstract]
11. Bury RB, Corn PS. 1988. Comparison of aquatic amphibians in logged and old-growth forests.
Bulletin of the Ecological Society of America 69(suppl):88. [abstract]
12. Vertucci, FA, Corn PS. 1993. Episodic acidification is not responsible for amphibian declines in the
Rocky Mountains, USA. Bulletin of the Ecological Society of America 74(suppl):470. [abstract]
13. Corn PS. 1993. Bufo boreas (boreal toad). Predation. Herpetological Review 24:57.
14. Corn PS. 1994. Common factors and contradictions in amphibian declines. Northwest Science
68:121. [abstract]
15. Vertucci FA, Corn S. 1994. Acidification and salamander recruitment: reply. BioScience 44(3):126,
cover 3. [letter]
16. Corn PS. 1995. Wildlife 2001: populations. Copeia 1995:1001–1002. [book review]
17. Johnson TL, Muths E, Corn PS. 1997. Experimental methods for restoring boreal toad populations in
Rocky Mountain National Park. In Proceedings of the Conference on Research and Resource
Management in Parks and on Public Lands. 1997 George Wright Society Biennial Conference. p
255–257.
18. Corn S. 2000. Amphibian Research and Monitoring Initiative. Froglog 41:1–2.
P.S. CORN
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19. Corn PS. 2001. A plague of frogs: the horrifying true story. Quarterly Review of Biology. 76:64–65.
[book review]
20. Corn PS. 2001. Perspectives from the Aldo Leopold Wilderness Research Institute: Amphibians and
wilderness. International Journal of Wilderness 7(2):25.
21. Jennings M, Beiswenger R, Corn S, Parker M, Pessier A, Spencer B, Miller PS, editors. 2001. Population and
habitat viability assessment for the Wyoming toad (Bufo baxteri). Final Workshop Report. Apple Valley, MN:
IUCN/SSC Conservation Breeding Specialist Group. 104 p.
22. Corn PS. 2003. Endangered toads in the mountains. Vancouver, University of British Columbia
Centre for Alpine Research. Science Highlight (published online; no longer available).
23. Pilliod DS, Corn PS. 2003. Changes in stream amphibian populations following large fires in Idaho.
Northwestern Naturalist 84:110–111. [abstract]
24. Hossack BR, Corn PS. 2004. Responses of pond-breeding amphibians to wildfire in Glacier National
Park. Northwestern Naturalist 85:78. [abstract]
25. Karraker NF, Pilliod DS, Bull EL, Corn PS, Diller LV, Dupuis LA, Hayes MP, Hossack BR, Hodgson GR,
Hyde EJ, Lohman K, Norman BR, Ollivier LM, Pearl CA, Peterson CR. 2004. Taxonomic and geographic
variation in the oviposition of tailed frogs (Ascaphus spp.). Northwestern Naturalist 85:79. [abstract]
26. Pilliod DS, Corn PS, Bury RB, Hyde EJ. 2004. Effects of wildland fires on stream amphibian
populations in the greater Northwest. Northwestern Naturalist 85:85–86. [abstract]
27. Hossack BR, Corn PS. 2005. Changes in counts of Rocky Mountain tailed frog tadpoles after wildfire in
Glacier National Park, Montana. Northwestern Naturalist 86:99–100. [abstract]
28. Pilliod DS, Jain TB, Evans JS, Bury RB, Corn PS. 2005. Effects of burn severity on stream amphibian
populations. Northwestern Naturalist 86:113. [abstract]
29. Corn PS. 2006. Assessing bias in the estimation of abundance using distance sampling.
Northwestern Naturalist 87:156. [abstract]
30. Pounds A, Carnaval ACOQ, Corn S. 2007. Climate change, biodiversity loss, and amphibian declines.
In Gascon C, Collins JP, Moore RD, Church DR, McKay JE, Mendelson JR III, editors. Amphibian
conservation action plan. Gland, Switzerland and Cambridge, UK: IUCN/SSC Amphibian Specialist
Group. p 19–20.
31. Corn S. 2008. Climate change impacts to amphibians and reptiles. Northwestern Naturalist 89:126–
127. [abstract]
32. Corn PS. 2008. Perspectives from the Aldo Leopold Wilderness Research Institute: The U.S.
Geological Survey and wilderness research. International Journal of Wilderness 14(2):24, 33.
33. Corn S, Hossack B, Pilliod, D. 2009. Amphibians and fire in the northern Rockies. In Masters RE,
Galley KEM, Despain DG, editors. The ’88 fires: Yellowstone and beyond, conference proceedings.
Tall Timbers Research Station Miscellaneous Publication 16:89. [abstract]
34. Patla DA, Peterson CR, Corn PS. 2009. Amphibian decline in Yellowstone National Park. Proceedings
of the National Academy of Sciences, USA. 106(9):E22 [www.pnas.org_cgi_doi_10.1073_pnas.0812670106].
[letter]
35. Corn S, Muths E, Kissel A, Scherer R. 2011. How well do call indices represent abundance of breeding
anurans? Northwestern Naturalist 92:140–141. [abstract]
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CURRICULUM VITAE
36. Hossack BR, Lowe WH, Corn PS. 2011. Wildfire and fragmentation: effects on amphibian populations
and associated nematodes. Northwestern Naturalist 92:148. [abstract]
INVITED PAPERS
Trends in amphibian populations in US National Parks on the Continental Divide. (also coauthor on
Interactive effects of wildfire, forest management, and isolation on amphibian and parasite abundance).
Amphibian Monitoring Across the U.S.: Tracking Declining Populations in Diverse Habitats and Unique
Assemblages. 7th World Congress of Herpetology and Joint Meeting of Ichthyologists and Herpetologists.
Vancouver, BC, August 2012.
How well do call indices represent abundance of breeding anurans? (also coauthor on Wildfire and
fragmentation: effects on amphibian populations and associated nematodes). USGS ARMI in the West:
Populations, Stressors, Methods. Society for Northwestern Vertebrate Biology/Washington Chapter of
The Wildlife Society. Gig Harbor, WA, March 2011.
Fire and herpetofauna in the Northwest. Dry Forests and Dependent Wildlife: Yesterday, Today, and in
the Future. The Nature Conservancy. Bend, OR, November 2009.
Climate Change and Amphibians in Snow-Dominated Landscapes. Symposium: Amphibian Responses to
Climate Change. 10th International Congress of Ecology (INTECOL). Brisbane, Australia, August 2009.
Amphibians in the Crown of the Continent Ecosystem. Crown Managers Forum. Crown of the Continent
Managers Partnership. Pincher Creek, AB, February 2009.
Amphibians and Fire in the Northern Rockies. The ’88 Fires: Yellowstone and Beyond. International
Association of Wildland Fire/9th Yellowstone Science Conference. Jackson, WY, September 2008.
Climate Change Impacts to Amphibians and Reptiles. Northwest Connections: Sustaining Our Wildlife
Populations in the Face of Climate Change, Human Population Growth, and Energy Development.
Montana Chapter of the Wildlife Society/Society for Northwestern Vertebrate Biology, Missoula, MT,
February 2008.
Climate Change and Amphibians. MTNCLIM 2006. Consortium for Integrated Climate Research on
Western Mountains, Mt. Hood, OR, September 2006.
Current and Future Threats to Amphibian Populations. 2nd Alaska Amphibian Conference, Juneau, AK,
February 2006.
Amphibian Research in Glacier National Park. Glacier Science Conference, Glacier National Park, August
2004.
Endangered Toads in the Rockies. Ecological and Earth Sciences in Mountain Areas. Banff, AB,
September 2002.
Current and Future Effects of Climate Change on Montane Amphibians. Species, Water Quality, &
Climate (session). American Geophysical Union, Washington, DC, May 2002.
Amphibian Monitoring in National Parks. Symposium: Monitoring in National Parks. The Wildlife
Society, Reno, NV, September 2001.
Amphibian Declines: Status, Causes, & Cautions. Workshop: Global Decline of Amphibian Populations.
An Integrated Analysis of Multiple Stressor Effects. Society of Environmental Toxicology and Chemistry
and the Johnson Foundation, Wingspread, Racine WI., August 2001.
Status of amphibians in the western United States. Symposium: Declining Amphibians. Society for the
Study of Amphibians and Reptiles, Indianapolis, IN, July 2001.
P.S. CORN
15
Amphibian declines and global climate change. The Big Unknowns in Global Change: Climatic, Biotic,
Human Systems. University of Georgia State-of-the-Art Conference, Athens, GA, April 2001.
Effects of ultraviolet radiation on amphibians. Annual meeting of the NPS/EPA PRIMENet project.
Shenandoah National Park, VA, November 2000.
Effects of ultraviolet radiation on amphibians, and Department of Interior FY2000 amphibian initiative.
Annual meeting of the NPS/EPA PRIMENet project. Sequoia National Park, CA, November 1999.
Techniques for estimating abundance of desert tortoises. Interagency Management Oversight Group,
Las Vegas, NV, June 1999.
Declining Amphibians in Western North America: Research Addressing Park Service and Fish and Wildlife
Service Information Needs. USGS Midcontinent Ecological Science Center Client Coordination Meeting.
Ft. Collins, CO, February 1998.
Case Study: Effects of Global Change on Amphibians. USGS Biological Resources Division Scoping
Symposium on Global Change and Wetlands Ecology. Phoenix, AZ, February 1998
Keynote Address: Ultraviolet Radiation and Amphibians: Evaluation of Some Hypotheses. Southwest
U.S. Working Group, Declining Amphibian Populations Task Force. Phoenix, AZ, January 1998
Introduction (and session organizer): Survey and Monitoring Techniques for Amphibians. Conference on
Declining and Sensitive Amphibians in the Rocky Mountains and Pacific Northwest. Boise, ID,
November 1996
Monitoring amphibians: can we get the answers to the questions we ask? Symposium: Inventory and
Monitoring of Vertebrates. Society for Northwestern Vertebrate Biology, Corvallis, OR, March 1996
Declines of pond-breeding amphibians in the West: causes and recovery. Symposium: Candidate
Species. The Wildlife Society, Portland, OR, September 1995
What we know and don’t know about amphibian declines in the West. Conference on Sustainable
Ecological Systems. Flagstaff, AZ, July, 1993
Distribution and Status of Amphibians in Rocky Mountain National Park. Symposium: Amphibian
Monitoring in Western National Parks. Society for Northwestern Vertebrate Biology, Astoria, OR,
March, 1993
Declining amphibian populations. Western States and Provinces Nongame Symposium. Missoula, MT,
September, 1992
Potential for effects of acidification on amphibians in western North America. Symposium: Effects of
acidification on amphibians in North America. Society for the Study of Amphibians and Reptiles and
Herpetologists' League, Pennsylvania State University, 1991
Regional assessment of the status of amphibian populations: Rocky Mountains. Symposium: Declining
amphibian populations. Society for the Study of Amphibians and Reptiles and Herpetologists' League,
New Orleans, 1990
Declining amphibians in the Rockies. Workshop: Declining amphibian populations: a global
phenomenon? National Research Council Board on Biology, Irvine, CA, 1990
Small mammal communities in the Oregon Coast Range. Symposium: Old-growth Douglas-fir forests:
wildlife communities and habitat relationships. U.S. Forest Service, Portland OR, 1989
Terrestrial amphibian communities in the Oregon Coast Range. Symposium: Old-growth Douglas-fir
forests: wildlife communities and habitat relationships. U.S. Forest Service, Portland OR, 1989
16
CURRICULUM VITAE
The effects of timber harvest on aquatic amphibians. Symposium: Streamside management: riparian
wildlife and forestry interactions. University of Washington Institute of Forest Resources, Seattle, 1987
OFFERED PAPERS AND POSTERS
Pacific Northwest Partners in Amphibian and Reptile Conservation, Hood River, OR, March 2012
10th Yellowstone Science Conference, Mammoth, Wyoming, October 2010
Joint Meeting of Ichthyologists and Herpetologists (ASIH, HL, SSAR), Portland, Oregon, July 2009
Montana Chapters of The Wildlife Society and American Fisheries Society, Kalispell, Montana, February 2009*
Oregon Chapter of The Wildlife Society, Gleneden Beach, Oregon, February 2008*
Amphibian Declines & Chytridiomycosis (Partners in Amphibian and Reptile Conservation), Tempe, Arizona,
November 2007
George Wright Society, St. Paul, Minnesota, April 2007*
Northwest Science Association/ Washington-Oregon Chapter of The Wildlife Society, Pendleton, Oregon, April
2007*
Joint Meeting of Ichthyologists and Herpetologists (ASIH, HL, SSAR), New Orleans, Louisiana, July 2006
Society for Northwestern Vertebrate Biology, Olympia, Washington, March 2006
Northwest Science Association/ Idaho Chapter of The Wildlife Society, Boise, Idaho, March 2006
Society for Northwestern Vertebrate Biology, Corvallis, Oregon, March 2005
Ecological Society of America, Portland, Oregon, August 2004
Joint Meeting of Ichthyologists and Herpetologists (ASIH, HL, SSAR), Norman, Oklahoma, May 2004
Society of Environmental Toxicology and Chemistry, Austin, Texas, November 2003
American Society for Limnology and Oceanography, Salt Lake City, Utah, February 2003
Joint Meeting of Ichthyologists and Herpetologists (ASIH, HL, SSAR), Kansas City, Missouri, July 2002.
International Conference on Distance Sampling, St. Andrews, Scotland, August 2001.
Society for the Study of Amphibians and Reptiles/Herpetologists’ League/American Society of
Ichthyologists and Herpetologists, La Paz, Baja California Sur, June 2000.
Symposium, Wilderness Science in a time of Change, Missoula, Montana, May 1999.
Joint Meeting of Ichthyologists and Herpetologists (ASIH, HL, SSAR), Guelph, Ontario, July 1998
Joint Meeting of Ichthyologists and Herpetologists (ASIH, HL, SSAR), Seattle, Washington, June 1997
Desert Tortoise Council, Las Vegas, Nevada, April 1997*
Society for the Study of Amphibians and Reptiles, Lawrence, Kansas, July 1996
North American Amphibian Monitoring Program, Burlington, Ontario, September 1995
Society for Northwestern Vertebrate Biology, Orcas Island, Washington, March 1995
Society for Northwestern Vertebrate Biology, Ellensburg, Washington, March 1994
Society for the Study of Amphibians and Reptiles, Bloomington, Indiana, August, 1993
Ecological Society of America, Madison, Wisconsin, August, 1993*
Society for the Study of Amphibians and Reptiles, El Paso, Texas, August, 1992
Symposium, Biodiversity in the Rocky Mountains. Ft. Collins, Colorado, 1991
Desert Tortoise Council, Las Vegas, Nevada, 1991
Colorado Chapter of The Wildlife Society, Ft. Collins, 1991
Guild of Rocky Mountain Population Biologists, Univ Colo Mtn Research Station, 1989
First World Congress of Herpetology, Canterbury, England, 1989*
Ecological Society of America, University of California, Davis, 1988
U.S. Forest Service Symposium, Management of amphibians, reptiles, and small mammals in North
America. Flagstaff, Arizona, 1988
Colorado Chapter of The Wildlife Society, Fort Collins, 1987
National Acid Precipitation Assessment Program, Aquatic Effects Peer Review. New Orleans, 1987
Society for the Study of Amphibians and Reptiles/Herpetologists' League, Springfield, Missouri, 1986
P.S. CORN
American Society of Mammalogists, Arcata California, 1984
Society for the Study of Amphibians and Reptiles/Herpetologists' League, Salt Lake City, 1983
Ecological Society of America, Pennsylvania State University, 1982
Colorado-Wyoming Academy of Sciences, Ft. Collins, Colorado, 1982
Colorado-Wyoming Academy of Sciences, Denver, Colorado, 1980
AAAS, Southwestern and Rocky Mountain Division, Durango, Colorado, 1979
AAAS, Southwestern and Rocky Mountain Division, Phoenix, Arizona, 1976
*coauthor—not in attendance
SEMINARS & LECTURES
USGS Northern Rocky Mountain Science Center, Bozeman, Montana, 2013
US Fish and Wildlife Service Fish Technology Center, Bozeman, Montana, 2007
Flathead Audubon Society, Kalispell, Montana, 2005
Five Valleys Audubon Society, Missoula, Montana, 2004
University of Montana, 1998
University of Wyoming/National Park Service Research Center, 1998
Colorado State University, Society for Conservation Biology, 1995
Colorado Herpetological Society, 1991
Rocky Mountain Biological Laboratory, 1990
Colorado State University, Colloquium in the Life Sciences, 1987
Oregon State University, Dept. of Forest Sciences, 1986
17